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Indoseges malkangiri sp. nov. Choudhury, Siliwal, Das & Giroti urn:lsid:zoobank.org:act: 5EB57046-CF2F-4E1E-890A-2E259273866A (Fig. 1–5) Type specimens: INDIA. holotype ♂, 25.v.2017, near Malkangiri town, Malkangiri district, Odisha (18° 21’ 46.3”N, 81° 54’ 43.8”E, elev. 235m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-554; paratypes 1 ♂ same data as holotype, IPU-17-ARA-555, 2 ♂ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda IPU-18-ARA-1272, IPU-18-ARA-1275, 2 ♀ same data as holotype, IPU-17-ARA-557, IPU-17-ARA-558, 3 ♀ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda, IPU-18-ARA-1274, IPU-18-ARA-1276, IPU-18-ARA-1277. Diagnosis: Males of I. malkangiri differs from I. sushildutta by having thorn-like macrosetae on the ventral side of the distal half of metatarsi I (in I. sushildutta cuspule-like macrosetae present) (Fig. 2 A,B); tibia Iwith a row of short macrosetae on the retrolateral side (in I. sushildutta on the ventro-retrolateral side) (Fig. 2 B, 7 A) and lacks any apophysis (in I. sushildutta, tibiae Iwith distinct pointed basal apophysis) (Fig. 2 A–C, 7 A–C); femur Iwith a cluster of stridulatory macrosetae but without any tubercle-like apophysis (in I. sushildutta, prominent tubercle-like femoral apophysis present covered with a cluster of stridulatory spines on the distal face) (Fig. 2 A–D, 7 A–C); leg formula 2143 (in I. sushildutta leg formula in male 1243); palpal tibiae distinctly incrassate and embolus at the base twists 90° along with distal bulb (in I. sushildutta, slightly incrassate and embolus at the base twists 110°) (Fig. 3 A–D, 8 A–D). Females of I. malkangiri differs from all other congers by having two prolateral macrosetae on metatarsi II (in all other species one prolateral macroseta), and a distinct internal genitalia: anterior receptaculum hornlike with a small spherical base and pointed apex; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold narrow, curved and strongly sclerotized forming a very conspicuous and well developed median flap; posterior receptaculum pear shaped (Fig. 5 A–E). Females also differ from rest of the species except I. satkosia by having three ventral macrosetae on tibiae IV (in all other species one ventral macroseta) and from I. sushildutta having leg formula 1423 (in I. sushildutta leg formula in female 1243). Etymology: The species epithet is a noun in apposition, referring to the type locality, Malkangiri in Odisha. Description of holotype male (IPU-17-ARA-554) (Fig. 1–3). Total length 7.50; carapace 3.89 long, 2.58 wide; abdomen 3.61 long, 2.28 wide; carapace covered with long and short brown hairs on caput and posterior margins; clypeus 0.25 high; eyes posterior row straight (Fig. 1 D); ocular group 0.27 long, 0.78 wide; eye diameter ALE 0.14, PME 0.14, PLE 0.14; inter-distances PLE-PME 0.13, ALE-ALE 0.28; abdomen ventrally pale greyish mottled with pale dots; chelicerae retrolaterally with two blunt tubercles at base; labium 0.90 long, 0.51 wide; endites 1.41 long, 0.58 wide; sternum 2.25 long, 1.45 wide, anteriorly marked by thick pale band (Fig. 1 B); legs metatarsi Iin ventral distal half with a thorn-like macroseta; tibiae Iwith a row of short macrosetae on retrolateral side (Fig. 2 B) and lacks any apophysis (Fig. 2 A–C); femur Iwith a cluster of stridulatory macrosetae and lacks prominent tuberclelike apophysis (Fig. 2 A–D); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 3.00, 1.12, 2.61, 2.55, 1.31, 10.59; leg II: 3.21, 1.09, 3.86, 2.78, 1.27, 12.21; leg III: 2.36, 0.86, 1.51, 2.14, 0.82, 7.69; leg IV: 2.98, 1.18, 2.56, 2.51, 1.01, 10.24; palp: 1.25, 0.58, 0.95, -, 0.71, 3.49; leg formula 2143; spines or macrosetae: leg I: fep 16, tip 1-1-1-1, r 1-1-1-1, vp 1-1-1-1, vr 1-1-1-1-1-1-1, mtv 2-0-2; leg II: fep 1-1, tip 1-1-1, r 1-1-1-1, vp 0-0-1, vr 1-1-1, mtvp 1-0-1, vr 1-1-1; leg III: fep 1, tip 1-1-1, r 1-0-1, v 2-1-1-2, mtp 1-0-1, r 1-1-1, v 1-1-3; leg IV: fe d 1-1, ti v 1-1-2, mt vp 1-1-1-2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: no spine; teeth on paired claws of legs: I: 10, II: 8, III: 7, IV: 6; palp brownish-black; patella and tibia distinctly incrassate; cymbium; bulb pear shaped; embolic duct distinct; embolus at the base twists 90° along with distal bulb; embolus long, tubular, tapers and gradually bends dorsally, and spatulated distally with embolic opening on one side (Fig. 3 A–D). Remaining characters same as generic character. Variation in males [n=4; measurements: range (mean±standard deviation)]. Total length 6.88–7.50 (7.26±0.28); carapace. 3.32–3.89 (3.63±0.24) long, 2.09–2.58 (2.28±0.22) wide; abdomen 3.51–3.83 (3.63±0.14) long, 2.01–2.31 (2.21±0.14) wide; clypeus 0.18–0.28 (0.23±0.05) high; ocular group 0.25–0.31 (0.29±0.02) long, 0.61–0.78 (0.69±0.07) wide; eye diameter ALE 0.12–0.19 (0.15±0.03), PME 0.12–0.15 (0.14±0.01), PLE 0.12–0.19 (0.15±0.03); inter-distances PLE-PME 0.08–0.15 (0.12±0.03), ALE-ALE 0.25–0.28 (0.26±0.01); labium 0.63–0.90 (0.78±0.13) long, 0.44–0.51 (0.48±0.04) wide; endites 1.16–1.41 (1.29±0.12) long, 0.43–0.58 (0.51±0.07) wide; sternum 2.00–2.25 (2.13±0.11) long, 1.22–1.45 (1.33±0.12) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.42–3.00 (2.77±0.25), 0.77–1.18 (1.00±0.19), 2.12–2.61 (2.45±0.23), 2.02–2.55 (2.37±0.24), 1.10–1.34 (1.25±0.11), 8.43–10.59 (9.84±0.98); leg II: 2.62–3.21 (2.95±0.25), 1.01–1.19 (1.10±0.07), 2.43–3.86 (2.43±0.62), 2.61–2.88 (2.77±0.11), 1.19–1.34 (1.26±0.07), 9.86–12.21 (11.05±0.98); leg III: 2.02–2.36 (2.21±0.14), 0.83–0.87 (0.85±0.02), 1.24–1.51 (1.41±0.13), 1.80–2.14 (2.00±0.14), 0.81–0.88 (0.83±0.03), 6.70–7.69 (7.30±0.43); leg IV: 2.55–2.98 (2.97±0.01), 1.04–1.25 (1.22±0.05), 2.06–2.56 (2.43±0.19), 2.18–2.62 (2.57±0.08), 0.79–1.01 (0.96±0.08), 8.62–10.24 (9.61±0.72); palp: 1.07–1.25 (1.14±0.08), 0.54–0.58 (0.56±0.02), 0.87–0.95 (0.92±0.03), -, 0.60–0.71 (0.67±0.05), 3.15–3.49 (3.28±0.15). Description of paratype female (IPU-18-ARA-1274) (Fig. 4, 5). Total length 10.96; carapace.4.55 long, 2.75 wide; abdomen 6.41 long, 3.62 wide; carapace covered with long and short brown hairs, relatively more on caput; clypeus 0.30 high; eyes posterior row straight (Fig. 4 D); ocular group 0.29 long, 0.88 wide; eye diameter ALE 0.18, PME 0.15, PLE 0.18; inter-distances PLE-PME 0.18, ALE-ALE 0.46; abdomen ventrally pale greyish mottled with pale dots; labium 0.77 long, 0.58 wide; endites 1.45 long, 0.67 wide; sternum 2.64 long, 1.66 wide, anteriorly marked by a thick pale band (Fig. 4 B); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.87, 1.23, 2.10, 1.91, 0.99, 9.10; leg II: 2.74, 1.21, 2.14, 1.95, 1.00, 9.04; leg III: 2.17, 1.00, 1.21, 1.54, 0.72, 6.64; leg IV: 2.88, 1.31, 2.23, 1.93, 0.72, 9.08; palp: 1.10, 0.65, 0.59, -, 0.92, 3.26; leg formula 1423; macrosetae: leg I: fep 1, tip 1-1-1, v 2-2-2, mtvp 1-0-1, vr 1-1-1; leg II: fep 1, tip 1-1-1, vp 1-0-1, vr 0-0-1, mtp 1-1-0, v 2-1-1-2; leg III: ti p 1-1, v 1-2, r 1-1, mt p 1-1-1-1, r 1-1-1-1, v 1-1-1-3; leg IV: ti vp 1-1, vr 0-1, mt vp 1-1- 2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 1-1, ta p 13; teeth on paired claws of legs: I: 15, II: 13, III: 11, IV: 9; female genitalia distinct; internal genitalia with a horn-like anterior receptaculum having small spherical base and pointed apex; dorsal lobe spherical; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold narrow, curved and strongly sclerotized forming a very conspicuous and well developed median flap; pear shaped posterior receptaculum (Fig. 5 A–E). Remaining characters same as generic character. Variation in females [n=5, measurements: range (mean±standard deviation)]. Total length 8.61–12.80 (10.57±1.61); carapace. 3.69–4.55 (4.17±0.31) long, 2.5–2.75 (2.57±0.13) wide; abdomen 4.92–8.58 (6.40±1.43) long, 2.93–3.62 (3.29±0.26) wide; clypeus 0.24–0.40 (0.32±0.08) high; ocular group 0.25–0.34 (0.30±0.03) long, 0.79–0.94 (0.83±0.10) wide; eye diameter ALE 0.14–0.18 (0.15±0.02), PME 0.13–0.18 (0.15±0.02), PLE 0.14–0.18 (0.15±0.02), inter-distances PLE-PME 0.12–0.18 (0.16±0.03), ALE-ALE 0.36–0.46 (0.39±0.04); labium 0.72–0.78 (0.74±0.03) long, 0.54–0.59 (0.57±0.02) wide; endites 1.26–1.45 (1.35±0.08) long, 0.53–0.67 (0.61±0.05) wide; sternum 2.27–2.64 (2.42±0.16) long, 1.44–1.69 (1.55±0.12) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.56–2.92 (2.75±0.14), 1.08–1.23 (1.15±0.06), 1.94–2.19 (2.08±0.10), 1.79–2.03 (1.94±0.09), 0.97–1.04 (0.99±0.03), 8.85–9.30 (8.92±0.36); leg II: 2.34–2.81 (2.65±0.19), 1.17–1.27 (1.22±0.04), 1.95–2.15 (2.131±0.11), 1.85–2.08 (2.00±0.10), 0.96–1.02 (0.97±0.05), 9.03–9.37 (8.98±0.42); leg III: 1.91–2.22 (2.08±0.12), 0.98–1.02 (0.98±0.06), 1.07–1.22 (1.17±0.06), 1.38–1.54 (1.49±0.07), 0.65–0.73 (0.69±0.03), 6.00–6.72 (6.41±0.30); leg IV: 2.66–2.97 (2.82±0.12), 1.28–1.31 (1.31±0.05), 2.02–2.34 (2.20±0.12), 1.76–1.93 (1.87±0.09), 0.66–0.77 (0.71±0.05), 8.38–9.44 (8.91±0.41); palp: 0.89–1.10 (1.04±0.09), 0.56–0.65 (0.60±0.04), 0.53–0.59 (0.56±0.03), -, 0.78–1.01 (0.90±0.09), 2.76–3.26 (3.09±0.21); teeth on paired claws of legs: I: 14–15, II: 12–13., Published as part of Choudhury, Sudhir Ranjan, Siliwal, Manju, Das, Sanjay Keshari & Giroti, André Marsola, 2021, Description of a new genus and five new species of tube-dwelling spider family Segestriidae (Araneae: Synspermiata) from Odisha, India, pp. 91-114 in Zootaxa 4963 (1) on pages 94-98, DOI: 10.11646/zootaxa.4963.1.4, http://zenodo.org/record/4719875

Indoseges gen. nov. Siliwal, Das, Choudhury & Giroti urn:lsid:zoobank.org:act: 4A00F387-0EBB-4B58-A2DA-66056B230B2F Diagnosis: Indoseges gen. nov. resembles Ariadninae genera, but differs from Segestriinae genera Gippsicola Hogg, 1900 and Segestria Latreille, 1804 by having labium narrowed distally (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), posterior eyes in a straight line or slightly recurved (Fig. 1 D, 4 D, 6 D, 9 D, 11 D, 13 D, 15 C), lateral basal transverse ridge present on chelicerae, and chelicerae with three promarginal and one retromarginal teeth (Fig. 6 E, 11 E, 13 E, 15 D) (Beatty, 1970; Grismado, 2008; Ramírez, 2014; Giroti & Brescovit, 2015,2017,2018; Marsh et al. 2018; Fomichev & Marusik, 2020). Among Ariadninae, it differs from both Ariadna and Citharoceps by having: male tarsi I–II incrassate and with ventral scopulae (Fig. 2 F,G), presence of 4–5 ventral spines on metatarsi Iof males, and female femur I–II with one macrosetae. Additional characters that separates it from Ariadna: labrum as long as labium and distally with a row of dorsal median microsetae (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), but in Ariadna labrum longer than the labium and without dorsal median microsetae; males lack apophysis on metatarsi I (Fig. 2 A,B), but in Ariadna apophysis may or may not present on metatarsi Iof males; males with a cluster of stridulatory macrosetae on femur I, which could be on tubercle-like apophysis (Fig. 2 A–D, 7 A–C), but in Ariadna a cluster of stridulatory macrosetae may or may not present on femur Iof males and apophysis absent; female palp tarsi with more than ten macrosetae (Fig. 4 F, 9 F, 11 G, 13 G, 15 E), but in Ariadna number of macrosetae five or more; and presence of conspicuous median flap in female genitalia (Fig. 5 C,E, 10 C,E, 12 B,D, 14 B,D, 16 B,D), but absent in Ariadna. Additional characters that separates it from Citharoceps: stridulatory apparatus in the cephalic area absent, but present in Citharoceps; a stridulatory thorn in the prolateral region of femur I absent, but present in Citharoceps; presence of tubular invaginations on posterior receptaculum in females (Fig. 5 B–E, 10 B–E, 12 A–C, 14 A–C, 16 A–D), but absent in Citharoceps; and labium-sternum junction length is greater than the endite-sternum junction; but in Citharoceps labium-sternum junction is equal or smaller than the length of endite-sternum junction (Beatty, 1970; Grismado, 2008; Ramírez, 2014; Giroti & Brescovit, 2015, 2018; Siliwal et al. 2017; Marsh et al. 2018). Description (Fig. 1–17): Colouration: Spider in life blackish-brown with glossy integument (17 A,B), overall males darker than females. In ethanol, carapace greenish-brown (thoracic area) to brownish-black (cephalic area) (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A); median eyes translucent and lateral eyes pale, all eyes with black rim (Fig. 4 D, 6D, 9D); legs and palp orangish-green except brownish-black femora and greenish-brown tibia, few small black patches on patella and tibia (Fig. 1 A, 3, 4 A, 6 A, 8, 11 A, 13 A, 15 A), ventrally pale greenish-brown (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); abdomen dorsally greyish-brown with few pale patches (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A), ventrally pale greyish mottled with pale dots or without any pattern, two orange spots and black patches on epigastric area (Fig. 5 A, 10 A); labium orangish-brown, lighter anteriorly (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); endites orangish-brown except for white proximal end (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); sternum orangish-brown with faint black patches laterally and medially (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); chelicerae brownish-black (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B). Morphometry and setae (male, female): Total length 6.82–7.50 (5.09–12.80), carapace 3.32–3.89 long, 2.09–2.58 wide (2.48–4.89 long, 0.99–2.82 wide), abdomen 3.50–3.83 long, 2.00–2.31 wide (2.61–8.58 long, 1.67–4.33 wide). Carapace: longer than broad, narrowed anteriorly; cephalic area higher than thoracic area; integument rough, covered with long and short brown hairs; caput slightly raised; thoracic fovea indistinct (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A). Eyes: six, grouped in three pairs, anterior median eyes absent, lateral eyes placed on a single tubercle, posterior row straight or slightly recurved (Fig. 1 D, 4 D, 6 D, 9 D, 11 D, 13 D, 15 C). Chelicerae: conical with rough integument due to files or hairs; having distinct retrolateral basal transverse ridge; covered with stiff hairs with tuberculated base except for posterior and basal lateral sides; prominent depression on the posterior distal region to fit distal part of endites; fangs short, shaft with an internal serrula and about 2.5 times longer than wide, with three promarginal teeth and one retromarginal tooth (Fig. 6 E, 11 E, 13 E, 15 D). Labium: longer than wide, narrowed down in anterior one-third, separated from sternum by partially membranous suture, covered with few long and short black hairs (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B). Labrum: well developed and is about length of the labium (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), distal end having a row of median microsetae. Endites: broader at the base, narrowed medially and rounded proximally, covered with long black hairs (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), dorso-distally depressed to fit cheliceral distal end, serrula with a single row of fine minute teeth forming a ridge-like structure. Sternum: longer than wide, margins with precoxal triangles, concave anteriorly, rebordered and not fused with carapace or pedicel, covered with long black hairs (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B). Abdomen: longer than wide, uniformly hairy (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A), coated with short setae, and scattered long setae among them; book lung and tracheal opening weakly sclerotized (Fig. 5 A, 10 A). Spinnerets: three pairs, covered with brown hairs; anterior lateral spinnerets with three segments, the basal segment crossed by a diagonal membranous stripe, colulus distinct, short and covered with long hairs (Fig. 1 C, 4 C, 6 C, 9 C, 11 C, 13 C). Legs: robust, prograde, with third pair directed forward; each leg with three tarsal claws, two paired claws with long dentitions (Fig. 4 E, 9 E, 11 F, 13 F), teeth largest on leg I, unpaired claw with one small tooth (Fig. 4 E, 9 E, 11 F, 13 F), true claw tufts absent; leg pairs I–II with more setae and macrosetae than pairs III–IV; metatarsi and tarsi IV with ventral brush of thickened and serrated setae and metatarsi IV with preening comb on distal ventro or ventro-retrolateral (Fig.7 D); trichoboth- ria on dorsal subdistal region of metatarsi I–IV and dorsal basal half of tibia I–IV; male tarsi I–II incrassate and with ventral scopulae or tenant setae (Fig. 2 F,G); in males femur, tibiae and metatarsi of all legs with macrosetae; male metatarsi Ilacks any apophysis (Fig. 2 A,B) and with thorn-like or cuspule-like macrosetae on ventral side of distal half; males with or without distinct pointed basal tibial apophysis on prolateral side of tibiae I (Fig. 2 A–C, Fig. 7 A–C), male tibiae Ialso with a row of short macrosetae on retrolateral side or ventro-retrolateral side (Fig. 2 B, 7 A); males with a cluster of stridulatory macrosetae on femur I, that could be on tubercle-like apophysis (Fig. 2 A–D, 7 A–C); in females tarsi I–II with ventro-prolateral lines of thickened tactile setae, interspersed with lines of chemosensory setae; females tibiae and metatarsi of all legs with macrosetae, but only ventro-prolateral, ventroretrolateral, and sometimes prolateral and retrolateral; female femur I–II with one macroseta. Leg formula in male 1243 or 2143 and female 1243 or 1423. Palp: tibiae of male palp incrassate/swollen basally slightly to distinctly, longer than cymbium, with one dorsal trichobothrium and without any macrosetae; cymbium with distal notch dividing it into two unequal short processes, prolateral portion slightly longer than retrolateral one and without any scopulae or sensory setae; bulb pear-shaped, abruptly narrowed in mid-portion between bulb and embolus, embolus long, tubular, tapers and gradually bends dorsally, twists at the base 90° or 110° along with distal bulb and spatulated distally with an embolic opening on one side; spermatic duct broad, emerging at the base of bulb, coiling gradually towards mid-portion and enters embolus (Fig. 3 A–D, 8 A–D); female palp short and robust, with one smooth claw, turned inward almost 90°; macrosetae present on prolateral sides of tibiae and tarsi, tarsi with more than ten macrosetae (Fig. 4 F, 9 F, 11 G, 13 G, 15 E); tibiae with a single dorsal trichobothrium. Female genitalia: epigynal plate distinct and sclerotized; internal genitalia composed by anterior receptaculum, uterus externus, interpulmonary fold and posterior receptaculum; anterior receptaculum divided into dorsal and ventral lobe; dorsal lobe small, weakly sclerotized and transparent; ventral lobe strongly sclerotized and with multiple elongated entapophyses; glandular duct plate sclerotized and located in central area at the junction of both lobes; uterus externus indistinct or weakly sclerotized and transparent; interpulmonary fold strongly sclerotized forming a median flap; posterior receptaculum sac-like or shape variable, and with small tubular invaginations (Fig. 5 A–E, 10 A–E, 12 A–D, 14 A–D, 16 A–D). Etymology: The name is a combination of two words, ‘Indo’ and ‘seges’, Indo, relates to India from where the new genus is discovered and seges, refers to ‘Segestriidae’, the family name to which the new genus belongs. Type species: Indoseges malkangiri sp. nov., based on male and female specimens deposited at Indraprastha University Museum (IPUM), New Delhi. Distribution: Odisha, India, Published as part of Choudhury, Sudhir Ranjan, Siliwal, Manju, Das, Sanjay Keshari & Giroti, André Marsola, 2021, Description of a new genus and five new species of tube-dwelling spider family Segestriidae (Araneae: Synspermiata) from Odisha, India, pp. 91-114 in Zootaxa 4963 (1) on pages 92-94, DOI: 10.11646/zootaxa.4963.1.4, http://zenodo.org/record/4719875, {"references":["Beatty, J. (1970) The spider genus Ariadna in the Americas (Araneae, Dysderidae). Bulletin of the Museum of Comparative Zoology, 139 (8), 433 - 518.","Grismado, C. J. (2008) A taxonomic revision of the spider genus Ariadna Audouin, 1826 in Argentina and Chile, with the description of five new species (Arachnida, Araneae, Segestriidae). Zoosystema, 30, 333 - 360.","Ramirez, M. J. (2014) The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, (390), 1 - 374. https: // doi. org / 10.1206 / 821.1","Marsh, J. R., Baehr, B. C., Glatz, R. V. & Framenau, V. W. (2018) New species of tube web spiders of the genus Ariadna from South Australia (Araneae, Segestriidae). Evolutionary Systematics, 2, 137 - 149. https: // doi. org / 10.3897 / evolsyst. 2.29172","Fomichev, A. A. & Marusik, Y. M. (2020) Notes on the spider genus Segestria Latreille, 1804 (Araneae: Segestriidae) in the East Palaearctic with description of three new species. Zootaxa, 4758 (2), 330 - 346. https: // doi. org / 10.11646 / zootaxa. 4758.2.7","Giroti, A. M. & Brescovit, A. D. (2015) Revalidation of the spider genus Citharoceps Chamberlin, 1924 (Araneae, Segestriidae). ZooKeys, 495, 1 - 19. https: // doi. org / 10.3897 / zookeys. 495.8950","Giroti, A. M. & Brescovit, A. D. (2018) The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae). Zootaxa, 4400 (1), 1 - 114. [& Erratum: vol. 4438 (no. 3): 600] https: // doi. org / 10.11646 / zootaxa. 4400.1.1","Siliwal, M., Yadav, A. & Kumar, D. (2017) Three new species of tube-dwelling spider genus Ariadna Audouin, 1826 (Araneae: Segestriidae) from India. Zootaxa, 4362 (3), 433 - 441. https: // doi. org / 10.11646 / zootaxa. 4362.3.7"]}

Indoseges gen. nov. Siliwal, Das, Choudhury & Giroti urn:lsid:zoobank.org:act: 4A00F387-0EBB-4B58-A2DA-66056B230B2F Diagnosis: Indoseges gen. nov. resembles Ariadninae genera, but differs from Segestriinae genera Gippsicola Hogg, 1900 and Segestria Latreille, 1804 by having labium narrowed distally (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), posterior eyes in a straight line or slightly recurved (Fig. 1 D, 4 D, 6 D, 9 D, 11 D, 13 D, 15 C), lateral basal transverse ridge present on chelicerae, and chelicerae with three promarginal and one retromarginal teeth (Fig. 6 E, 11 E, 13 E, 15 D) (Beatty, 1970; Grismado, 2008; Ramírez, 2014; Giroti & Brescovit, 2015,2017,2018; Marsh et al. 2018; Fomichev & Marusik, 2020). Among Ariadninae, it differs from both Ariadna and Citharoceps by having: male tarsi I–II incrassate and with ventral scopulae (Fig. 2 F,G), presence of 4–5 ventral spines on metatarsi Iof males, and female femur I–II with one macrosetae. Additional characters that separates it from Ariadna: labrum as long as labium and distally with a row of dorsal median microsetae (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), but in Ariadna labrum longer than the labium and without dorsal median microsetae; males lack apophysis on metatarsi I (Fig. 2 A,B), but in Ariadna apophysis may or may not present on metatarsi Iof males; males with a cluster of stridulatory macrosetae on femur I, which could be on tubercle-like apophysis (Fig. 2 A–D, 7 A–C), but in Ariadna a cluster of stridulatory macrosetae may or may not present on femur Iof males and apophysis absent; female palp tarsi with more than ten macrosetae (Fig. 4 F, 9 F, 11 G, 13 G, 15 E), but in Ariadna number of macrosetae five or more; and presence of conspicuous median flap in female genitalia (Fig. 5 C,E, 10 C,E, 12 B,D, 14 B,D, 16 B,D), but absent in Ariadna. Additional characters that separates it from Citharoceps: stridulatory apparatus in the cephalic area absent, but present in Citharoceps; a stridulatory thorn in the prolateral region of femur I absent, but present in Citharoceps; presence of tubular invaginations on posterior receptaculum in females (Fig. 5 B–E, 10 B–E, 12 A–C, 14 A–C, 16 A–D), but absent in Citharoceps; and labium-sternum junction length is greater than the endite-sternum junction; but in Citharoceps labium-sternum junction is equal or smaller than the length of endite-sternum junction (Beatty, 1970; Grismado, 2008; Ramírez, 2014; Giroti & Brescovit, 2015, 2018; Siliwal et al. 2017; Marsh et al. 2018). Description (Fig. 1–17): Colouration: Spider in life blackish-brown with glossy integument (17 A,B), overall males darker than females. In ethanol, carapace greenish-brown (thoracic area) to brownish-black (cephalic area) (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A); median eyes translucent and lateral eyes pale, all eyes with black rim (Fig. 4 D, 6D, 9D); legs and palp orangish-green except brownish-black femora and greenish-brown tibia, few small black patches on patella and tibia (Fig. 1 A, 3, 4 A, 6 A, 8, 11 A, 13 A, 15 A), ventrally pale greenish-brown (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); abdomen dorsally greyish-brown with few pale patches (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A), ventrally pale greyish mottled with pale dots or without any pattern, two orange spots and black patches on epigastric area (Fig. 5 A, 10 A); labium orangish-brown, lighter anteriorly (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); endites orangish-brown except for white proximal end (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); sternum orangish-brown with faint black patches laterally and medially (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B); chelicerae brownish-black (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B). Morphometry and setae (male, female): Total length 6.82–7.50 (5.09–12.80), carapace 3.32–3.89 long, 2.09–2.58 wide (2.48–4.89 long, 0.99–2.82 wide), abdomen 3.50–3.83 long, 2.00–2.31 wide (2.61–8.58 long, 1.67–4.33 wide). Carapace: longer than broad, narrowed anteriorly; cephalic area higher than thoracic area; integument rough, covered with long and short brown hairs; caput slightly raised; thoracic fovea indistinct (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A). Eyes: six, grouped in three pairs, anterior median eyes absent, lateral eyes placed on a single tubercle, posterior row straight or slightly recurved (Fig. 1 D, 4 D, 6 D, 9 D, 11 D, 13 D, 15 C). Chelicerae: conical with rough integument due to files or hairs; having distinct retrolateral basal transverse ridge; covered with stiff hairs with tuberculated base except for posterior and basal lateral sides; prominent depression on the posterior distal region to fit distal part of endites; fangs short, shaft with an internal serrula and about 2.5 times longer than wide, with three promarginal teeth and one retromarginal tooth (Fig. 6 E, 11 E, 13 E, 15 D). Labium: longer than wide, narrowed down in anterior one-third, separated from sternum by partially membranous suture, covered with few long and short black hairs (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B). Labrum: well developed and is about length of the labium (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), distal end having a row of median microsetae. Endites: broader at the base, narrowed medially and rounded proximally, covered with long black hairs (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B), dorso-distally depressed to fit cheliceral distal end, serrula with a single row of fine minute teeth forming a ridge-like structure. Sternum: longer than wide, margins with precoxal triangles, concave anteriorly, rebordered and not fused with carapace or pedicel, covered with long black hairs (Fig. 1 B, 4 B, 6 B, 9 B, 11 B, 13 B, 15 B). Abdomen: longer than wide, uniformly hairy (Fig. 1 A, 4 A, 6 A, 9 A, 11 A, 13 A, 15 A), coated with short setae, and scattered long setae among them; book lung and tracheal opening weakly sclerotized (Fig. 5 A, 10 A). Spinnerets: three pairs, covered with brown hairs; anterior lateral spinnerets with three segments, the basal segment crossed by a diagonal membranous stripe, colulus distinct, short and covered with long hairs (Fig. 1 C, 4 C, 6 C, 9 C, 11 C, 13 C). Legs: robust, prograde, with third pair directed forward; each leg with three tarsal claws, two paired claws with long dentitions (Fig. 4 E, 9 E, 11 F, 13 F), teeth largest on leg I, unpaired claw with one small tooth (Fig. 4 E, 9 E, 11 F, 13 F), true claw tufts absent; leg pairs I–II with more setae and macrosetae than pairs III–IV; metatarsi and tarsi IV with ventral brush of thickened and serrated setae and metatarsi IV with preening comb on distal ventro or ventro-retrolateral (Fig.7 D); trichoboth- ria on dorsal subdistal region of metatarsi I–IV and dorsal basal half of tibia I–IV; male tarsi I–II incrassate and with ventral scopulae or tenant setae (Fig. 2 F,G); in males femur, tibiae and metatarsi of all legs with macrosetae; male metatarsi Ilacks any apophysis (Fig. 2 A,B) and with thorn-like or cuspule-like macrosetae on ventral side of distal half; males with or without distinct pointed basal tibial apophysis on prolateral side of tibiae I (Fig. 2 A–C, Fig. 7 A–C), male tibiae Ialso with a row of short macrosetae on retrolateral side or ventro-retrolateral side (Fig. 2 B, 7 A); males with a cluster of stridulatory macrosetae on femur I, that could be on tubercle-like apophysis (Fig. 2 A–D, 7 A–C); in females tarsi I–II with ventro-prolateral lines of thickened tactile setae, interspersed with lines of chemosensory setae; females tibiae and metatarsi of all legs with macrosetae, but only ventro-prolateral, ventroretrolateral, and sometimes prolateral and retrolateral; female femur I–II with one macroseta. Leg formula in male 1243 or 2143 and female 1243 or 1423. Palp: tibiae of male palp incrassate/swollen basally slightly to distinctly, longer than cymbium, with one dorsal trichobothrium and without any macrosetae; cymbium with distal notch dividing it into two unequal short processes, prolateral portion slightly longer than retrolateral one and without any scopulae or sensory setae; bulb pear-shaped, abruptly narrowed in mid-portion between bulb and embolus, embolus long, tubular, tapers and gradually bends dorsally, twists at the base 90° or 110° along with distal bulb and spatulated distally with an embolic opening on one side; spermatic duct broad, emerging at the base of bulb, coiling gradually towards mid-portion and enters embolus (Fig. 3 A–D, 8 A–D); female palp short and robust, with one smooth claw, turned inward almost 90°; macrosetae present on prolateral sides of tibiae and tarsi, tarsi with more than ten macrosetae (Fig. 4 F, 9 F, 11 G, 13 G, 15 E); tibiae with a single dorsal trichobothrium. Female genitalia: epigynal plate distinct and sclerotized; internal genitalia composed by anterior receptaculum, uterus externus, interpulmonary fold and posterior receptaculum; anterior receptaculum divided into dorsal and ventral lobe; dorsal lobe small, weakly sclerotized and transparent; ventral lobe strongly sclerotized and with multiple elongated entapophyses; glandular duct plate sclerotized and located in central area at the junction of both lobes; uterus externus indistinct or weakly sclerotized and transparent; interpulmonary fold strongly sclerotized forming a median flap; posterior receptaculum sac-like or shape variable, and with small tubular invaginations (Fig. 5 A–E, 10 A–E, 12 A–D, 14 A–D, 16 A–D). Etymology: The name is a combination of two words, ‘Indo’ and ‘seges’, Indo, relates to India from where the new genus is discovered and seges, refers to ‘Segestriidae’, the family name to which the new genus belongs. Type species: Indoseges malkangiri sp. nov., based on male and female specimens deposited at Indraprastha University Museum (IPUM), New Delhi. Distribution: Odisha, India

Indoseges sushildutta sp. nov. Siliwal, Das, Choudhury, & Giroti urn:lsid:zoobank.org:act: 333AD004-862B-4F37-B27D-121C3E12AAFD (Fig. 6–10) Type specimens: INDIA- holotype ♂, 25.v.2017, Govindpali ghati, Malkangiri district, Odisha (18° 36’ 34.3”N, 82° 18’ 12.3”E, elev. 248m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-560; paratypes 1 ♂ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda, IPU-18-ARA-1286, 4 ♀ same data as holotype, IPU- 17-ARA-548, IPU-17-ARA-549, IPU-17-ARA-550, IPU-17-ARA-559, 3 ♀ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda, IPU-18-ARA-1281, IPU-18-ARA-1287, IPU-18-ARA-1290. Diagnosis: Males of I. sushildutta differs from I. malkangiri by having capsule-like macrosetae on the ventral side of the distal half of metatarsi I (in I. malkangiri thorn-like macrosetae present); tibiae Iwith a row of short macrosetae on the ventro-retrolateral side (in I. malkangiri on the retrolateral side) (Fig. 2 B, 7 A) and a distinct pointed prolateral basal apophysis (in I. malkangiri, tibiae Ilacks any apophysis) (Fig. 2 A–C, 7 A–C); femur Iwith a distinct triangular tubercle-like prolateral femoral apophysis covered by a cluster of stridulatory macrosetae on the distal face (in I. malkangiri, femur Iwith a cluster of stridulatory macrosetae and lacks any apophysis) (Fig. 2 A–D, 7 A–C); leg formula 1243 (in I. malkangiri leg formula in male 2143); palpal tibiae slightly incrassate and embolus at the base twists 110° along with distal bulb (in I. malkangiri, distinctly incrassate and embolus at the base twists 90°) (Fig. 3 A–D, 8 A–D). Females of I. sushildutta differs from all other congers by having leg formula of 1243 (in all other species it is 1423), four ventral macrosetae on metatarsi I (in all other species five ventral macrosetae), and a distinct internal genitalia: anterior receptaculum horn-like with a broad, elongated base and gradually tapered apex; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide, nearly straight, strongly sclerotized forming a conspicuous, but short median flap; posterior receptaculum with irregular edges (Fig. 10 A–E). Etymology: The species epithet is a noun in apposition, in honor of renowned herpetologist, Prof. Sushil Kumar Dutta for encouraging and supporting our studies on spiders in Odisha. Description of holotype male (IPU-17-ARA-560) (Fig. 6–8). Total length 7.37; carapace.3.87 long, 2.54 wide; abdomen 3.50 long, 3.20 wide; clypeus 0.23 high; ocular group 0.31 long, 0.81 wide; eye diameter ALE 0.17, PME 0.17, PLE 0.17; inter-distances PLE-PME 0.11, ALE-ALE 0.33; chelicerae retrolaterally with one blunt tubercle at base; labium 0.82 long, 0.53 wide; endites 1.44 long, 0.52 wide; sternum 2.30 long, 1.44 wide; metatarsi Iwith a single row of small cuspule like ventral macrosetae on distal half; tibiae Iwith a row of short macrosetae on ventro-retrolateral side (Fig. 7 A), and a distinct pointed basal prolateral apophysis (Fig. 7 A–C); femur Iwith a distinct triangular tubercle-like prolateral femoral apophysis covered by a cluster of stridulatory spines on distal face (Fig. 7 A–C); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.93, 1.00, 3.40, 2.64, 1.34, 11.31; leg II: 2.85, 1.11, 2.73, 2.56, 1.18, 10.43; leg III: 2.14, 0.87, 1.37, 1.74, 0.88, 7.00; leg IV: 2.70, 1.22, 2.29, 1.24, 0.75, 9.20; palp: 1.10, 0.52, 0.81, -, 0.65, 3.08; Leg formula 1243; macrosetae: leg I: fe p 1-1, v 23, tip 1-1-1, r 1-1-1-1-1-1, vp 1-1, vr 1-1-1-1-1-1-1-1-1-1-1-1, mtvp 1-1-1, vr 1-0-1; leg II: fep 1, v 1, ti p 1-1-1, r 1-1-1, vp 1, vr 1-1-1-1-1-1-1-1-1-1, mtp 1,vp 1, vr 1-1-1-1-1; leg III: fep 1, tid 2-2, vp 0-0-1, vr 1-1-1, mt d 2-2, vp 0-0-2, vr 1-1-1; leg IV: fe d 1-1, ti vp 1-1-1, vr 0-0-1 mt vp 1-0-1, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV (Fig. 7 D); palp: no spine; teeth on paired claws of legs: I: 12, II: 11, III: 8, IV: 8. Palp patella and tibia slightly incrassate; embolus at the base twists 110° along with distal bulb (Fig. 8 A-D). Rest of the characters same as in the male of I. malkangiri. Variation in males [n=2; measurements: range]. Total length 7.09–7.37; carapace 3.41–3.87 long, 2.29–2.54 wide; abdomen 3.50–3.68 long, 2.00–3.20 wide; clypeus height 0.18–0.23; ocular group 0.30–0.31 long, 0.73– 0.81 wide; eye diameter ALE 0.12–0.17, PME 0.15–0.17, PLE 0.16–0.17; inter-distances PLE-PME 0.10–0.11, ALE-ALE 0.30–0.33; labium: 0.66–0.82 long, 0.43–0.53 wide; endites: 1.20–1.44 long, 0.46–0.52 wide; sternum: 2.00–2.30 long, 1.32–1.44 wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.71–2.93, 0.89–1.00, 2.99–3.40, 2.26–2.64, 1.20–1.34, 10.05–11.31; leg II: 2.61–2.85, 0.98–1.11, 2.44–2.73, 2.27–2.56, 1.18–1.22, 9.42–10.43; leg III: 1.91–2.14, 0.82–0.87, 1.19–1.37, 1.63–1.74, 0.67–0.88, 6.22–7.00; leg IV: 2.48–2.70, 1.02–1.22, 2.02–2.29, 1.24–2.06, 0.74–0.75, 8.32–9.20; palp: 1.09–1.10, 0.50–0.52, 0.68–0.81, -, 0.55–0.65, 2.82–3.08. Description of paratype female (IPU-17-ARA-550) (Fig. 9, 10). Total length 10.21; carapace.4.51 long, 2.81 wide; abdomen 5.70 long, 3.37 wide; carapace posterior border slightly concave, covered with few short hairs expect for long hairs in ocular area and few on caput; clypeus 0.30 high; eyes posterior row slightly recurved (Fig. 9 D); ocular group 0.28 long, 0.87 wide; eye diameter ALE 0.15, PME 0.15, PLE 0.16; inter-distances PLE-PME 0.15, ALE-ALE 0. 44; labium 0.81 long, 0.62 wide; endites 1.42 long, 0.62 wide; sternum 2.58 long, 1.64 wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.89, 1.16, 2.31, 2.03, 0.96, 9.35; leg II: 2.80, 1.25, 2.25, 2.01, 0.95, 9.26; leg III: 2.18, 0.95, 1.16, 1.57, 0.67, 6.53; leg IV: 2.95, 1.28, 2.31, 1.98, 0.66, 9.18; palp: 1.08, 0.63, 0.53, -, 0.88, 3.12; leg formula 1243; macrosetae: leg I: fep 1, tip 1-1-1, v 2-2-2, mt v 0-2-2; leg II: fep 1, tip 0-1-1, vp 0-0-0, vr 0-0-1, mtp 1, vp 0-0-1, vr 1-0-1; leg III: tip 1-1-0, r 1-1-1, vp 0-0-0, vr 0-0-1, mt d 2-2, vp 0-0-2, vr 1-1-1; leg IV: ti vp 0-0-0, vr 0-0-1, mt vp 1-1-2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 1-2-2, ta p 15; teeth on paired claws of legs: I: 17, II: 14, III: 9, IV: 9; female genitalia distinct; internal genitalia with a horn-like anterior receptaculum having broad, elongated base and gradually tapered apex; dorsal lobe spherical; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide, nearly straight, strongly sclerotized forming a conspicuous, but short median flap; posterior receptaculum with irregular edges (Fig. 10 A–E). Remaining characters same as I. malkangiri female. Variation in females [n=7, measurements: range (mean±standard deviation)]. Total length 6.69–12.25 (9.72±1.52); carapace. 3.92–4.89 (4.36±0.28) long, 2.38–2.82 (2.63±0.16) wide; abdomen 4.88–7.36 (5.75±0.74) long, 2.47–4.33 (3.36±0.52) wide; clypeus 0.28–0.35 (0.31±0.02) high; ocular group 0.28–0.55 (0.34±0.09) long, 0.80–0.90 (0.85±0.04) wide; eye diameter ALE 0.15–0.17 (0.16±0.01), PME 0.13–0.15 (0.15±0.01), PLE 0.15–0.18 (0.16±0.01), inter-distances PLE-PME 0.10–0.17 (0.14±0.03), ALE-ALE 0.35–0.55 (0.41±0.06); labium 0.72–0.87 (0.80±0.06) long, 0.51–0.62 (0.59±0.04) wide; endites 1.32–1.58 (1.43±0.09) long, 0.49–0.65 (0.58±0.05) wide; Sternum 2.24–2.70 (2.49±0.14) long, 1.36–1.64 (1.55±0.09) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.51–3.31 (2.88±0.22), 1.08–1.40 (1.20±0.10), 2.07–2.39 (2.25±0.09), 1.83–2.18 (1.99±0.10), 0.93–1.14 (1.02±0.10), 8.42–10.26 (9.34±0.50); leg II: 2.31–3.10 (2.69±0.22), 1.09–1.44 (1.23±0.10), 1.93–2.40 (2.17±0.14), 1.79–2.23 (1.98±0.13), 0.91–1.02 (0.98±0.04), 8.04–10.17 (9.05±0.59); leg III: 1.94–2.31 (2.13±0.11), 0.94–1.07 (0.98±0.05), 1.09–1.34 (1.20±0.09), 1.31–1.72 (1.51±0.12), 0.59–0.71 (0.66±0.04), 5.92–7.15 (6.48±0.34); leg IV: 2.61–3.28 (2.88±0.19), 1.17–1.45 (1.29±0.08), 2.04–2.46 (2.25±0.12), 1.70–2.16 (1.89±0.14), 0.66–0.83 (0.72±0.05), 8.25–10.18 (9.04±0.54); palp: 0.09–1.17 (1.05±0.08), 0.63–0.69 (0.65±0.02), 0.47–0.64 (0.53±0.05), -, 0.81–0.99 (0.91±0.08), 2.82–3.48 (3.14±0.21); preening comb with 3–4 ventro-retrolateral macrosetae on metatarsi IV; teeth on paired claws of all legs 9–17., Published as part of Choudhury, Sudhir Ranjan, Siliwal, Manju, Das, Sanjay Keshari & Giroti, André Marsola, 2021, Description of a new genus and five new species of tube-dwelling spider family Segestriidae (Araneae: Synspermiata) from Odisha, India, pp. 91-114 in Zootaxa 4963 (1) on pages 99-103, DOI: 10.11646/zootaxa.4963.1.4, http://zenodo.org/record/4719875

Indoseges narayani sp. nov. Choudhury, Siliwal, Das & Giroti urn:lsid:zoobank.org:act: 3DC6F099-4A3B-4296-81FA-EF6C3A8DA732 (Fig. 13, 14, 17A) Type specimens: INDIA. holotype ♀, 14.x.2017, Narayani, Ganjam district, Odisha (19° 41’ 52.3”N, 85° 09’ 13.5”E, elev. 138m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-952; paratypes 2 ♀ same data as holotype, IPU-17-ARA-927, IPU-17-ARA-928, same data as holotype. Diagnosis: Females of I. narayani differs from all other congers by having distinct internal genitalia; anterior receptaculum screw-like having a broad base; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide and strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions and having ‘nearly V-shaped’ inner margins; pos- terior receptaculum globular (Fig. 14 A–D). Females also differ from rest of species except I. chilika by having two ventral macrosetae on metatarsi IV (four in I. malkangiri and I. sushildutta, five in I. satkosia) and from I. chilika in absence of dorsal macrosetae on tibiae III (four in I. chilika). Etymology: The species epithet is a noun in apposition, referring to the type locality Narayani. Description of holotype female (IPU-17-ARA-952) (Fig. 13, 14, 17A). Total length 7.35; carapace.3.38 long, 2.10 wide; abdomen 3.97 long, 2.50 wide; carapace anteriorly darker, covered with long and short brown hairs, relatively more on caput; clypeus 0.26 high; ocular group 0.25 long, 0.74 wide; eye diameter ALE 0.13, PME 0.13, PLE 0.13; inter-distances PLE-PME 0.15, ALE-ALE 0.35; abdomen ventrally pale greyish without any pattern; labium 0.60 long, 0.48 wide; endites 1.06 long, 0.46 wide; sternum 1.91 long, 1.26 wide, anterior end darkish without any distinct band (Fig. 13 B); leg measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.00, 0.92, 1.75, 1.61, 0.83,7.11; leg II: 1.96, 0.91, 1.60, 1.53, 0.75, 6.75; leg III: 1.54, 0.77, 0.97, 1.15, 0.51, 4.94; leg IV: 2.12, 1.02, 1.68, 1.37, 0.57, 6.76; palp: 0.71, 0.43, 0.45, -, 0.66, 2.25; leg formula:1423; macrosetae: leg I: fep 1, tip 1- 1-1, v 2-2-2, mtvp 1-0-1, vr 1-1-1; leg II: fep 1, tip 1-1-1, vp 0-0-1, vr 1-0-1, mtp 1, vp 1-0-1, vr 1-1-1; leg III: ti p 1-1-0, r 1-1-0, vp 0-0-0, vr 0-0-1, mt d 2-2, vp 0-0-1,vr 0-1-1; leg IV: ti vp 0-0-0, vr 0-0-1, mt vp 0-1-1, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 2-2, ta p 17; teeth on paired claws of legs: I: 18, II: 15, III: 12, IV: 10; Female genitalia distinct; internal genitalia with a screw-like anterior receptaculum having broad base, dorsal lobe spherical, ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide and strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions having ‘nearly V-shaped’ inner margins; posterior receptaculum globular (Fig. 14 A–D). Remaining characters same as I. malkangiri female. Variation in females [n=3, measurements: range (mean±standard deviation)]. Total length 5.09–7.35 (6.13±1.14). Carapace. 2.48–3.38 (2.80±0.51) long, 0.99–1.26 (1.79±0.27) wide; abdomen 2.61–3.97 (3.34±0.68) long, 1.67–2.50 (2.06±0.42) wide. ocular group 0.20–0.25 (0.23±0.03) long, 0.58–0.74 (0.66±0.08) wide; eye diameter ALE 0.11–0.13 (0.12±0.01), PME 0.12–0.13 (0.12±0.01), PLE 0.12–0.13 (0.12±0.01), inter-distances PLE- PME 0.08–0.15 (0.09±0.05), ALE-ALE 0.22–0.35 (0.27±0.07); clypeus 0.20–0.26 (0.22±0.03) high. Labium 0.46– 0.6 (0.51±0.08) long, 0.35–0.48 (0.40±0.07) wide, Endites 0.8–1.06 (0.90±0.14) long, 0.33–0.46 (0.38±0.07) wide, Sternum 1.47–1.91 (1.63±0.24) long, 0.99–1.26 (1.09±0.15) wide. Leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): Leg I: 1.71–2.00 (1.81±0.17), 0.72–0.92 (0.81±0.10), 1.32–1.75 (1.46±0.25), 1.23–1.61 (1.38±0.20), 0.65–0.83 (0.72±0.10), 5.63–7.11 (6.18±0.81); Leg II: 1.53–1.96 (1.71±0.22), 0.75–0.91 (0.81±0.09), 1.18–1.60 (1.33±0.23), 1.20–1.53 (1.32±0.18), 0.59–0.75 (0.65±0.09), 5.26–6.75 (5.83±0.81); Leg III: 1.28–1.54 (1.37±0.15), 0.59–0.77 (0.68±0.09), 0.70–0.97 (0.80±0.15), 0.92–1.15 (1.03±0.12), 0.41–0.51 (0.44±0.06), 3.91–4.94 (4.32±0.55); Leg IV: 1.72–2.12 (1.85±0.23), 0.81–1.02 (0.88±0.12, 1.34–1.68 (1.47±0.18), 1.12–1.37 (1.22±0.13), 0.45–0.57 (0.50±0.06, 5.44–6.76 (5.93±0.73); Palp: 0.62–0.71 (0.66±0.05), 0.40–0.43 (0.41±0.02), 0.32–0.45 (0.08±0.32), -, 0.55–0.66 (0.59±0.06), 1.89–2.25 (2.02±0.20); preening comb with 3–4 ventroretrolateral macrosetae on metatarsi IV.

Indoseges malkangiri sp. nov. Choudhury, Siliwal, Das & Giroti urn:lsid:zoobank.org:act: 5EB57046-CF2F-4E1E-890A-2E259273866A (Fig. 1–5) Type specimens: INDIA. holotype ♂, 25.v.2017, near Malkangiri town, Malkangiri district, Odisha (18° 21’ 46.3”N, 81° 54’ 43.8”E, elev. 235m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-554; paratypes 1 ♂ same data as holotype, IPU-17-ARA-555, 2 ♂ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda IPU-18-ARA-1272, IPU-18-ARA-1275, 2 ♀ same data as holotype, IPU-17-ARA-557, IPU-17-ARA-558, 3 ♀ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda, IPU-18-ARA-1274, IPU-18-ARA-1276, IPU-18-ARA-1277. Diagnosis: Males of I. malkangiri differs from I. sushildutta by having thorn-like macrosetae on the ventral side of the distal half of metatarsi I (in I. sushildutta cuspule-like macrosetae present) (Fig. 2 A,B); tibia Iwith a row of short macrosetae on the retrolateral side (in I. sushildutta on the ventro-retrolateral side) (Fig. 2 B, 7 A) and lacks any apophysis (in I. sushildutta, tibiae Iwith distinct pointed basal apophysis) (Fig. 2 A–C, 7 A–C); femur Iwith a cluster of stridulatory macrosetae but without any tubercle-like apophysis (in I. sushildutta, prominent tubercle-like femoral apophysis present covered with a cluster of stridulatory spines on the distal face) (Fig. 2 A–D, 7 A–C); leg formula 2143 (in I. sushildutta leg formula in male 1243); palpal tibiae distinctly incrassate and embolus at the base twists 90° along with distal bulb (in I. sushildutta, slightly incrassate and embolus at the base twists 110°) (Fig. 3 A–D, 8 A–D). Females of I. malkangiri differs from all other congers by having two prolateral macrosetae on metatarsi II (in all other species one prolateral macroseta), and a distinct internal genitalia: anterior receptaculum hornlike with a small spherical base and pointed apex; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold narrow, curved and strongly sclerotized forming a very conspicuous and well developed median flap; posterior receptaculum pear shaped (Fig. 5 A–E). Females also differ from rest of the species except I. satkosia by having three ventral macrosetae on tibiae IV (in all other species one ventral macroseta) and from I. sushildutta having leg formula 1423 (in I. sushildutta leg formula in female 1243). Etymology: The species epithet is a noun in apposition, referring to the type locality, Malkangiri in Odisha. Description of holotype male (IPU-17-ARA-554) (Fig. 1–3). Total length 7.50; carapace 3.89 long, 2.58 wide; abdomen 3.61 long, 2.28 wide; carapace covered with long and short brown hairs on caput and posterior margins; clypeus 0.25 high; eyes posterior row straight (Fig. 1 D); ocular group 0.27 long, 0.78 wide; eye diameter ALE 0.14, PME 0.14, PLE 0.14; inter-distances PLE-PME 0.13, ALE-ALE 0.28; abdomen ventrally pale greyish mottled with pale dots; chelicerae retrolaterally with two blunt tubercles at base; labium 0.90 long, 0.51 wide; endites 1.41 long, 0.58 wide; sternum 2.25 long, 1.45 wide, anteriorly marked by thick pale band (Fig. 1 B); legs metatarsi Iin ventral distal half with a thorn-like macroseta; tibiae Iwith a row of short macrosetae on retrolateral side (Fig. 2 B) and lacks any apophysis (Fig. 2 A–C); femur Iwith a cluster of stridulatory macrosetae and lacks prominent tuberclelike apophysis (Fig. 2 A–D); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 3.00, 1.12, 2.61, 2.55, 1.31, 10.59; leg II: 3.21, 1.09, 3.86, 2.78, 1.27, 12.21; leg III: 2.36, 0.86, 1.51, 2.14, 0.82, 7.69; leg IV: 2.98, 1.18, 2.56, 2.51, 1.01, 10.24; palp: 1.25, 0.58, 0.95, -, 0.71, 3.49; leg formula 2143; spines or macrosetae: leg I: fep 16, tip 1-1-1-1, r 1-1-1-1, vp 1-1-1-1, vr 1-1-1-1-1-1-1, mtv 2-0-2; leg II: fep 1-1, tip 1-1-1, r 1-1-1-1, vp 0-0-1, vr 1-1-1, mtvp 1-0-1, vr 1-1-1; leg III: fep 1, tip 1-1-1, r 1-0-1, v 2-1-1-2, mtp 1-0-1, r 1-1-1, v 1-1-3; leg IV: fe d 1-1, ti v 1-1-2, mt vp 1-1-1-2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: no spine; teeth on paired claws of legs: I: 10, II: 8, III: 7, IV: 6; palp brownish-black; patella and tibia distinctly incrassate; cymbium; bulb pear shaped; embolic duct distinct; embolus at the base twists 90° along with distal bulb; embolus long, tubular, tapers and gradually bends dorsally, and spatulated distally with embolic opening on one side (Fig. 3 A–D). Remaining characters same as generic character. Variation in males [n=4; measurements: range (mean±standard deviation)]. Total length 6.88–7.50 (7.26±0.28); carapace. 3.32–3.89 (3.63±0.24) long, 2.09–2.58 (2.28±0.22) wide; abdomen 3.51–3.83 (3.63±0.14) long, 2.01–2.31 (2.21±0.14) wide; clypeus 0.18–0.28 (0.23±0.05) high; ocular group 0.25–0.31 (0.29±0.02) long, 0.61–0.78 (0.69±0.07) wide; eye diameter ALE 0.12–0.19 (0.15±0.03), PME 0.12–0.15 (0.14±0.01), PLE 0.12–0.19 (0.15±0.03); inter-distances PLE-PME 0.08–0.15 (0.12±0.03), ALE-ALE 0.25–0.28 (0.26±0.01); labium 0.63–0.90 (0.78±0.13) long, 0.44–0.51 (0.48±0.04) wide; endites 1.16–1.41 (1.29±0.12) long, 0.43–0.58 (0.51±0.07) wide; sternum 2.00–2.25 (2.13±0.11) long, 1.22–1.45 (1.33±0.12) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.42–3.00 (2.77±0.25), 0.77–1.18 (1.00±0.19), 2.12–2.61 (2.45±0.23), 2.02–2.55 (2.37±0.24), 1.10–1.34 (1.25±0.11), 8.43–10.59 (9.84±0.98); leg II: 2.62–3.21 (2.95±0.25), 1.01–1.19 (1.10±0.07), 2.43–3.86 (2.43±0.62), 2.61–2.88 (2.77±0.11), 1.19–1.34 (1.26±0.07), 9.86–12.21 (11.05±0.98); leg III: 2.02–2.36 (2.21±0.14), 0.83–0.87 (0.85±0.02), 1.24–1.51 (1.41±0.13), 1.80–2.14 (2.00±0.14), 0.81–0.88 (0.83±0.03), 6.70–7.69 (7.30±0.43); leg IV: 2.55–2.98 (2.97±0.01), 1.04–1.25 (1.22±0.05), 2.06–2.56 (2.43±0.19), 2.18–2.62 (2.57±0.08), 0.79–1.01 (0.96±0.08), 8.62–10.24 (9.61±0.72); palp: 1.07–1.25 (1.14±0.08), 0.54–0.58 (0.56±0.02), 0.87–0.95 (0.92±0.03), -, 0.60–0.71 (0.67±0.05), 3.15–3.49 (3.28±0.15). Description of paratype female (IPU-18-ARA-1274) (Fig. 4, 5). Total length 10.96; carapace.4.55 long, 2.75 wide; abdomen 6.41 long, 3.62 wide; carapace covered with long and short brown hairs, relatively more on caput; clypeus 0.30 high; eyes posterior row straight (Fig. 4 D); ocular group 0.29 long, 0.88 wide; eye diameter ALE 0.18, PME 0.15, PLE 0.18; inter-distances PLE-PME 0.18, ALE-ALE 0.46; abdomen ventrally pale greyish mottled with pale dots; labium 0.77 long, 0.58 wide; endites 1.45 long, 0.67 wide; sternum 2.64 long, 1.66 wide, anteriorly marked by a thick pale band (Fig. 4 B); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.87, 1.23, 2.10, 1.91, 0.99, 9.10; leg II: 2.74, 1.21, 2.14, 1.95, 1.00, 9.04; leg III: 2.17, 1.00, 1.21, 1.54, 0.72, 6.64; leg IV: 2.88, 1.31, 2.23, 1.93, 0.72, 9.08; palp: 1.10, 0.65, 0.59, -, 0.92, 3.26; leg formula 1423; macrosetae: leg I: fep 1, tip 1-1-1, v 2-2-2, mtvp 1-0-1, vr 1-1-1; leg II: fep 1, tip 1-1-1, vp 1-0-1, vr 0-0-1, mtp 1-1-0, v 2-1-1-2; leg III: ti p 1-1, v 1-2, r 1-1, mt p 1-1-1-1, r 1-1-1-1, v 1-1-1-3; leg IV: ti vp 1-1, vr 0-1, mt vp 1-1- 2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 1-1, ta p 13; teeth on paired claws of legs: I: 15, II: 13, III: 11, IV: 9; female genitalia distinct; internal genitalia with a horn-like anterior receptaculum having small spherical base and pointed apex; dorsal lobe spherical; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold narrow, curved and strongly sclerotized forming a very conspicuous and well developed median flap; pear shaped posterior receptaculum (Fig. 5 A–E). Remaining characters same as generic character. Variation in females [n=5, measurements: range (mean±standard deviation)]. Total length 8.61–12.80 (10.57±1.61); carapace. 3.69–4.55 (4.17±0.31) long, 2.5–2.75 (2.57±0.13) wide; abdomen 4.92–8.58 (6.40±1.43) long, 2.93–3.62 (3.29±0.26) wide; clypeus 0.24–0.40 (0.32±0.08) high; ocular group 0.25–0.34 (0.30±0.03) long, 0.79–0.94 (0.83±0.10) wide; eye diameter ALE 0.14–0.18 (0.15±0.02), PME 0.13–0.18 (0.15±0.02), PLE 0.14–0.18 (0.15±0.02), inter-distances PLE-PME 0.12–0.18 (0.16±0.03), ALE-ALE 0.36–0.46 (0.39±0.04); labium 0.72–0.78 (0.74±0.03) long, 0.54–0.59 (0.57±0.02) wide; endites 1.26–1.45 (1.35±0.08) long, 0.53–0.67 (0.61±0.05) wide; sternum 2.27–2.64 (2.42±0.16) long, 1.44–1.69 (1.55±0.12) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.56–2.92 (2.75±0.14), 1.08–1.23 (1.15±0.06), 1.94–2.19 (2.08±0.10), 1.79–2.03 (1.94±0.09), 0.97–1.04 (0.99±0.03), 8.85–9.30 (8.92±0.36); leg II: 2.34–2.81 (2.65±0.19), 1.17–1.27 (1.22±0.04), 1.95–2.15 (2.131±0.11), 1.85–2.08 (2.00±0.10), 0.96–1.02 (0.97±0.05), 9.03–9.37 (8.98±0.42); leg III: 1.91–2.22 (2.08±0.12), 0.98–1.02 (0.98±0.06), 1.07–1.22 (1.17±0.06), 1.38–1.54 (1.49±0.07), 0.65–0.73 (0.69±0.03), 6.00–6.72 (6.41±0.30); leg IV: 2.66–2.97 (2.82±0.12), 1.28–1.31 (1.31±0.05), 2.02–2.34 (2.20±0.12), 1.76–1.93 (1.87±0.09), 0.66–0.77 (0.71±0.05), 8.38–9.44 (8.91±0.41); palp: 0.89–1.10 (1.04±0.09), 0.56–0.65 (0.60±0.04), 0.53–0.59 (0.56±0.03), -, 0.78–1.01 (0.90±0.09), 2.76–3.26 (3.09±0.21); teeth on paired claws of legs: I: 14–15, II: 12–13.

Indosegeschilika sp. nov. Siliwal, Das, Choudhury & Giroti urn:lsid:zoobank.org:act: 444BC9A6-7775-4023-8F8D-76B106163CEF (Fig. 11, 12) Type specimens: INDIA. holotype ♀, 14.x.2017, Barkud Island, Chilika lake, Ganjam district, Odisha (19° 33’ 22.6”N, 85° 08’ 47.8”E, elev. 05m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-945; paratypes 3 ♀ same data as holotype, IPU-17-ARA-946, IPU-17-ARA-956, IPU-17-ARA-957. Diagnosis: Females of I. chilika differs from all other congers by having two prolateral macrosetae on tibiae I (in all other species three prolateral macrosetae), four dorsal macrosetae on tibiae III (in all other species dorsal macrosetae absent on tibiae III), and a distinct internal genitalia: anterior receptaculum horn-like with a broad, elongated base and pointed apex; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold very wide and eye-like, strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions having ‘clear V-shaped’ inner margins; posterior receptaculum oval (Fig. 12 A–D). Etymology: The species epithet is a noun in apposition, referring to the Chilika lake, where the type locality, Barkud Island is located. Description of holotype female (IPU-17-ARA-945) (Fig. 11, 12). Total length 6.25; carapace.3.10 long, 1.94 wide; abdomen 3.15 long, 2.13 wide; carapace covered with long and short brown hairs, relatively more on caput; clypeus 0.29 high; ocular group 0.20 long, 0.74 wide; eye diameter ALE 0.12, PME 0.12, PLE 0.13; inter-distances PLE-PME 0.13, ALE-ALE 0.32; abdomen ventrally pale greyish without any pattern; labium 0.60 long, 0.43 wide; endites 1.14 long, 0.45 wide; sternum 1.84 long, 1.23 wide; leg measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.13, 0.91, 1.66, 1.59, 0.75,7.04; leg II: 1.99, 0.90, 1.53, 1.46, 0.63, 6.51; leg III: 1.52, 0.67, 0.98, 1.19, 0.48, 4.84; leg IV: 2.08, 0.92, 1.70, 1.40, 0.52, 6.62; palp: 0.82, 0.52, 0.41, -, 0.71, 2.46; leg formula 1423; macrosetae: leg I: fep 1, tip 1-1, v 2-2-2, mtvp 1-0-1, vr 1-1-1; leg II: fep 1, tip 1-1, vp 0-0-1, vr 1-0-1, mt p 1, vp 0-0-1, vr 1-1-1; leg III: ti d 2-2, vp 0-0-0, vr 0-0-1, mt d 2-2, vp 0-0-1,vr 1-1-1; leg IV: ti vp 0-0-0, vr 0-0- 1, mt vp 1-0-1, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 2-2-2, ta p 12; teeth on paired claws of legs: I: 20, II: 16, III: 10, IV: 9; female genitalia distinct; external genitalia in the form of sclerotized plate; internal genitalia with a horn-like anterior receptaculum having broad, elongated base and pointed apex; dorsal lobe spherical; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold very wide and eye-like, strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions having ‘clear V-shaped’ inner margins; posterior receptaculum oval (Fig. 12 A–D). Remaining characters same as I. malkangiri female. Variation in females [n=4, measurements: range (mean±standard deviation)]. Total length 6.25–6.84 (6.50±0.26), carapace. 2.60–3.37 (2.92±0.38) long, 1.65–2.22 (1.88±0.26) wide; abdomen 3.15–3.94 (3.58±0.35) long, 2.13–2.32 (2.23±0.08) wide; clypeus 0.24–0.29 (0.28±0.02) high; ocular group 0.20–0.25 (0.24±0.03) long, 0.62–0.77 (0.70±0.07) wide; eye diameter ALE 0.12–0.13 (0.12±0.01), PME 0.12–0.13 (0.12±0.01), PLE 0.12–0.13 (0.12±0.01), inter-distances PLE-PME 0.08–0.13 (0.10±0.02), ALE-ALE 0.26–0.32 (0.30±0.03); labium 0.45–0.61 (0.54±0.08) long, 0.37–0.49 (0.42±0.05) wide, endites 0.84–1.14 (0.99±0.15) long, 0.35–0.45 (0.41±0.05) wide; sternum 1.50–2.02 (1.72±0.25) long, 1.01–1.35 (1.16±0.16) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 1.68–2.28 (1.97±0.29), 0.68–0.98 (0.82±0.15), 1.30–1.66 (1.52±0.21), 1.21–1.62 (1.42±0.21), 0.64–0.80 (0.72±0.07), 5.51–7.04 (6.44±0.92); leg II: 1.54–2.01 (1.80±0.24), 0.66–0.98 (0.82±0.14), 1.18–1.60 (1.39±0.21), 1.18–1.52 (1.34±0.17), 0.57–0.72 (0.64±0.06), 5.13–6.83 (5.98±0.82); leg III: 1.24–1.65 (1.43±0.19), 0.59–0.74 (0.66±0.06), 0.67–0.98 (0.81±0.14), 0.85–1.23 (1.05±0.19), 0.46–0.51 (0.47±0.03), 3.78–4.99 (4.42±0.59); leg IV: 1.70–2.31 (1.96±0.29), 0.80–1.10 (0.92±0.13), 1.34–1.70 (1.56±0.22), 1.11–1.51 (1.29±0.19), 0.45–0.58 (0.51±0.06), 5.42–6.62 (6.23±0.88); palp: 0.61–0.82 (0.71±0.10), 0.35–0.52 (0.45±0.08), 0.32–0.43 (0.37±0.06), -, 0.52–0.71 (0.63±0.10), 1.85–2.46 (2.16±0.33).

Indosegeschilika sp. nov. Siliwal, Das, Choudhury & Giroti urn:lsid:zoobank.org:act: 444BC9A6-7775-4023-8F8D-76B106163CEF (Fig. 11, 12) Type specimens: INDIA. holotype ♀, 14.x.2017, Barkud Island, Chilika lake, Ganjam district, Odisha (19° 33’ 22.6”N, 85° 08’ 47.8”E, elev. 05m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-945; paratypes 3 ♀ same data as holotype, IPU-17-ARA-946, IPU-17-ARA-956, IPU-17-ARA-957. Diagnosis: Females of I. chilika differs from all other congers by having two prolateral macrosetae on tibiae I (in all other species three prolateral macrosetae), four dorsal macrosetae on tibiae III (in all other species dorsal macrosetae absent on tibiae III), and a distinct internal genitalia: anterior receptaculum horn-like with a broad, elongated base and pointed apex; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold very wide and eye-like, strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions having ‘clear V-shaped’ inner margins; posterior receptaculum oval (Fig. 12 A–D). Etymology: The species epithet is a noun in apposition, referring to the Chilika lake, where the type locality, Barkud Island is located. Description of holotype female (IPU-17-ARA-945) (Fig. 11, 12). Total length 6.25; carapace.3.10 long, 1.94 wide; abdomen 3.15 long, 2.13 wide; carapace covered with long and short brown hairs, relatively more on caput; clypeus 0.29 high; ocular group 0.20 long, 0.74 wide; eye diameter ALE 0.12, PME 0.12, PLE 0.13; inter-distances PLE-PME 0.13, ALE-ALE 0.32; abdomen ventrally pale greyish without any pattern; labium 0.60 long, 0.43 wide; endites 1.14 long, 0.45 wide; sternum 1.84 long, 1.23 wide; leg measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.13, 0.91, 1.66, 1.59, 0.75,7.04; leg II: 1.99, 0.90, 1.53, 1.46, 0.63, 6.51; leg III: 1.52, 0.67, 0.98, 1.19, 0.48, 4.84; leg IV: 2.08, 0.92, 1.70, 1.40, 0.52, 6.62; palp: 0.82, 0.52, 0.41, -, 0.71, 2.46; leg formula 1423; macrosetae: leg I: fep 1, tip 1-1, v 2-2-2, mtvp 1-0-1, vr 1-1-1; leg II: fep 1, tip 1-1, vp 0-0-1, vr 1-0-1, mt p 1, vp 0-0-1, vr 1-1-1; leg III: ti d 2-2, vp 0-0-0, vr 0-0-1, mt d 2-2, vp 0-0-1,vr 1-1-1; leg IV: ti vp 0-0-0, vr 0-0- 1, mt vp 1-0-1, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 2-2-2, ta p 12; teeth on paired claws of legs: I: 20, II: 16, III: 10, IV: 9; female genitalia distinct; external genitalia in the form of sclerotized plate; internal genitalia with a horn-like anterior receptaculum having broad, elongated base and pointed apex; dorsal lobe spherical; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold very wide and eye-like, strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions having ‘clear V-shaped’ inner margins; posterior receptaculum oval (Fig. 12 A–D). Remaining characters same as I. malkangiri female. Variation in females [n=4, measurements: range (mean±standard deviation)]. Total length 6.25–6.84 (6.50±0.26), carapace. 2.60–3.37 (2.92±0.38) long, 1.65–2.22 (1.88±0.26) wide; abdomen 3.15–3.94 (3.58±0.35) long, 2.13–2.32 (2.23±0.08) wide; clypeus 0.24–0.29 (0.28±0.02) high; ocular group 0.20–0.25 (0.24±0.03) long, 0.62–0.77 (0.70±0.07) wide; eye diameter ALE 0.12–0.13 (0.12±0.01), PME 0.12–0.13 (0.12±0.01), PLE 0.12–0.13 (0.12±0.01), inter-distances PLE-PME 0.08–0.13 (0.10±0.02), ALE-ALE 0.26–0.32 (0.30±0.03); labium 0.45–0.61 (0.54±0.08) long, 0.37–0.49 (0.42±0.05) wide, endites 0.84–1.14 (0.99±0.15) long, 0.35–0.45 (0.41±0.05) wide; sternum 1.50–2.02 (1.72±0.25) long, 1.01–1.35 (1.16±0.16) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 1.68–2.28 (1.97±0.29), 0.68–0.98 (0.82±0.15), 1.30–1.66 (1.52±0.21), 1.21–1.62 (1.42±0.21), 0.64–0.80 (0.72±0.07), 5.51–7.04 (6.44±0.92); leg II: 1.54–2.01 (1.80±0.24), 0.66–0.98 (0.82±0.14), 1.18–1.60 (1.39±0.21), 1.18–1.52 (1.34±0.17), 0.57–0.72 (0.64±0.06), 5.13–6.83 (5.98±0.82); leg III: 1.24–1.65 (1.43±0.19), 0.59–0.74 (0.66±0.06), 0.67–0.98 (0.81±0.14), 0.85–1.23 (1.05±0.19), 0.46–0.51 (0.47±0.03), 3.78–4.99 (4.42±0.59); leg IV: 1.70–2.31 (1.96±0.29), 0.80–1.10 (0.92±0.13), 1.34–1.70 (1.56±0.22), 1.11–1.51 (1.29±0.19), 0.45–0.58 (0.51±0.06), 5.42–6.62 (6.23±0.88); palp: 0.61–0.82 (0.71±0.10), 0.35–0.52 (0.45±0.08), 0.32–0.43 (0.37±0.06), -, 0.52–0.71 (0.63±0.10), 1.85–2.46 (2.16±0.33)., Published as part of Choudhury, Sudhir Ranjan, Siliwal, Manju, Das, Sanjay Keshari & Giroti, André Marsola, 2021, Description of a new genus and five new species of tube-dwelling spider family Segestriidae (Araneae: Synspermiata) from Odisha, India, pp. 91-114 in Zootaxa 4963 (1) on pages 104-106, DOI: 10.11646/zootaxa.4963.1.4, http://zenodo.org/record/4719875

Indoseges sushildutta sp. nov. Siliwal, Das, Choudhury, & Giroti urn:lsid:zoobank.org:act: 333AD004-862B-4F37-B27D-121C3E12AAFD (Fig. 6–10) Type specimens: INDIA- holotype ♂, 25.v.2017, Govindpali ghati, Malkangiri district, Odisha (18° 36’ 34.3”N, 82° 18’ 12.3”E, elev. 248m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-560; paratypes 1 ♂ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda, IPU-18-ARA-1286, 4 ♀ same data as holotype, IPU- 17-ARA-548, IPU-17-ARA-549, IPU-17-ARA-550, IPU-17-ARA-559, 3 ♀ 08.vi.2018, same locality as holotype, coll. S. R. Choudhury and N. Panda, IPU-18-ARA-1281, IPU-18-ARA-1287, IPU-18-ARA-1290. Diagnosis: Males of I. sushildutta differs from I. malkangiri by having capsule-like macrosetae on the ventral side of the distal half of metatarsi I (in I. malkangiri thorn-like macrosetae present); tibiae Iwith a row of short macrosetae on the ventro-retrolateral side (in I. malkangiri on the retrolateral side) (Fig. 2 B, 7 A) and a distinct pointed prolateral basal apophysis (in I. malkangiri, tibiae Ilacks any apophysis) (Fig. 2 A–C, 7 A–C); femur Iwith a distinct triangular tubercle-like prolateral femoral apophysis covered by a cluster of stridulatory macrosetae on the distal face (in I. malkangiri, femur Iwith a cluster of stridulatory macrosetae and lacks any apophysis) (Fig. 2 A–D, 7 A–C); leg formula 1243 (in I. malkangiri leg formula in male 2143); palpal tibiae slightly incrassate and embolus at the base twists 110° along with distal bulb (in I. malkangiri, distinctly incrassate and embolus at the base twists 90°) (Fig. 3 A–D, 8 A–D). Females of I. sushildutta differs from all other congers by having leg formula of 1243 (in all other species it is 1423), four ventral macrosetae on metatarsi I (in all other species five ventral macrosetae), and a distinct internal genitalia: anterior receptaculum horn-like with a broad, elongated base and gradually tapered apex; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide, nearly straight, strongly sclerotized forming a conspicuous, but short median flap; posterior receptaculum with irregular edges (Fig. 10 A–E). Etymology: The species epithet is a noun in apposition, in honor of renowned herpetologist, Prof. Sushil Kumar Dutta for encouraging and supporting our studies on spiders in Odisha. Description of holotype male (IPU-17-ARA-560) (Fig. 6–8). Total length 7.37; carapace.3.87 long, 2.54 wide; abdomen 3.50 long, 3.20 wide; clypeus 0.23 high; ocular group 0.31 long, 0.81 wide; eye diameter ALE 0.17, PME 0.17, PLE 0.17; inter-distances PLE-PME 0.11, ALE-ALE 0.33; chelicerae retrolaterally with one blunt tubercle at base; labium 0.82 long, 0.53 wide; endites 1.44 long, 0.52 wide; sternum 2.30 long, 1.44 wide; metatarsi Iwith a single row of small cuspule like ventral macrosetae on distal half; tibiae Iwith a row of short macrosetae on ventro-retrolateral side (Fig. 7 A), and a distinct pointed basal prolateral apophysis (Fig. 7 A–C); femur Iwith a distinct triangular tubercle-like prolateral femoral apophysis covered by a cluster of stridulatory spines on distal face (Fig. 7 A–C); leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.93, 1.00, 3.40, 2.64, 1.34, 11.31; leg II: 2.85, 1.11, 2.73, 2.56, 1.18, 10.43; leg III: 2.14, 0.87, 1.37, 1.74, 0.88, 7.00; leg IV: 2.70, 1.22, 2.29, 1.24, 0.75, 9.20; palp: 1.10, 0.52, 0.81, -, 0.65, 3.08; Leg formula 1243; macrosetae: leg I: fe p 1-1, v 23, tip 1-1-1, r 1-1-1-1-1-1, vp 1-1, vr 1-1-1-1-1-1-1-1-1-1-1-1, mtvp 1-1-1, vr 1-0-1; leg II: fep 1, v 1, ti p 1-1-1, r 1-1-1, vp 1, vr 1-1-1-1-1-1-1-1-1-1, mtp 1,vp 1, vr 1-1-1-1-1; leg III: fep 1, tid 2-2, vp 0-0-1, vr 1-1-1, mt d 2-2, vp 0-0-2, vr 1-1-1; leg IV: fe d 1-1, ti vp 1-1-1, vr 0-0-1 mt vp 1-0-1, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV (Fig. 7 D); palp: no spine; teeth on paired claws of legs: I: 12, II: 11, III: 8, IV: 8. Palp patella and tibia slightly incrassate; embolus at the base twists 110° along with distal bulb (Fig. 8 A-D). Rest of the characters same as in the male of I. malkangiri. Variation in males [n=2; measurements: range]. Total length 7.09–7.37; carapace 3.41–3.87 long, 2.29–2.54 wide; abdomen 3.50–3.68 long, 2.00–3.20 wide; clypeus height 0.18–0.23; ocular group 0.30–0.31 long, 0.73– 0.81 wide; eye diameter ALE 0.12–0.17, PME 0.15–0.17, PLE 0.16–0.17; inter-distances PLE-PME 0.10–0.11, ALE-ALE 0.30–0.33; labium: 0.66–0.82 long, 0.43–0.53 wide; endites: 1.20–1.44 long, 0.46–0.52 wide; sternum: 2.00–2.30 long, 1.32–1.44 wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.71–2.93, 0.89–1.00, 2.99–3.40, 2.26–2.64, 1.20–1.34, 10.05–11.31; leg II: 2.61–2.85, 0.98–1.11, 2.44–2.73, 2.27–2.56, 1.18–1.22, 9.42–10.43; leg III: 1.91–2.14, 0.82–0.87, 1.19–1.37, 1.63–1.74, 0.67–0.88, 6.22–7.00; leg IV: 2.48–2.70, 1.02–1.22, 2.02–2.29, 1.24–2.06, 0.74–0.75, 8.32–9.20; palp: 1.09–1.10, 0.50–0.52, 0.68–0.81, -, 0.55–0.65, 2.82–3.08. Description of paratype female (IPU-17-ARA-550) (Fig. 9, 10). Total length 10.21; carapace.4.51 long, 2.81 wide; abdomen 5.70 long, 3.37 wide; carapace posterior border slightly concave, covered with few short hairs expect for long hairs in ocular area and few on caput; clypeus 0.30 high; eyes posterior row slightly recurved (Fig. 9 D); ocular group 0.28 long, 0.87 wide; eye diameter ALE 0.15, PME 0.15, PLE 0.16; inter-distances PLE-PME 0.15, ALE-ALE 0. 44; labium 0.81 long, 0.62 wide; endites 1.42 long, 0.62 wide; sternum 2.58 long, 1.64 wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.89, 1.16, 2.31, 2.03, 0.96, 9.35; leg II: 2.80, 1.25, 2.25, 2.01, 0.95, 9.26; leg III: 2.18, 0.95, 1.16, 1.57, 0.67, 6.53; leg IV: 2.95, 1.28, 2.31, 1.98, 0.66, 9.18; palp: 1.08, 0.63, 0.53, -, 0.88, 3.12; leg formula 1243; macrosetae: leg I: fep 1, tip 1-1-1, v 2-2-2, mt v 0-2-2; leg II: fep 1, tip 0-1-1, vp 0-0-0, vr 0-0-1, mtp 1, vp 0-0-1, vr 1-0-1; leg III: tip 1-1-0, r 1-1-1, vp 0-0-0, vr 0-0-1, mt d 2-2, vp 0-0-2, vr 1-1-1; leg IV: ti vp 0-0-0, vr 0-0-1, mt vp 1-1-2, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 1-2-2, ta p 15; teeth on paired claws of legs: I: 17, II: 14, III: 9, IV: 9; female genitalia distinct; internal genitalia with a horn-like anterior receptaculum having broad, elongated base and gradually tapered apex; dorsal lobe spherical; ventral lobe with few shallow entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide, nearly straight, strongly sclerotized forming a conspicuous, but short median flap; posterior receptaculum with irregular edges (Fig. 10 A–E). Remaining characters same as I. malkangiri female. Variation in females [n=7, measurements: range (mean±standard deviation)]. Total length 6.69–12.25 (9.72±1.52); carapace. 3.92–4.89 (4.36±0.28) long, 2.38–2.82 (2.63±0.16) wide; abdomen 4.88–7.36 (5.75±0.74) long, 2.47–4.33 (3.36±0.52) wide; clypeus 0.28–0.35 (0.31±0.02) high; ocular group 0.28–0.55 (0.34±0.09) long, 0.80–0.90 (0.85±0.04) wide; eye diameter ALE 0.15–0.17 (0.16±0.01), PME 0.13–0.15 (0.15±0.01), PLE 0.15–0.18 (0.16±0.01), inter-distances PLE-PME 0.10–0.17 (0.14±0.03), ALE-ALE 0.35–0.55 (0.41±0.06); labium 0.72–0.87 (0.80±0.06) long, 0.51–0.62 (0.59±0.04) wide; endites 1.32–1.58 (1.43±0.09) long, 0.49–0.65 (0.58±0.05) wide; Sternum 2.24–2.70 (2.49±0.14) long, 1.36–1.64 (1.55±0.09) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.51–3.31 (2.88±0.22), 1.08–1.40 (1.20±0.10), 2.07–2.39 (2.25±0.09), 1.83–2.18 (1.99±0.10), 0.93–1.14 (1.02±0.10), 8.42–10.26 (9.34±0.50); leg II: 2.31–3.10 (2.69±0.22), 1.09–1.44 (1.23±0.10), 1.93–2.40 (2.17±0.14), 1.79–2.23 (1.98±0.13), 0.91–1.02 (0.98±0.04), 8.04–10.17 (9.05±0.59); leg III: 1.94–2.31 (2.13±0.11), 0.94–1.07 (0.98±0.05), 1.09–1.34 (1.20±0.09), 1.31–1.72 (1.51±0.12), 0.59–0.71 (0.66±0.04), 5.92–7.15 (6.48±0.34); leg IV: 2.61–3.28 (2.88±0.19), 1.17–1.45 (1.29±0.08), 2.04–2.46 (2.25±0.12), 1.70–2.16 (1.89±0.14), 0.66–0.83 (0.72±0.05), 8.25–10.18 (9.04±0.54); palp: 0.09–1.17 (1.05±0.08), 0.63–0.69 (0.65±0.02), 0.47–0.64 (0.53±0.05), -, 0.81–0.99 (0.91±0.08), 2.82–3.48 (3.14±0.21); preening comb with 3–4 ventro-retrolateral macrosetae on metatarsi IV; teeth on paired claws of all legs 9–17.

Indoseges satkosia sp. nov. Das, Siliwal, Choudhury & Giroti urn:lsid:zoobank.org:act: AD447F5B-2E55-4524-BC25-4F82A889F958 (Fig. 15, 16, 17B) Type specimens: INDIA. holotype ♀, 02.iii.2017, Tuluka road, Satkosia, Anugul district, Odisha (20° 36’ 19.9”N, 84° 57’ 22.0”E, elev. 178m), coll. S.K. Das, S. R. Choudhury and N. Panda, IPU-17-ARA-460; paratypes 2 ♀ same data as holotype, IPU-17-ARA-455, IPU-17-ARA-456. Diagnosis: Females of I. satkosia differs from all other congers by having 2-4 arrangement of ventral macrosetae on tibiae I (in all other species it is 3-3 arrangement of ventral macrosetae), five ventral macrosetae on metatarsi IV (four in I. malkangiri and I. sushildutta, two in I. chilika and I. satkosia), and a distinct internal genitalia: anterior receptaculum stump-like with a broad, elongated base and pointed apex being twisted distally; ventral lobe with few shallow entapophyses; uterus externus indistinct; interpulmonary fold sclerotized forming a very conspicuous and well developed median flap and with crossed lateral extensions; posterior receptaculum with conical edges (Fig. 16 A-D). Females also differ from rest of the species except I. malkangiri by having three ventral macrosetae on tibiae IV (in all other species one ventral macroseta) Etymology: The species epithet is a noun in apposition, referring to the type locality, Satkosia. Description of holotype female (IPU-17-ARA-460) (Fig. 15, 16, 17B). Total length 9.16; carapace 4.36 long, 2.68 wide; abdomen 4.80 long, 2.95 wide; carapace covered with long and short brown hairs, relatively more on caput; clypeus 0.30 high; ocular group 0.25 long, 0.90 wide; eye diameter ALE 0.15, PME 0.15, PLE 0.16; inter-distances PLE-PME 0.15, ALE-ALE 0.47; abdomen ventrally pale greyish mottled with pale dots in radiating pattern; labium 0.86 long, 0.57 wide; endites 1.42 long, 0.54 wide; sternum 2.36 long, 1.47 wide; leg measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.79, 1.16, 2.19, 2.02, 0.94, 9.10; leg II: 2.69, 1.16, 2.20, 1.97, 0.95, 8.99; leg III: 2.11, 0.99, 1.13, 1.48, 0.65, 6.36; leg IV: 2.88, 1.31, 2.27, 1.90, 0.65, 9.01; palp: 1.01, 0.58, 0.52, -, 0.93, 3.04; leg formula 1423; macrosetae: leg I: fep 1, ti p1-1-1, vp 1-0-0-1, vr 1-1-1-1, mtvp 1-1-1, vr 1-0-1; leg II: fep 1, tip 1-1-1, vp 0-0-1, vr 1-1-1, mtp 1, vp 0-1-1, vr 1-1-1; leg III: tip 1-1, r 1-1, v 0-0-2, mtp 1-1-0, r 1-1-0, vp 0-0-2, vr 1-1-2; leg IV: ti vp 0-0-1, vr 0-1-1, mt vp 0-0-0-0, vr 1-1-1-2, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 1-1-1, ta p 11; teeth on paired claws of legs: I: 15, II: 13, III: 10, IV: 9; female genitalia distinct; internal genitalia with a stump-like anterior receptaculum having a broad, elongated base and pointed apex being twisted distally; dorsal lobe spherical; ventral lobe with few shallow entapophyses; uterus externus indistinct; interpulmonary fold sclerotized forming a very conspicuous and well developed median flap and with crossed lateral extensions; posterior receptaculum with conical edges (Fig. 16 A–D). Remaining characters same as in females of I. malkangiri. Variation in females [n=3, measurements: range (mean±standard deviation)]. Total length 8.80–9.16 (9.01±0.19); carapace. 4.00–4.36 (4.14±0.20) long, 2.39–2.68 (2.51±0.15) wide; abdomen 4.75–5.08 (4.88±0.18) long, 2.80–3.24 (3.00±0.22) wide; clypeus 0.30 (0.30±0.00) high; ocular group 0.25–0.32 (0.29±0.04) long, 0.82– 0.90 (0.85±0.04) wide; eye diameter ALE 0.15–0.18 (0.17±0.02), PME 0.15–0.18 (0.17±0.02), PLE 0.16–0.18 (0.17±0.01); inter-distances PLE-PME 0.09–0.15 (0.11±0.03), ALE-ALE 0.35–0.47 (0.41±0.06); labium 0.73–0.86 (0.78±0.07) long, 00.52–0.57 (0.53±0.03) wide; endites 1.24–1.42 (1.31±0.10) long, 0.49–0.54 (0.52±0.03) wide; sternum 2.28–2.36 (2.31±0.05) long, 1.39–1.47 (1.42±0.04) wide; leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.50–2.79 (2.60±0.16), 1.09–1.16 (1.12±0.04), 1.86–2.19 (1.99±0.18), 1.81–2.02 (1.88±0.12), 0.94–0.99 (0.97±0.03), 8.33–9.10 (8.56±0.47); leg II: 2.38–2.69 (2.51±0.16), 1.10–1.16 (1.13±0.03), 1.88–2.22 (2.00±0.19), 1.70–1.97 (1.85±0.14), 0.92–0.95 (0.93±0.02), 8.05–8.99 (8.43±0.50); leg III: 1.88–2.11 (1.96±0.13), 0.85–0.99 (0.91±0.07), 1.07–1.13 (1.10±0.03), 1.35–1.48 (1.41±0.07), 0.57–0.65 (0.62±0.04), 5.79–6.36 (6.00±0.31); leg IV: 2.58–2.88 (2.68±0.17), 1.23–1.31 (1.26±0.04), 2.00–2.27 (2.12±0.14), 1.68–1.90 (1.77±0.11), 0.65–0.66 (0.66±0.01), 8.15–9.01 (8.49±0.46); palp: 0.89–1.01 (0.96±0.06), 0.56–0.58 (0.57±0.01), 0.48–0.53 (0.51±0.03), 0.78–0.93 (0.83±0.08), -, 0.78–0.93 (0.83±0.08), 2.78–3.04 (2.88±0.14).

Indoseges narayani sp. nov. Choudhury, Siliwal, Das & Giroti urn:lsid:zoobank.org:act: 3DC6F099-4A3B-4296-81FA-EF6C3A8DA732 (Fig. 13, 14, 17A) Type specimens: INDIA. holotype ♀, 14.x.2017, Narayani, Ganjam district, Odisha (19° 41’ 52.3”N, 85° 09’ 13.5”E, elev. 138m), coll. S. R. Choudhury and N. Panda, IPU-17-ARA-952; paratypes 2 ♀ same data as holotype, IPU-17-ARA-927, IPU-17-ARA-928, same data as holotype. Diagnosis: Females of I. narayani differs from all other congers by having distinct internal genitalia; anterior receptaculum screw-like having a broad base; ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide and strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions and having ‘nearly V-shaped’ inner margins; pos- terior receptaculum globular (Fig. 14 A–D). Females also differ from rest of species except I. chilika by having two ventral macrosetae on metatarsi IV (four in I. malkangiri and I. sushildutta, five in I. satkosia) and from I. chilika in absence of dorsal macrosetae on tibiae III (four in I. chilika). Etymology: The species epithet is a noun in apposition, referring to the type locality Narayani. Description of holotype female (IPU-17-ARA-952) (Fig. 13, 14, 17A). Total length 7.35; carapace.3.38 long, 2.10 wide; abdomen 3.97 long, 2.50 wide; carapace anteriorly darker, covered with long and short brown hairs, relatively more on caput; clypeus 0.26 high; ocular group 0.25 long, 0.74 wide; eye diameter ALE 0.13, PME 0.13, PLE 0.13; inter-distances PLE-PME 0.15, ALE-ALE 0.35; abdomen ventrally pale greyish without any pattern; labium 0.60 long, 0.48 wide; endites 1.06 long, 0.46 wide; sternum 1.91 long, 1.26 wide, anterior end darkish without any distinct band (Fig. 13 B); leg measurements (femur, patella, tibia, metatarsus, tarsus, total length): leg I: 2.00, 0.92, 1.75, 1.61, 0.83,7.11; leg II: 1.96, 0.91, 1.60, 1.53, 0.75, 6.75; leg III: 1.54, 0.77, 0.97, 1.15, 0.51, 4.94; leg IV: 2.12, 1.02, 1.68, 1.37, 0.57, 6.76; palp: 0.71, 0.43, 0.45, -, 0.66, 2.25; leg formula:1423; macrosetae: leg I: fep 1, tip 1- 1-1, v 2-2-2, mtvp 1-0-1, vr 1-1-1; leg II: fep 1, tip 1-1-1, vp 0-0-1, vr 1-0-1, mtp 1, vp 1-0-1, vr 1-1-1; leg III: ti p 1-1-0, r 1-1-0, vp 0-0-0, vr 0-0-1, mt d 2-2, vp 0-0-1,vr 0-1-1; leg IV: ti vp 0-0-0, vr 0-0-1, mt vp 0-1-1, vr 0-0-0, preening comb with four ventro-retrolateral macrosetae on metatarsi IV; palp: ti p 2-2, ta p 17; teeth on paired claws of legs: I: 18, II: 15, III: 12, IV: 10; Female genitalia distinct; internal genitalia with a screw-like anterior receptaculum having broad base, dorsal lobe spherical, ventral lobe with many elongated entapophyses; uterus externus weakly sclerotized and transparent; interpulmonary fold wide and strongly sclerotized forming a very conspicuous and well developed median flap, with prominent lateral extensions having ‘nearly V-shaped’ inner margins; posterior receptaculum globular (Fig. 14 A–D). Remaining characters same as I. malkangiri female. Variation in females [n=3, measurements: range (mean±standard deviation)]. Total length 5.09–7.35 (6.13±1.14). Carapace. 2.48–3.38 (2.80±0.51) long, 0.99–1.26 (1.79±0.27) wide; abdomen 2.61–3.97 (3.34±0.68) long, 1.67–2.50 (2.06±0.42) wide. ocular group 0.20–0.25 (0.23±0.03) long, 0.58–0.74 (0.66±0.08) wide; eye diameter ALE 0.11–0.13 (0.12±0.01), PME 0.12–0.13 (0.12±0.01), PLE 0.12–0.13 (0.12±0.01), inter-distances PLE- PME 0.08–0.15 (0.09±0.05), ALE-ALE 0.22–0.35 (0.27±0.07); clypeus 0.20–0.26 (0.22±0.03) high. Labium 0.46– 0.6 (0.51±0.08) long, 0.35–0.48 (0.40±0.07) wide, Endites 0.8–1.06 (0.90±0.14) long, 0.33–0.46 (0.38±0.07) wide, Sternum 1.47–1.91 (1.63±0.24) long, 0.99–1.26 (1.09±0.15) wide. Leg and palp measurements (femur, patella, tibia, metatarsus, tarsus, total length): Leg I: 1.71–2.00 (1.81±0.17), 0.72–0.92 (0.81±0.10), 1.32–1.75 (1.46±0.25), 1.23–1.61 (1.38±0.20), 0.65–0.83 (0.72±0.10), 5.63–7.11 (6.18±0.81); Leg II: 1.53–1.96 (1.71±0.22), 0.75–0.91 (0.81±0.09), 1.18–1.60 (1.33±0.23), 1.20–1.53 (1.32±0.18), 0.59–0.75 (0.65±0.09), 5.26–6.75 (5.83±0.81); Leg III: 1.28–1.54 (1.37±0.15), 0.59–0.77 (0.68±0.09), 0.70–0.97 (0.80±0.15), 0.92–1.15 (1.03±0.12), 0.41–0.51 (0.44±0.06), 3.91–4.94 (4.32±0.55); Leg IV: 1.72–2.12 (1.85±0.23), 0.81–1.02 (0.88±0.12, 1.34–1.68 (1.47±0.18), 1.12–1.37 (1.22±0.13), 0.45–0.57 (0.50±0.06, 5.44–6.76 (5.93±0.73); Palp: 0.62–0.71 (0.66±0.05), 0.40–0.43 (0.41±0.02), 0.32–0.45 (0.08±0.32), -, 0.55–0.66 (0.59±0.06), 1.89–2.25 (2.02±0.20); preening comb with 3–4 ventroretrolateral macrosetae on metatarsi IV., Published as part of Choudhury, Sudhir Ranjan, Siliwal, Manju, Das, Sanjay Keshari & Giroti, André Marsola, 2021, Description of a new genus and five new species of tube-dwelling spider family Segestriidae (Araneae: Synspermiata) from Odisha, India, pp. 91-114 in Zootaxa 4963 (1) on pages 107-110, DOI: 10.11646/zootaxa.4963.1.4, http://zenodo.org/record/4719875

Gupta, Neha, Ganeshkumar, M., Das, Sanjay Keshari, Siliwal, Manju (2013): Three new species of Idiops Perty, 1833 (Araneae: Idiopidae) from India. Zootaxa 3635 (3): 237-250, DOI: http://dx.doi.org/10.11646/zootaxa.3635.3.3

Idiops mettupalayam sp. nov. Ganeshkumar and Siliwal (Figs 3 A���H, Table 2) Type specimens. Holotype male. INDIA: Tamil Nadu: Mettupalayam, Coimbatore, [11 �� 32 ��� 28 ������N & 76 �� 83 ��� 43 ������E], 29 May 2006, coll. M. Chandrasekaran (WILD-06-ARA- 142). Paratypes. INDIA: Tamil Nadu: 2 males, same data as holotype, 2 June 2006, (WILD-06-ARA- 143, WILD- 06-ARA- 144); male, same data as holotype, 30 May 2006 (WILD-06-ARA- 145). Diagnosis. Males of Idiops mettupalayam sp. nov. resemble those of I. bombayensis and I. constructor in having a triangular spine on tibial spur on leg I (Fig. 3 E), but can be distinguished from those species by having mt I strongly incrassate for most of the length (Fig. 3 D) and leg I distinctly longer than leg IV (in I. bombayensis mt I is deeply incrassate in basal 3 / 4 th with indistinct prolateral process; in I. joida, mt I is slender and lacks prolateral process; in I. constructor mt I is strongly incrassate in basal 3 / 4 th and leg I and leg IV equal in length). Etymology. The species epithet is a name in apposition from the type locality, Mettupalayam, located at the foothills of the Western Ghats of Tamil Nadu. Description. Total length 8.53. Carapace 4.53 long, 3.70 wide; chelicerae 1.20 long; abdomen 4.00 long, 2.30 wide. Spinnerets: PMS, 0.27 long, 0.10 wide; PLS, 0.27 basal, 0.47 middle, 0.33 distal; midwidths 0.40, 0.27, 0.20 respectively; 1.07 total length. LEG I LEG II LEG III LEG IV PALP HT*= Holotype, PT= Paratypes Colour in alcohol. Carapace, leg I and palp reddish-brown, other legs yellowish-brown. Abdomen grayishbrown, pattern probably absent (as abdomen have shrunken, pattern not clear). AME encircled with black patch; black patches on caput and few irregular faint patches on rest of carapace. Carapace (Fig. 3 A). Oval, smooth except for few scattered wart-like tubercles along sides of striae; few row of hairs along posterior striae; black patch on caput; fovea deep, procurved. Eyes (Fig. 3 B). Eight in three rows, ALE situated far from AME on clypeal edge; posterior row procurved. Ocular group 1.00 long, 0.67 wide; MOQ almost square, 0.67 front and back width, 0.50 long. Diameter AME 0.28, PME 0.17, ALE 0.33, PLE 0.27; distance between ALE-AME 0.33, AME-AME 0.10, PME-PME 0.20, ALE- PLE 0.10, PLE-PME and ALE-ALE adjacent. Maxillae. 1.20 long anteriorly, 1.47 long posteriorly, 0.67 wide; cuspules absent; anterior lobe distinct, sparsely covered with long and short black bristles. Labium. 0.67 long, 0.60 wide, labiosternal groove shallow, slightly procurved, cuspules absent. Chelicerae (Fig. 3 C). 8 promarginal teeth in single row (size gradually decreasing from anterior to posterior end) on sclerotized ridge and single big tooth on outer margin of row of promarginal teeth; rastellum strong, raised on high triangular mound, consist of 3 large and 8 small thick, spines with 8 spinules on dorso-prolateral edge; many normal pointed spines vertically face and up dorsally on chelicerae; two glabrous bands for length of dorsal surface of chelicerae. Sternum. Broader between posterior coxae; yellowish-brown, elevated in centre, sloping laterally, covered with long black hair; row of long bristles on margins, posterior angle blunt but not separating coxae IV. Sigilla. Posterior sigilla absent; anterior and median pairs not visible due to shrinking of sternum. Legs. Leg I thicker than other legs; ti I inflated with pointed triangular spur facing upward on tubercle and small pointed process below tubercle (Figs 3 D���E); mt prolateraly incrassate 3 / 4 th length, with distinct prolateral process at 1 / 4 th from anterior end (Fig. 3 D); femorae and tibiae III clearly wider than others; metatarsi of all legs longer than tarsi. Legs cylindrical, covered with few scattered hair, bristles and few curved thick thorn-like spines. Two conspicuous glabrous bands for length of femora, patellae and tibiae (not very prominent on leg I). Leg formula 1423. Spines. More on promarginal and retromarginal sides of legs and palp. I: pa, v= 3; ti, v= 11, p= 1 + 2 spur; mt, p= 1, v= 5, r= 5; ta, p= 4, r= 9. II: fe, d= 7, p= 2, r= 3; pa, p= 2, v= 4; ti, p= 4 +broken, v= 9, r= 3; mt, p= 5, v= 10, r= 3; ta, p= 1, r= 6. III: fe, d= 11; pa, p= 14, d= 7, r= 2; ti, p= 8, r=d= 5, v= 7; mt, p= 6, v= 8, r= 6; ta, p= 1, r= 3. IV: fe, d= 8; pa, p= 18, d= 5 (many spinules p= 110, d= 25, r= 12); ti, v= 7, r= 6; mt, v= 7, r= 2; ta, p= 5. Palp: fe, d= 4; ti, r= 26; ta, d= 6. Trichobothria. Clavate absent; ta I, 12 long filiform in each of two zig-zag rows for length; ta II���III, 14 long filiform in 2 zig-zag rows for length; ta IV, 18 long filiform in basal two thirds; palp, 6 short filiform in centre of cymbium. Mt I, 5 short filiform in distal fourth; mt II���IV, 6���7 short filiform in distal fourth. Leg coxae. Coxa I���IV sparsely covered with long and short black bristles, more towards lateral than in centre. Coxae III with central patch without hair or spinules, others sparsely covered with long and short bristles. Coxae IV clearly broader than others, anterior edge curved, sparsely covered long bristles. Claws. Paired claw on leg I���III with unequal three teeth and leg IV with two unequal teeth; unpaired claw on all legs. Both claws (paired as well unpaired) on leg IV larger in size, well developed and prominent than on legs I���III. False claw tufts on each side of paired claws. Scopulae. On all leg tarsi; ta I, few scopuliform hairs in distal half, ta II���IV entire. Abdomen (Fig. 3 A). Grayish-brown covered with short and long black hair; cuticle appears leathery and slightly rough. Spinnerets. PMS digitiform covered with brown hair; PLS covered with brown hair, apical segment domed. Palp (Figs 3 F���H). Tarsi with two unequal lateral processes forming cavity on ventral. Median haematodocha extensive; embolus straight with distally twists about 90 o; tip of embolus flat, wedge-like with central notch. Remarks. Paratypes are like the holotype. The paratype specimens are alcohol dried up and therefore only leg morphometry is provided for paratype specimen WILD-06-ARA- 144 (Table 2). Natural history. This species was found in dry deciduous habitat, mostly dominated by bamboo. All the males were collected in pit-fall traps, during first monsoon showers. Efforts were made to find females but they could not be located; probably females make vertical burrow on the ground and therefore it was difficult to spot them. The soil was hard and red in colour., Published as part of Gupta, Neha, Ganeshkumar, M., Das, Sanjay Keshari & Siliwal, Manju, 2013, Three new species of Idiops Perty, 1833 (Araneae: Idiopidae) from India, pp. 237-250 in Zootaxa 3635 (3) on pages 244-246, DOI: 10.11646/zootaxa.3635.3.3, http://zenodo.org/record/216039

Idiops oriya sp. nov. Siliwal (Figs 4 A���E, Table 3) Type specimen. Holotype female. INDIA: Odhisa: Kapilas, Dhenkanal, [19 �� 44 ��� 39.1 ������N & 83 ��0 6 ��� 34.8 ������E], 0 6 August, 2007, elev. 280 m, coll. Saroj Behera (WILD-07-ARA- 198). Diagnosis. The new species resembles I. pylorus and I. madrasensis and differs from rest of the South and South-east Asian Idiops sp. in females having leg formula 4123. Females differs from those of I. pylorus by having ocular area longer than wide and those of I. pylorus and I. madrasensis in spermathecae structure being ice-cream cone-shaped (Fig. 4 E). Etymology. The species epithet refers to the vernacular language spoken in the type locality state, Odhisa. Description. Total length 16.66. Carapace 7.38 long, 5.97 wide; chelicerae 33.60 long; abdomen 9.28 long, 6.99 wide. Spinnerets: PMS, 1.02 long, 0.38 wide, 0.17 apart; PLS, 1.09 basal, 0.31 middle, 0.21 distal; midwidths 0.99, 0.77, 0.36 respectively; 1.61 total length. Colour in alcohol. Carapace, patellae, tibiae, metatarsi and tarsi of legs I���II and patella and tarsus of palp blackish-brown, other leg parts reddish-brown. Abdomen dorsally grayish-brown, mottled with yellowish spots; ventrally uniformly yellowish-brown. AME encircled with black patch. Carapace (Fig. 4 A). Glabrous, broader anteriorly (widest between legs II) and gradually narrowing posteriorly, striae prominent; fovea, procurved, deep. Bristles: 2 long and 2 short on caput; 2 short between AME- ALE; 3 short on clypeus edge. Eyes (Fig. 4 B). Eight in three rows, ALE situated far from AME on clypeal edge; posterior row procurved. Ocular group 1.75 long, 1.45 wide; MOQ not square, 0.64 front and 0.82 back width, 0.69 long. Diameter AME 0.22, PME 0.17, ALE 0.25, PLE 0.24; distance between ALE-AME 0.62, AME-AME 0.05, PME-PME 0.06, ALE- PLE 0.90, PLE-PME 0.43 and ALE-ALE adjacent. Maxillae (Fig. 4 C). 2.30 long anteriorly, 2.87 long posteriorly, 1.68 wide; ca. 50 cuspules distributed towards anterior edge; anterior lobe distinct. Labium (Fig. 4 C). 0.99 long, 1.37 wide, shallow labiosternal groove, slightly procurved, 1 large and 4 small cuspules on anterior edge. Chelicerae (Figs 4 C���D). 9 promarginal teeth, 8 retomarginal teeth; rastellum strong, raised on high triangular mound, consist of 16 thick, short spines with many bristles-like spines; two glabrous bands for length of dorsal surface of chelicerae. Sternum (Fig. 4 C). Broader between posterior coxae; reddish-brown, elevated in centre, sloping laterally, covered with long black hair; row of long bristles on margins, posterior angle blunt. Sigilla (Fig. 4 C). Posterior sigilla absent; median pair submarginal, 2.04 apart, 0.30 from margin and anterior pair round, marginal. Legs. Leg III thicker than other legs; femora III clearly wider than others; metatarsi of all legs longer than tarsi. Legs cylindrical except for metatarsi and tarsi of legs I���II and tibiae and tarsi of palp, dorsoventrally flattened, covered with few scattered hair, bristles and few curved, thick, thorn-like spines. Two conspicuous glabrous bands for length of femora, patellae and tibiae. Scopulae absent on all legs and palp. Leg formula 4123. Spines. More on promarginal and retromarginal sides of legs and palp. I: ti, p= 18, r= 22; mt, p= 22, v= 2, r= 19; ta, p= 8, v= 3, r= 10. II: ti, p= 10, v= 8, r= 5; mt, p= 11, v= 7, r= 9; ta, p= 7, v= 3, r= 4. III: pa, p= 9, r= 2; ti, p= 9, v= 2, r= 7; mt, p= 10, v= 6, r= 7; ta, v= 10. IV: pa, p= 11; ti, v= 2; mt, v= 7; ta, v= 9. Palp: pa, v= 1; ti, p= 11, r= 7; ta, p= 14, v= 5, r= 10. Trichobothria. Clavate absent; ta I, 12 long filiform in each of two zig-zag rows for basal 3 / 4 th; ta II, 9 long filiform; ta III���IV, 10 long filiform; palp, 22 short filiform for length. Mt I, 13 short filiform; mt II, 7 short filiform; mt III, 9 filiform; mt IV, 16 short filiform. Trichobothria on tarsi in two zig-zag rows in basal 3 / 4 th and trichobothria on metatarsi for length. Leg coxae. Reddish-brown; coxae I���IV covered with long black bristles, more dense on anterior pairs. Coxae III���IV with central patch glabrous, others sparsely covered with long and short bristles. Coxae IV clearly broader than others, anterior edge curved. Claws. Paired claw on all legs with single tooth and palp with bifid tooth; unpaired claw on leg I���IV; all claws (paired as well as unpaired) leg IV distinctly larger in size than claws on leg I���III. False claw tufts on each side of paired claws. Abdomen (Fig. 4 A). Dorsum, grayish-brown heavily mottled with yellowish spots; uniformly covered with short and long black hairs, cuticle appears leathery and slightly rough. Venter uniformly yellowish-brown, evenly covered with short black hairs, intermixed with few long black hairs. Spinnerets. PMS digitiform covered with brown hair; PLS covered with brown hair, apical segment domed. Spermathecae (Fig. 4 E). Two single lobes facing away from each other; each lobe, distally bulge with notch at tip, resembling teats in mammals, sclerotized, covered with pores, otherwise lobes transparent, gradually narrowing (cone shape) and opens ventrally; transparent sheet of inverted triangular shape covers basal half of lobes. Natural history. The spider was found on a roadside cutting in a deciduous sacred grove. It was found sympatric with Heligmomerus barkudensis (Gravely, 1921) in Kapilas, Odhisa. Only a single female was found and so their population seems to be low as compared to H. barkudensis. More information on habitat can be seen in Siliwal et al. 2010. The burrow of this species was simple tube-like with thick silk linning, measuring about 15.0mm in diameter and 55mm in depth. Burrow was having vertical on the ground with a gentle slope. Like Heligmomerus barkudensis, entrance of the burrow had a hinge door (diameter 13.5mm), inside of which covered with thick layer of silk with claw markings in the center and outer side covered with soil particles and debris., Published as part of Gupta, Neha, Ganeshkumar, M., Das, Sanjay Keshari & Siliwal, Manju, 2013, Three new species of Idiops Perty, 1833 (Araneae: Idiopidae) from India, pp. 237-250 in Zootaxa 3635 (3) on pages 247-249, DOI: 10.11646/zootaxa.3635.3.3, http://zenodo.org/record/216039