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Batrisodes hanmi sp. nov. (Figs 39, 81A, 98C, D) Chinese common name: 汗ṁqà甲 Type material (4 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, Hanmi (ca. 29°21’53”N, 95°07’46”E), alt. 2200 m; 19.viii.2005, Liang Tang leg. [ƱẪẸṘ汗ṁ]’ (SNUC). PARATYPE: CHINA: 2 ♂♂, 1 ♀, same data as that of holotype (SNUC). Diagnosis. Male. Body length 2.50–2.58 mm. Head sub-rectangular at base, approximately as broad as pronotum; head and antenna lacking modifications. Pronotum with five dorsally visible longitudinal sulci, discal sulci slightly curved at middle, one pair of marginal sulci present ventral to lateral expansion. Elytron with short discal stria. Protibia with tiny apical denticle, mesotrochanter with distinct ventral spine, metatrochanter with small ventral tooth, metatibia with blunt apical projection and long bunch of setae at apex. Aedeagus strongly asymmetric; median lobe with large basal capsule and foramen, dorsal lobe well-developed, with apex deeply forked, split to two lobes. Female. Body length approximately 2.5 mm; legs lacking spine or projection, genitalia as in Fig. 39I. Description. Male. Body (Fig. 39A) length 2.50–2.58 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence. Head (Fig. 39B) sub-rectangular at base, slight wider than long, length 0.50–0.51 mm, width across eyes 0.55– 0.57 mm; vertex roughly punctate, with large, asetose vertexal foveae (dorsal tentorial pits), with U-shaped sulcus connecting foveae, lateral vertexal carina faint, extending from occipital constriction to level of anterior margin of eyes, mediobasal carina short, extending anteriorly to level of middle of eyes; antennal tubercles moderately raised; frons impressed at middle, confluent with clypeus; clypeus with rough surface, its entire anterior margin carinate and raised; ocular-mandibular carina complete, distinct. Venter with granulate surface; small gular foveae (posterior tentorial pits) originating from shared opening, with distinct median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 ommatidia. Antenna lacking distinct club and modification, length 1.16–1.27 mm; antennomere 1 thick, subcylindrical, 2–7 each elongate, 8 smallest, 9 much larger than 8, 10 slightly broader but shorter than 9, 11 largest, longer than 9 and 10 combined, subconical. Pronotum (Fig. 39B) approximately as long as wide, length 0.57 mm, width 0.58 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, antebasal tubercles small, lacking discal spines, with one median, one pair of discal and one pair of lateral longitudinal sulci, discal sulci curved at middle; with one pair of marginal sulci beneath lateral expansions; with small asetose median pit and large, setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, well-demarcated from hypomera by pair of lateral carinae, which extend posterolaterally to form complete hypomeral ridges; lateral procoxal foveae distinct; anterior and lateral margin of coxal cavity carinate. Elytra wider than long, length 0.77–0.78 mm, width 0.89–0.96 mm; each elytron with three large, asetose basal foveae, inner two close; discal stria only broadly and shallowly present for short distance posterior to outer basal fovea; small subhumeral fovea present, with carinate marginal stria extending from fovea to posterior margin of elytron. Mesoventrite short, demarcated from metaventrite by thin carinae; median mesoventral foveae narrowly separated, lateral mesoventral foveae deeply forked internally. Metaventrite moderately impressed at middle, with welldeveloped lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Legs more or less robust. Protrochanter simple, protibia with tiny apical spur; mesotrochanter with distinct, slightly curved ventral spine (Fig. 39C); metatrochanter with tiny ventral tooth (Fig. 39D; note: the male paratype has ventral margin smooth and lacks this tooth, which was treated as an intraspecific variation), metatibia (Fig. 39E) with dense golden setae along mesal margin of apical 1/2, apex with small, blunt projection and elongate bunch of long setae. Abdomen widest at lateral margins of tergite 1 (IV), length 0.63–0.67 mm, width 0.85–0.89 mm. Tergite 1 (IV) approximately twice as long as 2 (V), with pair of short, triangular discal carinae, basal sulcus separated by one pair of mediobasal and two pairs of basolateral foveae, with thick marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) as long as tergites 2 and 3 combined, with pair of small tubercles at lateral margins; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular. Sternite 2 (IV) with one pair of small mediobasal and three pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly impressed, sternite 7 (IX) (Fig. 39F) with asymmetric, membranous basal part and semicircular, well-sclerotized apical part. Aedeagus (Fig. 39G, H) 0.67 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection relatively long, middle and apical portion of ventral stalk each with one long seta; dorsal lobe well-developed, apically narrowed and deeply forked, split into two lobes. Female. Similar to male in external morphology; antenna approximately as long as that of male; legs lacking spines or projections; each compound eye composed of approximately 12 ommatidia. Measurements (as for male): body length 2.47 mm; length/width of head 0.51/ 0.59 mm, pronotum 0.56/ 0.56 mm, elytra 0.77/ 0.93 mm; abdomen 0.74/ 0.87 mm; length of antenna 1.19 mm; genitalia (Fig. 39I) moderately sclerotized, rounded triangular, maximum width 0.38 mm. Comparative notes. The new species is morphologically similar to B. guoguotang by sharing most external characters and may be separated from the latter by the following characters: 1) slightly curved discal longitudinal sulci of the pronotum (strongly curved in B. guoguotang), 2) a large ventral spine of mesotrochanter (with small ventral denticle in B. guoguotang), 3) a small ventral tooth of the metatrochanter (large ventral projection in B. guoguotang), and 4) the apical portion of aedeagal dorsal lobe split into two lobes (split into three lobes in B. guoguotang). Distribution. Mêdog County, Tibet, SW China (Figs 81A, 98C, D). Etymology. The specific name refers to the type locality of the new species, i. e., Hanmi., Published as part of Yin, Zi-Wei, 2022, The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth's " Third Pole " (Coleoptera, Staphylinidae, Pselaphinae), pp. 1-211 in Zootaxa 5111 (1) on pages 92-94, DOI: 10.11646/zootaxa.5111.1.1, http://zenodo.org/record/6340729

Batrisodes Reitter, 1882 Chinese common name: qà甲¤ Batrisodes Reitter, 1882: 134; Park 1947 (North American species), 1948 (checklist of world species); Chandler 1997 (catalog of North American species); Besuchet 1981 (Palaearctic species); Newton & Chandler 1989 (discussion of type species, catalog); Nomura 2007 (Japanese species). Type species: Batrisus delaporti Aubé, 1833: 509 (subsequent designation by Lucas, 1920: 135). = Alytus Hampe, 1863: 285. Type species: Trichonyx adnexus Hampe, 1863: 285 (monotypy). Nomen oblatum, see discussion in Newton & Chandler 1989: 8. = Batrisodellus Jeannel, 1958: 37. Type species: Batrisodes nipponensis Raffray, 1909: 23 (original designation). Synonymized by Yin et al. 2015a: 53. In China, the genus is represented by 21 species (e. g., Nomura 2007; Yin & Nomura 2011; Yin et al. 2015a; Jiang & Yin 2016, 2017; Zhang & Yin 2021), with most of these described only recently. Some of the Chinese species are known inquilines of a wide range of ant genera including Aphaenogaster Mayr, Brachyponera Emery, Formica Linné, Lasius Fabricius, Myrmica Latreille, Nylanderia Emery, Odontomachus Latreille, Pheidole Westwood, and Pseudolasius Emery (Yin & Li 2013b; Yin & Shen 2015; Jiang & Yin 2016, 2017). Batrisodes guoguotang was collected by sifting leaf litter, and there is no information on the biology of B. hanmi. The type material of Batrisodes pruinosus Reitter (a female) was collected by Russian ethnographer Grigory Potanin from Amdo (ȐƷ) (Besuchet 1981), an area historically in a broader Tibet, but now belonging to Qinghai Province. This species is here excluded from the Tibetan fauna. The following key is provided to help with the separation of the two Tibetan species. Key to Tibetan species of Batrisodes (male) 1 Central part of vertex finely punctate (Fig. 38B); metatrochanter with strongly curved ventral projection (Fig. 38C); dorsal lobe of aedeagus split into three lobes near apex (Fig. 38F)....................................... B. guoguotang sp. nov. - Central part of vertex coarsely punctate (Fig. 39B); metatrochanter with slightly curved ventral spine (Fig. 38C); dorsal lobe of aedeagus split into two lobes near apex (Fig. 39H)............................................... B. hanmi sp. nov., Published as part of Yin, Zi-Wei, 2022, The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth's " Third Pole " (Coleoptera, Staphylinidae, Pselaphinae), pp. 1-211 in Zootaxa 5111 (1) on pages 89-90, DOI: 10.11646/zootaxa.5111.1.1, http://zenodo.org/record/6340729, {"references":["Reitter, E. (1882) Neue Pselaphiden und Scydmaeniden aus Brasilien. Deutsche Entomologische Zeitschrift, 26 (1), 129 - 152, pl. 5.","Park, O. (1947) Observations on Batrisodes (Coleoptera: Pselaphidae), with particular reference to the American species east of the Rocky Mountains. Bulletin of the Chicago Academy of Sciences, 8 (3), 45 - 132, pls. 1 - 11.","Chandler, D. S. (1997) A catalog of the Coleoptera of America north of Mexico. Family: Pselaphidae. United States Department of Agriculture, Agriculture Handbook 529 - 31. United States Department of Agriculture, Washington, D. C., ix + 118 pp.","Besuchet, C. (1981) Contribution a l'etude des Batrisodes palearctiques (Coleoptera: Pselaphidae). Revue Suisse de Zoologie, 88 (1), 275 - 296. https: // doi. org / 10.5962 / bhl. part. 82372","Newton, A. F. Jr. & Chandler, D. S. (1989) World catalog of the genera of Pselaphidae (Coleoptera). Fieldiana, New Series, Zoology, 53, 1 - 93. https: // doi. org / 10.5962 / bhl. title. 3209","Nomura, S. (2007) Two new species of the genus Batrisodes (Coleoptera, Staphylinidae, Pselaphinae) from Taiwan. Elytra, 35 (1), 76 - 84.","Aube, C. (1833) Note sur la famille des Pselaphiens. Annales de la Societe Entomologique de France, 2, 502 - 511.","Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Archiv fur Naturgeschichte, 84 (A), i - xxxi + 1 - 696 pp. [1918]","Hampe, C. (1863) Ein kleiner Beitrag zur gross-osterreichischen Kaferfauna. Wiener Entomologische Monatschrift, 7 (8), 285 - 290.","Jeannel, R. (1958) Revision des Pselaphides du Japon. Memoires du Museum National d'Histoire Naturelle, A: Zoologie, 18 (1), 1 - 138.","Raffray, A. (1909) Nouvelles especes de Pselaphides. Annales de la Societe Entomologique de France, 78, 15 - 52.","Jiang, R. - X. & Yin, Z. - W. (2016) Two new species of Batrisodes Reitter (Coleoptera: Staphylinidae: Pselaphinae) from China. Zootaxa, 4205 (2), 194 - 200. https: // doi. org / 10.11646 / zootaxa. 4205.2.9","Jiang, R. - X. & Yin, Z. - W. (2017) Eight new species and two new records of Batriscenellus Jeannel (Coleoptera: Staphylinidae: Pselaphinae) from China and India. Zootaxa, 4318 (3), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4318.3.8","Zhang, W. - X. & Yin, Z. - W. (2021) Two new species of Batrisodes Reitter from Shaanxi, China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4938 (2), 243 - 250. https: // doi. org / 10.11646 / zootaxa. 4938.2.5","Yin, Z. - W. & Li, L. - Z. (2013 b) Batrisodes (Excavodes) simplex sp. n. (Coleoptera: Staphylinidae: Pselaphinae): a second species of the genus from continental China. Deutsche Entomologische Zeitschrift, 60 (2), 261 - 266. https: // doi. org / 10.1002 / mmnd. 201300033","Yin, Z. - W., Shen, J. - W. & Li, L. - Z. (2015 a) New species and new combinations of Asian Batrisodes Reitter (Coleoptera, Staphylinidae, Pselaphinae), and synonymy of Batrisodellus Jeannel with Batrisodes. Deutsche Entomologische Zeitschrift, 62 (1), 45 - 54. https: // doi. org / 10.3897 / dez. 62.4539"]}

Batrisodes guoguotang sp. nov. (Figs 38, 81A, 99D, G) Chinese common name: 果果Ƌqà甲 Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, forest nr. Guoguotang Great Bend, 29°18’39.35”N, 95°16’25.75”E, 1180 m; 2019.vii.22, leaf litter, Z.-W. Yin leg. [ƱẪẸṘ果果ḙ]’ (SNUC). Diagnosis. Male. Body length 2.37 mm. Head sub-rectangular at base, approximately as broad as pronotum; head and antenna lacking modifications. Pronotum with five dorsally visible longitudinal sulci, discal sulci strongly curved at middle, one pair of marginal sulci present ventral to lateral expansion. Elytron with short discal stria. Mesofemur and mesotibia simple; metatrochanter with large ventral projection, metatibia with blunt apical projection and long bunch of setae at apex. Aedeagus strongly asymmetric; median lobe with large basal capsule and foramen, dorsal lobe well-developed, with narrowed apex deeply forked, split to three lobes. Description. Male. Body (Fig. 38A) length 2.37 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence. Head (Fig. 38B) sub-rectangular at base, slight wider than long, length 0.52 mm, width across eyes 0.55 mm; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), with U-shaped sulcus connecting foveae, with sparse large punctures along outer margin of sulcus, lateral vertexal carina faint, extending from occipital constriction to level of middle of eyes, mediobasal carina as long as lateral ones; antennal tubercles moderately raised; frons impressed at middle, confluent with clypeus; clypeus with rough surface, its entire anterior margin carinate and raised; ocular-mandibular carina indistinct. Venter with granulate surface; small gular foveae (posterior tentorial pits) originating from shared opening, with distinct median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 20 large ommatidia. Antenna lacking distinct club and modification, length 1.24 mm; antennomere 1 thick, subcylindrical, 2–7 each elongate, gradually shorter, 8 smallest, 9 much larger than 8, 10 slightly broader than 9, 11 largest, as long as 9 and 10 combined, subconical. Pronotum (Fig. 38B) approximately as long as wide, length 0.50 mm, width 0.53 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, antebasal tubercles small, lacking discal spines, with one median, one pair of discal and one pair of lateral longitudinal sulci, discal sulci strongly curved at middle; with one pair of marginal sulci beneath lateral expansions; with small asetose median pit and large, setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, well-demarcated from hypomera by pair of lateral carinae, which extend posterolaterally to form complete hypomeral ridges; lateral procoxal foveae small; anterior and lateral margin of coxal cavity carinate. Elytra wider than long, length 0.69 mm, width 0.86 mm; each elytron with three large, asetose basal foveae, inner two close; discal stria only broadly and shallowly present for short distance posterior to outer basal fovea; small subhumeral fovea present, with carinate marginal stria extending from fovea to posterior margin of elytron. Mesoventrite short, demarcated from metaventrite by thin carinae; median mesoventral foveae narrowly separated, lateral mesoventral foveae deeply forked internally. Metaventrite moderately impressed at middle, with welldeveloped lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Legs more or less robust, lacking distinct modifications except for metatrochanter. Pro- and mesotrochanter each with tiny tubercle on ventral margin, pro- and mesotibia with small apical denticle; metatrochanter (Fig. 38C) with large, curved ventral projection, metatibia (Fig. 38D) with dense golden setae along mesal margin of apical 1/2, apex with small, blunt projection and elongate bunch of long setae. Abdomen widest at lateral margins of tergite 1 (IV), length 0.68 mm, width 0.79 mm. Tergite 1 (IV) approximately twice as long as 2 (V), with pair of short, triangular discal carinae, basal sulcus separated by one pair of mediobasal and two pairs of basolateral foveae, with thick marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) as long as 2 and 3 combined, with pair of small tubercles at lateral margins; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular. Sternite 2 (IV) with one pair of small mediobasal and three pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly impressed, sternite 7 (IX) (Fig. 38E) with asymmetric, membranous basal part and semicircular, well-sclerotized apical part. Aedeagus (Fig. 38F, G) 0.52 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk with two long preapical setae; dorsal lobe well-developed, apically narrowed and deeply forked. Female. Unknown. Comparative notes. Batrisodes guoguotang is apparently allied with B. hanmi by sharing most external characters, as well as a similar form of the aedeagus. The new species may be separated from B. hanmi by the following characters: 1) more strongly curved discal longitudinal sulci of the pronotum, 2) a smaller ventral denticle of the mesotrochanter (much larger ventral spine in B. hanmi), 3) a large projection on the ventral margin of the metatrochanter (small tubercle in B. hanmi), and 4) the apical portion of the aedeagal dorsal lobe split into with three lobes (split into two lobes in B. hanmi). Distribution. Mêdog County, Tibet, SW China (Figs 81A, 99D, G). Etymology. The new species is named after the Guoguotang Great Bend of Yarlung Tsangpo Grand Canyon located near Mêdog County., Published as part of Yin, Zi-Wei, 2022, The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth's " Third Pole " (Coleoptera, Staphylinidae, Pselaphinae), pp. 1-211 in Zootaxa 5111 (1) on pages 90-92, DOI: 10.11646/zootaxa.5111.1.1, http://zenodo.org/record/6340729

Batrisodes grandiceps Zhang & Yin, sp. nov. (Figs 1, 2) Type material. HOLOTYPE: CHINA: ♂, ‘ China: Shaanxi, Ningshan County (ṪȐfl), Pingheliang Forest Park (ỸOiẌÑẎƙfin'ẘ), 33°28′14″N, 106°29′29″E, 13.vii.2012, 2050– 2150 m, mixed leaf litter, sifted, Y.-H. Pan leg.’ (SNUC). PARATYTYPES: CHINA: 4 ♂♂, 3 ♀♀, same label data as for holotype (SNUC); 1 ♂, 1 ♀, same label data as for holotype, except ‘ 12.vii.2012,’ (SNUC); 3 ♂♂, 2 ♀♀, ‘ China: Shaanxi, Ningshan County, Huoditang Forestry Centre (Ẋḿffifiá), 33°26′N, 108°27′E, 12.vii.2012, 1500–1700 m, mixed leaf litter, sifted, CHEN & LI leg.’ (SNUC). Diagnosis. Male. Body length 2.29–2.35 mm. Head base rounded, much wider than pronotum; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, sides of projection more or less parallel in dorsal view; antennomere 3 larger than antennomere 2. Pronotum finely punctate, with four distinct discal teeth. Mesofemur with long ventral spine near middle, mesotibia expanded for apical 2/5 of tibial length, with blunt projection before middle. Aedeagus dorso-ventrally symmetric, with symmetric, strongly sclerotized endophallus. Female. Similar to male in external morphology, lacking modifications on head and legs, antennomere 3 same size of antennomere 2; may be identified only by geographical association with male. Description. Male (Fig. 1A). Body reddish brown, length from anterior margin of clypeus to apex of abdomen 2.29–2.35 mm. Head (Fig. 2A) slight wider than long, length from anterior margin of clypeus to head base (excluding occipital constriction) 0.46–0.48 mm, width across eyes 0.53–0.55 mm; surface roughly punctate, with large, nude vertexal foveae; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, sides of projection more or less parallel in dorsal view, clypeal anterior margin rounded; lateral vertexal carina faint, extending from occipital constriction to level of anterior margin of fovea, mediobasal carina as long as lateral ones. Eyes relatively small, each composed of approximately 35 facets. Antenna lacking distinct club, antennomere 1 (scape) thick, subcylindrical, antennomeres 2–10 moniliform, antennomere 3 distinctly larger than 2, antennomeres 4–8 of similar width, antennomere 9 much wider than 8, antennomeres 9 and 10 successively broader, each strongly transverse, antennomere 11 largest, subconical, lacking hook-like spine. Pronotum slightly longer than wide, length along midline 0.48–0.50 mm, maximum width 0.45 mm; sides rounded, widest near middle, disc slightly convex, finely punctate, with four distinct teeth; median and lateral longitudinal sulci distinct; with small nude median and setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Elytra wider than long, length along suture 0.60–0.62 mm, maximum width 0.72–0.75 mm; each elytron with three large, nude basal foveae, inner two close; discal striae shallow and short, extending posteriorly for 1/3 of elytral length. Mesofemur (Fig. 2C) with long ventral spine near middle, mesotibia expanded for apical 1/3 of tibial length, with blunt ventral projection at apical 2/5, apex of projection rounded, with dense short setae. Abdomen widest at lateral margins of tergite 1 (IV), length of dorsally exposed part along midline 0.73–0.77 mm, maximum width 0.72 mm; tergite 1 (IV) with strongly oblique marginal carinae; discal carinae thick, tergite 5 (VIII) (Fig. 2D) semicircular; sternite 6 (VIII) (Fig. 2E) transverse. Aedeagus (Fig. 2F, G) 0.41 mm long, median lobe dorso-ventrally symmetrical and flattened, broadest near apex, apical part narrowing toward apex and curved ventrally in lateral view; endophallus well-sclerotized, lateral lobes broad at base and evenly narrowing apically. Female (Fig. 1B). General morphology similar to male, lacking modifications on head (Fig. 2B) and legs; antennomere 3 same size of antennomere 2; each eye composed of about 12 facets; genitalia (Fig. 2H) symmetric, weakly sclerotized, 0.21 mm wide. Measurements of body parts (as for male): body length 2.25–2.31mm, length/ width of head 0.46–0.48 mm / 0.52–0.53 mm, pronotum 0.48–0.50 mm / 0.44–0.46 mm, elytra 0.58 mm / 0.74 mm, abdomen 0.73–0.75 mm / 0.74–0.75 mm. Comparative notes. The new species is apparently most close to B. grossepunctatus sp. nov. described below by the males having a similar position and shape of the cephalic modifications and spination of the legs, as well as the presence of a well-sclerotized endophallus of the aedeagus. They can be clearly separated by the smaller body size (2.29–2.35 mm vs. 2.41–2.48 mm), somewhat parallel sides of the projection on the clypeus, the finely punctate pronotal disc, and the different shape of the lateral lobes of the aedeagal endophallus of the new species. Distribution. China: Shaanxi. Etymology. The new specific epithet refers to the large head of the new species.

Batrisodes grossepunctatus Zhang & Yin, sp. nov. (Figs 3, 4) Type material. HOLOTYPE: CHINA: ♂, ��� China: Shaanxi, Nanzheng County (��fflfl), Liping Forest Park (= ����������fin'���), 32��50���20���N, 106��36���18���E, 15.vii.2012, 1400���1600m, mixed leaf litter, sifted, Y.-H. Pan leg.��� (SNUC). PARATYPES: CHINA: 1 ♂, 3 ♀♀, same label data as for holotype (SNUC); 1 ♂, same label data as for holotype, except ��� 16.vii.2012, Li Li-Zheng leg.��� (SNUC). Diagnosis. Male. Body length 2.41���2.48 mm. Head base rounded, wider than pronotum; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, sides of projection narrowing posteriorly from apical 1/ 3 in dorsal view. Pronotum roughly punctate, with four distinct discal teeth. Mesofemur with long ventral spine near middle, mesotibia expanded for apical 1/3 of tibial length, with blunt projection before middle. Aedeagus dorso-ventrally symmetric, with symmetric, well-sclerotized endophallus. Female. Similar to male in external morphology, lacking modifications on head and legs; may be identified only by geographical association with male. Description. Male (Fig. 3A). Body reddish brown, length from anterior margin of clypeus to apex of abdomen 2.41���2.48 mm. Head (Fig. 4A) slight wider than long, length from anterior margin of clypeus to head base (excluding occipital constriction) 0.52���0.54 mm, width across eyes 0.57���0.59 mm; surface roughly punctate, with large, nude vertexal foveae; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, margins of projection narrowing posteriorly from apical 1/ 3 in dorsal view. Clypeal anterior margin rounded; lateral vertexal carina faint, extending from occipital constriction to level of posterior margin of eyes, mediobasal carina weakly extending from head base to level of anterior margin of eyes. Eyes relatively small, each composed of approximately 40 facets. Antenna lacking distinct club, antennomere 1 (scape) thick, subcylindrical, antennomeres 2���10 moniliform, antennomeres 2���8 of similar width, antennomere 9 wider than 8, antennomeres 9 and 10 successively broader, each strongly transverse, antennomere 11 largest, subconical, lacking hook-like spine. Pronotum approximately as long as wide, length along midline 0.50 mm, maximum width 0.49 mm; sides rounded, widest near the middle, disc slightly convex, roughly punctate, with four distinct teeth; median longitudinal sulci weekly present; lateral longitudinal sulci distinct; with small nude median and setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Elytra wider than long, length along suture 0.64���0.65 mm, maximum width 0.78���0.81 mm; each elytron with three large, nude basal foveae, inner two close; discal striae shallow and short, extending posteriorly for 1/3 of elytral length. Mesofemur (Fig. 4C) with thin but distinct ventral spine near middle, mesotibia expanded for apical 1/3 of tibial length, with blunt ventral projection at apical 2/5, apex of projection rounded, with dense short setae. Abdomen widest at lateral margins of tergite 1 (IV), length of dorsally exposed part along midline 0.74���0.79 mm, maximum width 0.75���0.79 mm; tergite 1 (IV) with strongly oblique marginal carinae, discal carinae thick, tergite 5 (VIII) (Fig. 4D) semicircular; sternite 6 (VIII) (Fig. 4E) transverse. Aedeagus (Fig. 4F, G) 0.45 mm long, median lobe dorso-ventrally symmetrical and flattened, broadest near apex, apical part narrowing toward apex and curved ventrally in lateral view; endophallus well-sclerotized, lateral lobes broad at bases, abruptly constricted at middle and then evenly narrowing apically. Female (Fig. 3B). General morphology similar to male, lacking modifications on head (Fig. 4B) and legs; each eye composed of about 20 facets; genital complex (Fig. 4H) symmetrical, weakly sclerotized, 0.27 mm wide. Measurements of body parts (as for male): body length 2.52���2.53 mm, length/width of head 0.51���0.52 mm / 0.57 mm, pronotum 0.52���0.53 mm / 0.49���0.50 mm, elytra 0.66���0.67 mm / 0.80���0.81 mm, abdomen 0.80���0.84 mm / 0.80 mm. Comparative notes. The new species is morphologically most similar to B. grandiceps sp. nov. described above by the male having a similar position and shape of the cephalic modifications and spination of the legs, as well as the presence of a well-sclerotized endophallus of the aedeagus. They can be clearly separated by the larger body size (2.41���2.48 mm vs. 2.29���2.35 mm), the posteriorly narrowing sides of the projection on the clypeus, the roughly punctate pronotal disc, and the different shape of the lateral lobes of the aedeagal endophallus of the new species. The modified male head is not an uncommon character state in Asian Batrisodes. However, a roundly and deeply excavated anterior margin of the frons occurs only in B. babaianus Nomura distributed in Meifeng (������), Taiwan. The two new species described here can be readily separated from B. babaianus by the presence of a large projection on the clypeus, which is lacking in the latter species. Distribution. China: Shaanxi. Etymology. The new specific epithet refers to the roughly punctate pronotum of the new species., Published as part of Zhang, Wen-Xuan & Yin, Zi-Wei, 2021, Two new species of Batrisodes Reitter from Shaanxi, China (Coleoptera Staphylinidae: Pselaphinae), pp. 243-250 in Zootaxa 4938 (2) on pages 247-248, DOI: 10.11646/zootaxa.4938.2.5, http://zenodo.org/record/4563882

Batrisodes grandiceps Zhang & Yin, sp. nov. (Figs 1, 2) Type material. HOLOTYPE: CHINA: ♂, ��� China: Shaanxi, Ningshan County (�����fl), Pingheliang Forest Park (���Oi����������fin'���), 33��28���14���N, 106��29���29���E, 13.vii.2012, 2050��� 2150 m, mixed leaf litter, sifted, Y.-H. Pan leg.��� (SNUC). PARATYTYPES: CHINA: 4 ♂♂, 3 ♀♀, same label data as for holotype (SNUC); 1 ♂, 1 ♀, same label data as for holotype, except ��� 12.vii.2012,��� (SNUC); 3 ♂♂, 2 ♀♀, ��� China: Shaanxi, Ningshan County, Huoditang Forestry Centre (������ffifi��), 33��26���N, 108��27���E, 12.vii.2012, 1500���1700 m, mixed leaf litter, sifted, CHEN & LI leg.��� (SNUC). Diagnosis. Male. Body length 2.29���2.35 mm. Head base rounded, much wider than pronotum; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, sides of projection more or less parallel in dorsal view; antennomere 3 larger than antennomere 2. Pronotum finely punctate, with four distinct discal teeth. Mesofemur with long ventral spine near middle, mesotibia expanded for apical 2/5 of tibial length, with blunt projection before middle. Aedeagus dorso-ventrally symmetric, with symmetric, strongly sclerotized endophallus. Female. Similar to male in external morphology, lacking modifications on head and legs, antennomere 3 same size of antennomere 2; may be identified only by geographical association with male. Description. Male (Fig. 1A). Body reddish brown, length from anterior margin of clypeus to apex of abdomen 2.29���2.35 mm. Head (Fig. 2A) slight wider than long, length from anterior margin of clypeus to head base (excluding occipital constriction) 0.46���0.48 mm, width across eyes 0.53���0.55 mm; surface roughly punctate, with large, nude vertexal foveae; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, sides of projection more or less parallel in dorsal view, clypeal anterior margin rounded; lateral vertexal carina faint, extending from occipital constriction to level of anterior margin of fovea, mediobasal carina as long as lateral ones. Eyes relatively small, each composed of approximately 35 facets. Antenna lacking distinct club, antennomere 1 (scape) thick, subcylindrical, antennomeres 2���10 moniliform, antennomere 3 distinctly larger than 2, antennomeres 4���8 of similar width, antennomere 9 much wider than 8, antennomeres 9 and 10 successively broader, each strongly transverse, antennomere 11 largest, subconical, lacking hook-like spine. Pronotum slightly longer than wide, length along midline 0.48���0.50 mm, maximum width 0.45 mm; sides rounded, widest near middle, disc slightly convex, finely punctate, with four distinct teeth; median and lateral longitudinal sulci distinct; with small nude median and setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Elytra wider than long, length along suture 0.60���0.62 mm, maximum width 0.72���0.75 mm; each elytron with three large, nude basal foveae, inner two close; discal striae shallow and short, extending posteriorly for 1/3 of elytral length. Mesofemur (Fig. 2C) with long ventral spine near middle, mesotibia expanded for apical 1/3 of tibial length, with blunt ventral projection at apical 2/5, apex of projection rounded, with dense short setae. Abdomen widest at lateral margins of tergite 1 (IV), length of dorsally exposed part along midline 0.73���0.77 mm, maximum width 0.72 mm; tergite 1 (IV) with strongly oblique marginal carinae; discal carinae thick, tergite 5 (VIII) (Fig. 2D) semicircular; sternite 6 (VIII) (Fig. 2E) transverse. Aedeagus (Fig. 2F, G) 0.41 mm long, median lobe dorso-ventrally symmetrical and flattened, broadest near apex, apical part narrowing toward apex and curved ventrally in lateral view; endophallus well-sclerotized, lateral lobes broad at base and evenly narrowing apically. Female (Fig. 1B). General morphology similar to male, lacking modifications on head (Fig. 2B) and legs; antennomere 3 same size of antennomere 2; each eye composed of about 12 facets; genitalia (Fig. 2H) symmetric, weakly sclerotized, 0.21 mm wide. Measurements of body parts (as for male): body length 2.25���2.31mm, length/ width of head 0.46���0.48 mm / 0.52���0.53 mm, pronotum 0.48���0.50 mm / 0.44���0.46 mm, elytra 0.58 mm / 0.74 mm, abdomen 0.73���0.75 mm / 0.74���0.75 mm. Comparative notes. The new species is apparently most close to B. grossepunctatus sp. nov. described below by the males having a similar position and shape of the cephalic modifications and spination of the legs, as well as the presence of a well-sclerotized endophallus of the aedeagus. They can be clearly separated by the smaller body size (2.29���2.35 mm vs. 2.41���2.48 mm), somewhat parallel sides of the projection on the clypeus, the finely punctate pronotal disc, and the different shape of the lateral lobes of the aedeagal endophallus of the new species. Distribution. China: Shaanxi. Etymology. The new specific epithet refers to the large head of the new species., Published as part of Zhang, Wen-Xuan & Yin, Zi-Wei, 2021, Two new species of Batrisodes Reitter from Shaanxi, China (Coleoptera Staphylinidae: Pselaphinae), pp. 243-250 in Zootaxa 4938 (2) on pages 244-246, DOI: 10.11646/zootaxa.4938.2.5, http://zenodo.org/record/4563882

Continuing surveys on the pselaphine fauna in China have revealed four undescribed species of the genus Batrisodes: Batrisodesangustelytratussp. n. (with Pachycondyla ant) and Batrisodeslongwangshanussp. n. from Zhejiang, Batrisodesbamiansp. n. (with Pseudolasius ant) from Hunan, and Batrisodessongxiaobinisp. n. (with Formica ant) from Yunnan. The genus Batrisodellus is placed as a junior synonym of Batrisodes, syn. n.; this taxonomic act results in eight new combinations:Batrisodescerberus, Batrisodescoprea, Batrisodescristatus, Batrisodesfengtingae, Batrisodeslaticollis, Batrisodeslongulus, Batrisodespalpalis, and Batrisodespunctipennis. The Russian Far East species Basitrodescornutus is transferred to Batrisodes, comb. n.

Eight new species of the genus Batrisodes Reitter are described from continental China, seven of which were found in association with ants: B. abdominalis sp. n. and B. tianmuensis sp. n. with an Ectomomyrmex ant from Zhejiang; B. grossus sp. n. with an Odontomachus ant from Guangxi; B. simianshanus sp. n. with an Aphaenogaster ant from Chongqing; B. qiului sp. n. with a Pheidole ant, B. xuhaoi sp. n. with a Lasius ant, and B. zhouchaoi sp. n. with Lasius and Nylanderia ants from Sichuan. Batrisodes zethus sp. n. was collected from a leaf litter sample.

Eight new species of the genus Batrisodes Reitter are described from continental China, seven of which were found in association with ants: B. abdominalis sp. n. and B. tianmuensis sp. n. with an Ectomomyrmex ant from Zhejiang; B. grossus sp. n. with an Odontomachus ant from Guangxi; B. simianshanus sp. n. with an Aphaenogaster ant from Chongqing; B. qiului sp. n. with a Pheidole ant, B. xuhaoi sp. n. with a Lasius ant, and B. zhouchaoi sp. n. with Lasius and Nylanderia ants from Sichuan. Batrisodes zethus sp. n. was collected from a leaf litter sample.

Batrisodes petalosus Jiang and Yin, new species (Figs 3���4) Type material. (2 ♂♂, 1 ♀). Holotype: CHINA: ♂, labeled ��� CHINA: Ningxia, Longde County (���������), Liupan Shan N. R. (������������������������), vi.2008, Bi Wen-Xuan leg.��� (SNUC). Paratypes: CHINA: 1 ♂, 1 ♀, same label data as the holotype (SNUC). Diagnosis of male. The new species is distinct among its Palaearctic congeners by possessing enlarged and flattened pedicel in the male, which is about four times as long as antennomere III. When combined with the frontal modifications (Fig. 4 A) and aedeagus (median lobe apically expanded; Fig. 4 F), it can be readily separated from all other European and East Asian species. Description. Male (Fig. 3 A). Body reddish brown, BL 2.86���2.90 mm. Head (Fig. 4 A) wider than long, roundly rectangular, HL 0.61���0.65 mm, HW 0.68���0.69 mm, with distinct, nude vertexal foveae, antennal tubercles prominent; anterior margin of frons strongly sinuate, with five bunches of thick setae anteromedially, area between antennal tubercles concave; clypeus medially impunctate, with round anterior margin; short lateral longitudinal carinae extending from level of vertexal foveae posteriorly to occipital constriction, lacking median vertexal carina. Each eye composed of about 50 facets. Antenna (Fig. 4 C) moniliform, with 11 antennomeres, club formed by three apical antennomeres; scape broad, slightly elongate, pedicel larger than scape, widest at apex. Pronotum about as long as wide, PL 0.57���0.60 mm, PW 0.61���0.62 mm, rounded laterally, disc slightly convex, with distinct median antebasal foveae, median and lateral longitudinal sulci present; outer and inner basolateral foveae small but distinct. Elytra slightly wider than long, EL 0.88���0.89 mm, EW 0.90���0.93 mm; each elytron with three large basal foveae; discal striae shallow and short. Mesofemur (Fig. 4 D) with tiny ventral spine near middle; mesotibiae (Fig. 4 E) with small ventral denticle at basal 3/5, and large triangular apical spine. Abdomen wider than long, AL 0.76��� 0.80 mm, AW 0.85���0.87 mm; tergite IV longest, about twice as long as next, with strongly oblique marginal carinae. Length of aedeagus (Fig. 4 F���G) 0.45 mm; median lobe simple, flattened, apically expanded, nearly symmetric. Female (Fig. 3 B). General habitus similar to male, frons (Fig. 4 B) unmodified; each eye composed of about 35 facets, and with one short ocular spine; pedicel not dilated as in male, only about twice length of antennomere III; legs lacking denticle and spine; tergite VIII (Fig. 4 H) semicircular; sternite VIII (Fig. 4 I) transverse; symmetric genital complex (Fig. 4 J) weakly sclerotized, width 0.27 mm. Measurements of body parts: BL 2.96 mm, HL 0.62 mm, HW 0.72 mm, PL 0.61 mm, PW 0.62 mm, EL 0.87 mm, EW 0.96 mm, AL 0.86 mm, AW 0.92 mm. Distribution. Northwestern China: Ningxia. Host ant. Myrmica sp. Etymology. The specific epithet is derived from the Greek adjective ���petalos���, meaning ���flat, spread out���, referring to the flattened and enlarged male pedicel of the new species., Published as part of Jiang, Ri-Xin & Yin, Zi-Wei, 2016, Two new species of Batrisodes Reitter (Coleoptera: Staphylinidae: Pselaphinae) from China, pp. 194-200 in Zootaxa 4205 (2) on pages 197-198, DOI: 10.11646/zootaxa.4205.2.9, http://zenodo.org/record/192959

Batrisodes linyejiei Jiang and Yin, new species (Figs 1��� 2) Type material ( 7 ♂♂, 5 ♀♀). Holotype: CHINA: ♂, labeled ��� China: Sichuan, Xiaojin County (���������), Jiajin Shan (���������), 30��48'49"N, 102��42'55"E, ant nest under rock, 2490 m, 20.vii.2015, Jiang, Peng, Tu & Zhou leg.��� (SNUC). Paratypes: CHINA: 4 ♂♂, 5 ♀♀, same label data as the holotype (SNUC); 2 ♂♂, labeled ��� China, Sichuan, Ya���an City (���������), Baoxing Hsien (���������), Shenmulei (���������������), 30��41'15"N, 102��42'13"E, 2300 m, 01.v.2016, Lin Ye-Jie & Chen Zhuo leg.��� (SNUC). Diagnosis of male. The new species can be separated from all other congeners by the following combination of characters: anterior margin of frons serrated, with a bunch of thick curved setae anteromedially; vertex with lateral carinae extending from above eyes to the occipital constriction; antennomere II subequal in length as antennomere I; antennomeres XI lacking a ventral spine; profemur with a tiny protuberance at ventral margin near the middle; mesotibia with a small ventral denticle at apical third, and a large triangular apical spine; aedeagus symmetric dorso-ventrally, flattened, simple. Description. Male. (Fig. 1 A), Body reddish brown, BL 2.94���2.95 mm. Head (Fig. 2 A) about as long as wide, rectangular, HL 0.55���0.58 mm, HW 0.59���0.60 mm, with large vertexal foveae, antennal tubercles prominent; frons with sinuate anterior margin and thick hook-like median setae, area between moderately raised antennal tubercles concave; clypeus medially impunctate, with round anterior margin; lateral longitudinal carinae extending from above eyes to occipital constriction, lacking median vertexal carina. Each eye composed of about 50 facets. Antenna (Fig. 2 C) with 11 antennomeres; club loosely formed by three apical antennomeres, scapes and pedicel enlarged and elongate, antennomeres III���XI moniliform. Pronotum about as long as wide, PL 0.61 mm, PW 0.58��� 0.60 mm, disc slightly convex; with distinct median antebasal fovea; median and lateral longitudinal sulci present. Elytra much wider than long, EL 0.94���0.95 mm, EW 1.02���1.04 mm; each elytron with three small but distinct basal foveae, inner two close; discal striae shallow and short. Mesofemora (Fig. 2 D) with thin but distinct ventral spine near middle; mesotibiae (Fig. 5E) with small ventral denticle at apical 2/5, and large triangular apical spine. Abdomen slightly wider than long, AL 0.81���0.84 mm, AW 0.83���0.85 mm; tergite IV (first visible tergite) longest, about 2.5 times as long as next, with strongly oblique marginal carinae. Length of aedeagus (Fig. 2 F���G) 0.37 mm; median lobe simple, flattened, nearly symmetric. Female (Fig. 1 B). General habitus similar to male, frons (Fig. 2 B) unmodified; each eye composed of about 35 facets; pedicel relatively shorter in relation to scape; legs lacking denticle and spine; tergite VIII (Fig. 2 H) semicircular; sternite VIII (Fig. 2 I) transverse; symmetric genital complex (Fig. 2 J) weakly sclerotized. Measurements of body parts: BL 3.02���3.06 mm, HL 0.58 mm, HW 0.60���0.62 mm, PL 0.62 mm, PW 0.60 mm, EL 0.96���0.98 mm, EW 1.02���1.06 mm, AL 0.86���0.88 mm, AW 0.82���0.83 mm. Distribution. Southwestern China: Sichuan. Host ant. Formica sp., Published as part of Jiang, Ri-Xin & Yin, Zi-Wei, 2016, Two new species of Batrisodes Reitter (Coleoptera: Staphylinidae: Pselaphinae) from China, pp. 194-200 in Zootaxa 4205 (2) on pages 194-197, DOI: 10.11646/zootaxa.4205.2.9, http://zenodo.org/record/192959