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Orientador: Fresia Soledad Ricardi Torres Branco Tese (doutorado) - Universidade Estadual de Campinas, Instituto de Geociências Resumo: Neste estudo se analisou a composição taxonômica e as características fisionômicas do material foliar de sete amostras de serapilheira da floresta ripária da Estação Ecológica de Mogi Guaçú, Nordeste do estado de São Paulo, assim como de cinco acumulações de macro-restos vegetais que se encontram ao longo de rios meandrantes do Nordeste, Leste e litoral Sul do estado de São Paulo, com a finalidade de se obter novas pautas para a análise e interpretação das associações fósseis de angiospermas. Também se analisou a relação entre a proporção de espécies sem dentes de uma flora e a temperatura média anual (TMA) de um conjunto de 121 localidades da América do Sul, com o objetivo de se obter uma nova equação baseada na Análise da Margem Foliar para a região. Ao analisar o material foliar das amostras de serapilheira observamos que a relação entre o número de espécies e de folhas/folíolos, assim como a composição florística, depende da localidade, do habitat e da dispersão produzida pelo vento. Deste material foliar 60-95% pertence a 2-3 espécies arbóreas. A maior parte das espécies identificadas pertence às famílias Bignoniaceae, Euphorbiaceae, Fabaceae, Myrtaceae e Sapindaceae. Das espécies identificadas 55-70% são arbóreas e o 20-40% restante são trepadeiras, as quais representam 8-35% da riqueza total de espécies arbóreas e 5-32% da riqueza total de espécies de trepadeiras da área. As principais características fisionômicas do material foliar nos permitiram estimar a TMA e a precipitação média anual (PMA). Os valores da TMA obtidos subestimaram o valor real em 0.8-1.8 °C e o superestimaram em 1.5-4.8 °C, e os valores da PMA obtidos subestimaram o valor real em 11-188 mm e o superestimaram em 44-83 mm. As acumulações de macro-restos vegetais estudadas podem ser consideradas um registro de alta resolução de El Niño. O material foliar que está presente nestas acumulações depende da localidade, do habitat, e do transporte produzido pelo vento e pela água. Deste material foliar 70-85% pertence a 2-3 espécies arbóreas. A maior parte das espécies identificadas pertence às famílias Bignoniaceae, Fabaceae e Sapindaceae. Das espécies identificadas 70% são arbóreas e o 30% restante são trepadeiras, as quais representam 25-35% da riqueza total de espécies arbóreas e 23-32% da riqueza total de espécies de trepadeiras da área. As principais características fisionômicas deste material foliar permitiram estimar a TMA e a PMA para o momento de sua deposição. Os valores da TMA obtidos subestimaram o valor real em 0.5-1.3 °C e o superestimaram em 2.2 °C, e os valores da PMA obtidos subestimaram o valor real em 332-1070 mm. A análise da relação entre a proporção de espécies sem dentes e a TMA de um conjunto de 121 localidades da América do Sul nos permitiu gerar uma nova equação de regressão linear, baseada na Análise da Margem Foliar, mais apta para a reconstrução da TMA de floras fósseis cenozoicas da região Abstract: In this study, we analyzed the taxonomic composition and physiognomic features of the leaf material of seven samples of leaf litter of the riparian forest of Mogi Guaçu Ecological Station, Northeastern of the Sao Paulo State, Brazil, as well as five accumulations of plant macroremains that are found along meandering rivers of the Northeastern, Eastern and the South coastline of Sao Paulo State, with the purpose of obtaining new guidelines for the analysis and interpretation of the fossil associations angiosperms. We also analyzed the relationship in between the proportion of the untoothed species of a flora and the mean annual temperature (MAT) of a dataset of 121 localities the South America, in attempt to establishing a new equation based on the Leaf Margin Analysis for the whole region. When analyzing the foliar material from the leaf litter, we observed that the relationship in between the number of species and the number of leaves/leaflets, as well as the floristic composition, depends on the locality, habitat and wind-induced dispersion. The 60-95% of these leaf material belongs to 2-3 tree species. Most of the identified species belongs to the Bignoniaceae, Euphorbiaceae, Fabaceae, Myrtaceae and Sapindaceae families. The 55-70% of the identified species are from trees and the remaining 20-40% are climbing species. These species represent a 8-35% of all tree species richness and a 5-32% of all climbing species richness of the área. The main physiognomic features of the leaf material allowed us to estimate the MAT and the mean annual precipitation (MAP). The obtained MAT values underestimated the real value in 0.8-1.8 °C and overestimated in 1.5-4.8 °C, and the obtained MAP values underestimated the real value in 11-188 mm and overestimated in 44-83 mm. The accumulations of plant macroremains studied can be considered a high-resolution record of the El Niño. The leaf material that is present in these accumulations depends on the locality, habitat and transport produced by wind and water. The 70-85% of this leaf material belongs to 2-3 tree species. Most of them are identified as part of the Bignoniaceae, Fabaceae and Sapindaceae families. The 70% of the identified species are from tree and the remaining 30% are climbing species. These species represent a 25-35% of all tree species richness and a 23-32% of all climbing species richness of the área. The main physiognomic features of the leaf material allowed estimation of the MAT and MAP at the moment of deposition. The obtained AMT values underestimated the real value in 0.5-1.3 °C and overestimated the real value in 2.2 °C, and the obtained values of MAP underestimated the real value in 332-1070mm. The relationship analysis in between the proportion of the untoothed species and the MAT of a dataset of 121 localities the South America allowed us to create a new linear regression equation based on the Leaf Margin Analysis, more robust to reconstruct the MAT of Cenozoic fossil flora of the region Doutorado Geologia e Recursos Naturais Doutor em Ciências CAPES FAPESP 2010/20379-6, 2013/22729-2 FAEPEX

Research in the Domuraty 2 section was focused on a series of lacustrine-river-swamp deposits in which the full spectrum of vegetation and climate changes was recognised in a detailed analysis of plant macroremains and a comparison with the results of pollen analysis. Based on plant macrofossil data, two (Dom II, Dom III) of three palynologically documented warm units were distinguished in the Domuraty succession. The palaeobotanical data from the Domuraty succession document several successive local vegetation changes in both interglacial and glacial periods, which can be related to global climate oscillations. The succession of local vegetation stands and changes in the vegetation, climate, water level, and trophy during the period of the Domuraty succession were analysed and compared with corresponding parameters of the Augustovian and Ferdynandovian interglacials. The taxonomically most diverse unit is Warm unit Dom II, with the highest share of taxa with high thermal requirements, which was compared to the older Augustovian (A II) and younger Ferdynandovian (F II) units. The comparison of the Domuratovian interglacial flora to that of the Korchevian interglacial in Belarus shows high similarity; most of the extinct taxa or taxa unknown in the present-day flora are common to the Korchevian and Domuratovian floras, suggesting similar age for these two communities. However, the Domuratovian flora lacks a few species important to the Korchevian flora, such as Stratiotes goretskyi, Carex rostrata-pliocenica, Brasenia sp., Caulinia antiqua, Aldrovanda borysthenica, and A. zusii.

Early Middle Pleistocene palaeolacustrine sediments of the Ferdynandów site (E Poland), serving as a stratotype for the Ferdynandovian interglacial, were subjected to a new drilling in 2011. The obtained profile, covering the late Sanian 1 glaciation, two interglacial successions (Ferdynandovian 1 and 2), the cold Ferdynandovian 1/2 interval, and the early Sanian 2 glaciation, permitted a high-resolution plant macroremains analysis, correlated with the results of a palynological examination. In detailed studies of plant macroremains from the profile, new taxa were discovered for the Polish Pleistocene flora: specifically, species that are extinct or not found nowadays in Poland. On the basis of the taxonomic diversity and occurrence of bioindicators, the division into warm and cold units of the Ferdynandovian interglacial as well as units associated with the Sanian 1 and 2 glaciations, already applied in pollen studies, were confirmed, and the periods were described in terms of plant macroremains. The Ferdynandovian succession is correlated with the Cromerian Complex (Cromerian III and IV) in the Early Middle Pleistocene of Western Europe, as well as with Marine Isotope Stages (MIS) 13-15. New geological and palaeobotanical data enabled a description of the palaeogeographic context and conditions of functioning, as well as the evolution of the interglacial lake at the stratotype site. The outcome of plant macroremains analysis is presented and interpreted in comparison with corresponding results previously obtained for the Ferdynandów B profile, studied by Janczyk-Kopikowa (1975). Particular consideration is given to similarities between the succession of Ferdynandów 2011 and the profiles of Ferdynandów B and the nearby drilled Łuków 3A. Study of the macrofossil flora of Ferdynandów 2011 revealed the presence of species absent in the archival Ferdynandów B. Taxa not formerly recorded in early Middle Pleistocene floras, such as Brasenia borysthenica, Aldrovanda borysthenica, Pilularia borysthenica, Caulinia goretskyi, and Potamogeton saryanensis, were identified. These species and other taxa of stratigraphic or climatic significance provided the basis for a more detailed description of the Ferdynandovian flora, afterwards compared with other stratigraphically corresponding Polish and European early Middle Pleistocene floras. The composition of characteristic taxa also was used for comparisons between two bimodal interglacial sequences, namely the Ferdynandovian (correlated with MIS 13-15) and Augustovian (correlated with MIS 19-21), and the Domuratovian succession (correlated with MIS 17).

International audience; This paper presents the archaeobotanical study of several Neolithic settlements located in the Valais, the upper part of the Rhone Valley, in Switzerland. The archaeological sites are dated between 5000 and 4200 cal bc, which corresponds to the Early and the Middle Neolithic. Most of the sites are situated around 500–600 m, overhanging the alluvial plain of the Rhone. First results of the macroremains analysis show that cereals, comprising mainly naked wheat (Triticum aestivum s.l./durum/turgidum) and barley (Hordeum distichum/vulgare), are predominant. In addition, pea (Pisum sativum) and opium poppy (Papaver somniferum) are recorded. Gathered plants are sparse, and this from the first establishment of farmers around 5000 cal bc. If we compare the obtained results with data from the Northern French Alps, not far from the Valais, we get a completely different picture of the Neolithic plant economy. In the northern French Alps the communities exploited all vegetation levels from the collinean to the subalpine belt, gathered plants playing an important role alongside the cultivated species. Our first results are giving a new insight into the first agro-pastoral communities in the Swiss Alps; they are allowing us to understand how plant resources were exploited in a mountainous context and to outline the catchment area of the settlements.

In this thesis, plant exploitation activities in Atlantic Iberia during the Early and Middle Holocene have been approached through the study of non-woody plant macroremain assemblages at Mesolithic and Neolithic archaeological sites in the Cantabrian region and Portugal. Little direct information was previously available; this thesis therefore contributes to the enrichment of current knowledge about the Cantabrian region and offers completely unprecedented data for Portugal, critical for the understanding of past human culture and essential for the comprehension of the different Neolithisation processes. The new archaeobotanical data shed light on the dynamics of the exploitation of plant resources, particularly regarding the phenomenon of the transition from wild resource gathering to domestic agriculture. A wide spectrum of wild resources and the earliest crops in Portugal, with an extensive range of cereals and possibly legumes, have been attested. RESUMEN: En este trabajo se ha estudiado la explotación de los recursos vegetales en la costa atlántica ibérica durante el Holoceno Inicial y Medio, a partir del estudio de restos carpológicos de yacimientos arqueológicos mesolíticos y neolíticos de la región Cantábrica y Portugal. La información previa disponible era muy limitada; esta tesis contribuye al enriquecimiento del conocimiento sobre la región Cantábrica y ofrece datos hasta ahora únicos para Portugal, para la comprensión de las culturas del pasado y de los diferentes procesos de neolitización. Los nuevos datos arqueobotánicos arrojan luz sobre las dinámicas de explotación de los recursos vegetales por las sociedades del pasado, y en particular en el marco de la transición de la recolección de recursos silvestres a la agricultura de domésticos. Se ha documentado un amplio espectro de recursos silvestres, y los primeros cultivos de Portugal, formados por diversos cereales y posiblemente leguminosas. Esta tesis se divide en cinco partes; además del índice general, cada parte tiene su propio índice. En la primera parte (Part I: Framework) se revisa el marco general de este trabajo, considerando las cuestiones teóricas, medioambientales, sociales e históricas. En primer lugar, se revisan los conceptos clave y las explicaciones sobre los procesos de domesticación y neolitización. A continuación, se describe el marco físico en que se enmarca el problema histórico estudiado, así como sus características socioeconómicas, particularmente al respecto de la explotación de recursos vegetales. Después, presenta el estado de la cuestión por región y método. A continuación, se plantean los objetivos a las que se enfrenta el autor en este trabajo y finalmente, se presentan los contextos estudiados ordenados por región y cronología. Los restos materiales estudiados en este trabajo son un tipo de restos de plantas conocidos como carporrestos (frutos y semillas), con un potencial y unas limitaciones que se debaten de forma extensa en el capítulo metodológico sobre arqueobotánica (Part II: Methods), junto con otros métodos utilizados para la elaboración del estado de la cuestión. Los resultados del estudio material se presentan en la tercera parte: la primera sección (Part III A: Results: the identified taxa) presenta la descripción de las características de los taxones identificados en todos los sitios estudiados; la segunda parte (Part III B: Results: the studied contexts) incluye la presentación detallada de los conjuntos carpológicos contexto por contexto. En la cuarta parte (Part IV: Discussion and conclusions), se ponen en relación los datos obtenidos aquí con los datos disponibles hasta el momento de realización de este trabajo y se discute su contribución a completar el estado del conocimiento para los periodos y regiones estudiadas. Finalmente, se presentan las conclusiones. Finalmente (Appendixes and references), se cita la lista de referencias utilizadas durante el desarrollo de este trabajo, junto con los programas informáticos utilizados y las fuentes de material vegetal donado por distintas instituciones, que han sido cruciales en la creación de la nueva colección. La mayor parte de esta tesis (2009-2013) ha sido desarollada gracias a la concesión de una becacontrato por la Universidad de Cantabria. Esta tesis se enmarca en los siguientes proyectos de investigación financiados por el Ministerio de Economía y Competitividad: • Coordinado “La implantación de las especies domésticas en la Europa atlántica” (DOMATLAN) (HAR2008-06477-C03-00/HIST) (I.P.: Pablo Arias), subproyecto 1: “La implantación de las especies domésticas en la Europa atlántica: Cronología e impacto en la dieta humana” (DOMATLANTICA) (HAR2008-06477-C03-01/HIST; • “Coastal transitions: A comparative approach to the processes of neolithisation in Atlantic Europe ” (COASTTRAN) (HAR2011-29907-C03-01/HIST) (I.P.: Pablo Arias), subproyecto 1: “From shell middens to crop fields: The transition to the Neolithic in the coastal regions of SW Europe”; • “Coastal societies in a changing world: A diachronic and comparative approach to the Prehistory of SW Europe from the late Palaeolithic to the Neolithic" (COCHANGE) (HAR2014-51830-P) (I.P.: Pablo Arias).

The research focused on brackish marsh soils located within the coastal zone of Puck Lagoon (the Beka botanical and ornithological nature reserve). They are characterised by the presence of layers rich with allochthonous organic matter, derived from washing out of peat deposits. The objective of the research was to determine sedimentary environments of the soil organic matter based on selected properties. The obtained results indicated that despite similar morphological characteristics, these soils might have developed in slightly different environmental conditions, determined mostly by the Baltic water levels and morphology of the coast. The organic matter of the studied soils was characterised by relatively low aromaticity, which may indicate the ongoing process of decomposition. The presence of both peat-bog, rush and ruderal plants, as well as trees in the species composition of macroremains proved the heterogeneity of sedimentary environments and confirmed the allochthonous origin of organic matter accumulated in the studied brackish marsh soils.

Palaeobotanical investigations were carried out with the aim of reconstructing the development of palaeovegetation and formation of sediments in the northeastern area of ancient Lake Burtnieks. Pollen and plant macroremain studies provide information on vegetation development in the surroundings of the lake, including Stone Age settlements of Braukšas I and Braukšas II. Results of the investigations indicate that the development of vegetation together with sedimentation conditions in the palaeolake have changed since the Younger Dryas until today. Vegetation composition varies in different parts of the ancient Lake Burtnieks area due to past changes in lake water level which reached different sites at different times. Data from the northern part of ancient Lake Burtnieks indicate its gradual overgrowing since the Preboreal. Deposition of minerogenic lacustrine sediments (silt, clayey silt and sand) lasted until the Boreal or the Atlantic time, depending on the water depth of the lake locality. Clastic sediments were overlain by gyttja, which in turn was later covered by well-decomposed fen (sedge, sedge–grass) peat that started to form at the end of Atlantic time. Pollen and plant macroremain composition of lacustrine sediments and fen peat sequences suggests that people have inhabited the area since Preboreal–Boreal times. However, weak traces of possible presence of people are found already at the very end of the Younger Dryas.Fluctuating curves of broadleaved tree pollen, a significant amount of pollen of cultivated plants and charcoal dust in sediments indicate activities of an early man and refer to start of crop growing in the area in the second half of the Atlantic chronozone.

A long-lived Anatolian archaeological site (Arslantepe, Malatya plain) was investigated under a palaeoclimate perspective. Identified plant charred macroremains underwent an isotopic analysis to determine carbon isotope composition. The results obtained are used to assess the degree of climate changes in the site and its surroundings.

Stanowisko w Witowie, położone ok. 50 kilometrów na północny wschód od Krakowa, jest znane z odsłonięcia osadów określanych mianem serii witowskiej. Były one przedmiotem badań paleobotanicznych, paleozoologicznych, sedymentologicznych i petrograficznych. W osadach serii witowskiej znaleziono oznaczalne makroszczątki roślin iglastych i okrytozalążkowych. Szczególnie interesujące są kopalne gatunki Pinus cf. palaeostrobus, Zelkova zelkovifolia i Spirematospermum wetzleri. Wyniki badań szczątków roślinnych wskazują na późnomioceński lub plioceński wiek serii witowskiej. Kopalne zbiorowiska roślinne, które występowały w okolicy Witowa można porównać do współczesnych podzwrotnikowych lasów wilgotnych obszaru Kolchidy i Niziny Tałyskiej na Zakaukaziu oraz środkowych Chin. Obecność szczątków roślinności ciepłolubnej wskazuje na cieplejszy niż obecnie klimat okresu sedymentacji serii witowskiej. Witów jest znany także ze stanowisk archeologicznych. Obszar ten od neolitu był wielokrotnie zasiedlany. Ślady najintensywniejszego osadnictwa pochodzą z końca epoki brązu i początków epoki żelaza (1000-600 BC) i są odnoszone do kultury łużyckiej. Na stanowisku w Witowie znaleziono także średniowieczny cmentarz datowany na koniec jedenastego wieku po Chrystusie, będący pierwszym dowodem na obecność średniowiecznego osadnictwa na tym terenie. Położenie Witowa na brzegu doliny Wisły zapewne dawało mu kontrolę nad okolicznymi szlakami oraz, być może, także nad brodami. Możliwe, że Witów sprawował także polityczną kontrolę na otaczających go terenach. Biorąc pod uwagę naukową wartość stanowiska w Witowie i wynikającą stąd konieczność jego ochrony, postulowane jest utworzenie na jego obszarze rezerwatu przyrody. The study area near village Witów situated about 50 kilometres north-east of the Kraków city is famous for its exposed deposits called the "Witów Series" on which palaeobotanical, micro- and macrofaunistic, malacological, sedimentological and petrographical studies have already been performed. From deposits of the Witów Series, plant macroremains belonging to gymnosperms and angiosperms were reported. Pinus cf. palaeostrobus, Zelkova zelkovifolia and Spirematospermum wetzleri are particularly interesting. Results of investigations on fossil plant remains point to the Late Miocene to Pliocene age of the Witów Series. Plant communities that existed in the vicinity of Witów could be compared to recent warm temperate and humid forests of the Colchis and Talysh Lowland in Transcaucasus and to forests of Central China. Presence of remains of termophilous taxa point to warmer climate of the period of sedimentation of the Witów Series than recent climate of Poland. The Witów region is also famous for archaeological localities that has been known since the turn of the twentieth century. The site was being repeatedly settled beginning with the late Stone Age (the Neolithic Period). Traces of the most intensive periods of settlement come from the late Bronze and early Iron Ages (1000-600 BC) and are attributed to the so-called Lusatian culture. The site also yielded a medieval cemetery that can be dated to the turn of the twelfth century AD. This is the first confirmation of the presence of medieval settlements on this site. Location of Witów in the Vistula river valley may have allowed this settlement to control nearby routes and possibly river fords and also probably exercised political control over the surrounding areas. Considering scientific value of the Witów locality and necessity of their protection, creation of natural reserve on this area is suggested.