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This upload contains the main dataset represented in the figures of the following publication: "Acoustic non-Hermitian skin effect from twisted winding topology" by Li Zhang et al.

Mycterothrips cangshanensis sp. n. (Figs 10–17) Female macroptera. Body brown (Fig. 10); antennae brown with the base and apex of segment III pale (Fig. 15); all legs brown with tarsus yellowish brown; fore wings brown except base pale, clavus brown with apex yellow (Fig. 16); body setae strong and dark brown. Head wider than long, slightly rounded at cheeks; ocellar setae I present, setae III situated between posterior ocelli, postocular setae I the longest (Fig. 12). Antennae 8-segmented, segment I with two dorsal apical median setae, II with microtrichial rows and a mid-dorsal seta below CPS on dorsal surface, segments III and IV with sense cone forked and tapering in distal neck-like third (Fig. 15). Pronotum wider than long, dorsal surface smooth; two pairs of posteromarginal setae, setae I slightly longer; two pairs of posteroangular setae, setae I subequal to setae II. Mesonotum with CPS anteromedially; median pair of setae near posterior margin. Metanotum with reticulate striations, but with irregular longitudinal striations laterally, median pair of setae situated at anterior margin, CPS absent (Fig. 12). Fore wing first vein with seven setae at base and two near apex, second vein with setal row complete; posterior fringe cilia slightly wavy; clavus with 5 marginal setae and one discal seta (Fig. 16). Abdominal tergites without ciliate microtrichia on lines of sculpture, smooth medially, but posterior margin of laterotergites often partly irregularly dentate; tergite II with 3 lateral marginal setae, tergites VI–VIII with S4 setae minute; tergite VIII with complete posteromarginal comb (Fig. 13); tergite IX with both anterior and posterior pairs of CPS; tergite X with longitudinally split in distal 1/3; sternites without discal setae; median pair of setae on sternite VII situated in front of posterior margin. Measurements (holotype female in microns). Body length 1477. Head, length 98; width across eyes 161. Ocellar setae III length 68. Pronotum, length 137; maximum width 196; posteroangular setae length, inner 66, outer 66. Metanotum median setae length 52. Fore wing length 960. Antennal segments III–VIII length 62, 54, 32, 37, 10, 16. Male macroptera. Body paler and smaller than female (Fig. 11); antennae 8-segmented, segment VI with microtrichia on dorsal and ventral surfaces, almost as long as that female; abdominal tergite VIII with complete posteromarginal comb; tergite IX with one pair of CPS, SB1 setae vestigial (Fig. 14); sternite without discal seta; hypomere expanded at apex (Fig. 17). Measurements (paratype male in microns). Body length 1115. Head, length 95; width across eyes 137. Ocellar setae III length 45. Pronotum, length 102; maximum width 161; posteroangular setae length, inner 55, outer 55. Metanotum median setae length 34. Fore wing length 716. Antennal segments III–VIII length 55, 52, 30, 38, 8, 16. Specimens examined. Holotype female: CHINA, Yunnan Province, Dali City, Cangshan Mountain, from Parthenocissus quinquefolia, 15.vii.2018 (Hui Liu), in collection of Yunnan Agricultural University, Kunming. Paratypes: 6 females, 2 males, collected with holotype. Etymology. The new species is named after collected place. Comments. This new species is similar to to M. weii from Iran, with ocellar setae I present, mesonotum with CPS anteromedially, abdominal tergites without ciliate microtrichia on lines of sculpture and tergite II with 3 lateral marginal setae (Mirab-Balou et al. 2011), but this new species has the body brown, ocellar setae III situated between the posterior ocelli, posterior margin of abdominal laterotergites often partly irregularly dentate and SB1 setae degenerate on tergite IX of male.

Mycterothrips hani sp.n. (Figs 36–42) Female macroptera. Body yellow to brown (Fig. 36); head and thorax yellow with brown lateral margins, mesonotum shaded at anterior margin; abdominal segments I & IX–X yellow, II–VIII brown with posterior margin of 1/3 yellow (Fig. 38); antennal segments I–II brown, the base and apex of segments III–IV yellow, and extreme base of V slightly paler, VI brown, VII–VIII light brown (Fig. 42); all legs yellowish brown, fore wings pale, sometimes with small brown patch on the sub-base and apex of clavus (Fig. 41); prominent body setae yellow. Head wider than long, slightly rounded at cheeks; ocellar setae I present, setae III situated near anterior margin of posterior ocelli, postocular setae I behind posterior ocelli (Fig. 38). Antennae 8-segmented, segment I with two dorsal apical median setae, II with 3 microtrichial rows on dorsal surface and a mid-dorsal seta below CPS on dorsal surface, III and IV widest at middle and tapering in distal neck-like third, III–VI with some rows of microtrichia on dorsal and ventral surfaces, VIII slightly longer than VII (Fig. 42). Pronotum wider than long, smooth medially, weakly sculptured with transverse striations on anterior and posterior part, with about 30 discal setae; two pairs of posteromarginal setae, setae I longer than setae II; inner posteroangular setae subequal to outer one (Fig. 38). Mesonotum with CPS anteromedially; median pair of setae situated near or on posterior margin. Metanotum with weak reticulations, but with longitudinal striations laterally, median pair of setae situated at anterior margin, CPS absent (Fig. 38). Fore wing first vein with seven setae at base and two near apex, second vein with setal row complete; posterior fringe cilia slightly wavy; clavus with 5 marginal setae and one discal seta (Fig. 41). Abdominal tergites and laterotergites without ciliate microtrichia; tergite II with 3 lateral marginal setae; tergites VI–VIII with S4 setae minute; tergite IX with both anterior and posterior pairs of CPS; tergite X split longitudinally in distal half (Fig. 39); sternites without discal setae; sternite VII with median pair of setae situated in front of posterior margin. Measurements (holotype female in microns). Body length 1379. Head, length 125; width across eyes 156. Ocellar setae III length 53. Pronotum, length 123; maximum width 193; posteroangular setae length, inner 60, outer 58. Metanotum median setae length 37. Fore wing length 754. Antennal segments III–VIII length 48, 55, 38, 48, 12, 18. Male macroptera. Body paler and smaller than female (Fig. 37); antennae 8-segmented, segment VI almost as long as that of female, with microtrichia on dorsal and ventral surfaces; abdominal tergite VIII with complete posteromarginal comb; IX with one pair of CPS and SB1vestigial (Fig. 40); abdominal sternites without discal setae; hypomere not expanded at apex. Measurements (paratype male in microns). Body length 1128. Head, length 92; width across eyes 133. Ocellar setae III length 42. Pronotum, length 97; maximum width 152; posteroangular setae length, inner 48, outer 42. Metanotum median setae length 35. Fore wing length 634. Antennal segments III–VIII length 47, 49, 34, 46, 10, 16. Specimens examined. Holotype female: CHINA, Chongqing City, Xiushan County, Fenghuangshan Forest Park, from Hemiboea subcapitata, 7.viii.2017 (Yajin Li), in collection of Yunnan Agricultural University, Kunming. Paratypes: 2 females, 2 males, collected with holotype; 5 females, 2 males from Oplismenus compositus, 7.viii.2017, same locality and collector. Etymology. In honor of Mr. Yunfa Han for his excellent work on thrips taxonomy in China. Comments. This species is similar to M. weii, but has antennal segments I–II brown, and the male with SB1 on tergite IX vestigial. In addition, the yellow head with brown lateral margins is unusual among Mycterothrips members., Published as part of Li, Yajin & Zhang, Hongrui, 2022, Five new species and two new records of Mycterothrips (Thysanoptera: Thripidae) from Southern China, pp. 544-566 in Zootaxa 5214 (4) on pages 544-566, DOI: 10.11646/zootaxa.5214.4.4, http://zenodo.org/record/7397755

Mycterothrips photiniae sp.n. (Figs 6–9) Female macroptera. Body brown (Fig. 8), all legs brown except tibia apex and tarsus yellowish brown; antennae brown with apex of segment III yellow (Fig. 6); fore wings brown except base pale, clavus brown with apex yellow. Head wider than long, slightly rounded at cheeks; ocellar setae I absent, setae III situated near anterior margin of posterior ocelli; five or six pairs of postocular setae (Fig. 7). Antennae 8-segmented, segment I with two dorsal apical median setae, II with 3 microtrichial rows on dorsal surface, III–VI with some rows of microtrichia on dorsal and ventral surfaces, VIII slightly longer than VII (Fig. 6). Pronotum wider than long, weakly sculptured with transverse striations, but smooth medially; one or two pairs of posteromarginal setae; two pairs of posteroangular setae, inner setae shorter than outer one. Mesonotum with CPS anteromedially; median pair of setae near posterior margin. Metanotum reticulate striate, but with irregular longitudinal striations laterally, CPS absent (Fig. 7). Fore wing first vein with seven setae at base and two near apex, second vein with setal row complete; posterior fringe cilia slightly wavy; clavus with 5 marginal setae and one discal seta. Abdominal tergites II–VIII without ciliate microtrichia on lines of sculpture, but often with a few large irregular microtrichia on sculpture lines at least on segment VIII; tergite II with 3 lateral marginal setae, tergites VI–VIII with S4 setae minute; tergite IX with posterior pair of CPS; tergite X split longitudinally in distal 1/4 (Fig. 9); sternites without discal setae; VII with median pair of setae situated in front of posterior margin. Measurements (holotype female in microns). Body length 1454. Head, length 100; width across eyes 154. Ocellar setae III length 42. Pronotum, length 125; maximum width 189; posteroangular setae length, inner 58, outer 68. Metanotum median setae length 54. Fore wing length 932. Antennal segments III–VIII length 59, 50, 38, 45, 10, 19. Male. Unknown. Specimens examined. Holotype female: CHINA, Yunnan Province, Shilin County, Long Lake, from Photinia serratifolia, 15.iii.2017 (Yajin Li), in collection of Yunnan Agricultural University, Kunming. Paratypes: 4 females, collected with holotype; 4 females, from Pinus sp., collected on the same locality. Etymology. In reference to the host plant of this species. Comments. This species runs to grandis in the key to world species of Mycterothrips, but this new species has the body brown, mesonotum with anteromedian CPS and abdominal tergite II with 3 lateral marginal setae., Published as part of Li, Yajin & Zhang, Hongrui, 2022, Five new species and two new records of Mycterothrips (Thysanoptera: Thripidae) from Southern China, pp. 544-566 in Zootaxa 5214 (4) on pages 544-566, DOI: 10.11646/zootaxa.5214.4.4, http://zenodo.org/record/7397755

Mycterothrips daweishanensis sp. n. (Figs 18–24) Female macroptera. Body brown (Fig. 18); antennae wholly brown; all legs brown except tibia apex and tarsus yellowish brown (Fig. 20); fore wings brown except base pale; clavus brown with apex pale (Fig. 24). Head wider than long, slightly rounded at cheeks; ocellar setae I present, setae III situated near anterior margin of posterior ocelli; 5–6 pairs of postocular setae (Fig. 19). Antennae 8-segmented, segment I with two dorsal apical median setae, II with 3 microtrichial rows on dorsal surface; III and IV widest at middle and tapering in distal necklike third; III–VI with some rows of microtrichia on dorsal and ventral surfaces; VIII slightly longer than VII (Fig. 20). Pronotum wider than long, weakly sculptured with transverse striations, inner posteroangular setae shorter than outer pair; two pairs of posteromarginal setae. Mesonotum with CPS absent; median pair of setae near posterior margin, Metanotum reticulate striate, but with irregular longitudinal striations laterally, CPS absent (Fig. 19). Fore wing first vein with seven setae at base and two near apex, second vein with setal row complete; posterior fringe cilia slightly wavy; clavus with 5 marginal setae and one discal seta (Fig. 24). Abdominal tergites and laterotergites without ciliate microtrichia on lines of sculpture; tergite II with 4 lateral marginal setae; tergites VI–VIII with S4 setae minute; tergite IX with only posterior pair of CPS; tergite X split longitudinally in distal 1/3 (Fig. 22); abdominal sternite IV–V with 0–1 discal setae, VI with 1–2 discal setae; sternite VII with median pair of setae situated in front of posterior margin (Fig. 21). Measurements (holotype female in microns). Body length 1557. Head, length 101; width across eyes 168. Ocellar setae III length 70. Pronotum, length 140; maximum width 194; posteroangular setae length, inner 85, outer 89. Metanotum median setae length 62. Fore wing length 895. Antennal segments III–VIII length 54, 57, 36, 44, 10, 20. Male Unknown. Specimens examined. Holotype female: CHINA, Yunnan Province, Pingbian County, Dawei Mountain, from Gardneria multiflora, 3.viii.2017 (Bo Kong), in collection of Yunnan Agricultural University, Kunming. Paratypes: 6 females, collected with holotype. Etymology. The new species is named after collected place. Comments. This species runs to M. auratus in the key to world species of Mycterothrips by Masumoto and Okajima (2006), and these two share the following character states: Ocellar setae III near anterior margin of hind ocelli; mesonotum without campaniform sensilla anteromedially; abdominal tergum II with four lateral marginal setae; sternites III–VII with discal setae. However, this new species differs in having antennae wholly brown, pronotum with two long discal setae near anterior margin, abdominal tergites without ciliate microtrichia along lines of sculpture, tergite IX with only posterior pair of CPS, sternites IV–VI with 0–2 discal setae. This species is also similar to M. nastarani, but that has the body uniformly yellowish white, mesonotum with CPS, and sternites III–VII with more than 3 discal setae.

Mycterothrips bicolor sp. n. (Figs 1–5) Female macroptera. Body bicolored (Fig. 1); head, thorax and abdominal segment I yellow, abdominal segment II brown, segments III–X dark brown; antennae brown, with apex of segment III pale (Fig. 4); all legs yellow; fore wings light brown with base yellow, clavus yellow (Fig. 5); prominent body setae brown. Head wider than long, slightly rounded at cheeks; ocellar setae I present, setae III situated near anterior margin of posterior ocelli; postocular setae I the longest (Fig. 2). Antennae 8-segmented, segment I with two dorsal apical median setae, II with 3 microtrichial rows on dorsal surface and a mid-dorsal seta below CPS on dorsal surface, III and IV widest at middle and tapering in distal neck-like third, III–VI with some rows of microtrichia on dorsal and ventral surfaces, VIII slightly longer than VII (Fig. 4). Pronotum wider than long, dorsal surface smooth, with a few transverse striations on posterior ¼; one pair of anteromarginal setae longer than discal setae; two pairs of posteromarginal setae, setae I longer than II, inner posteroangular setae subequal to outer one (Fig. 2). Mesonotum with CPS present; median pair of setae situated near or on posterior margin. Metanotum with very weak reticulations, but with irregular longitudinal striations laterally, median pair of setae situated near anterior margin, CPS absent. Fore wing first vein with seven setae at base and two near apex, second vein with setal row complete; posterior fringe cilia slightly wavy; clavus with 5 marginal setae and one discal seta (Fig. 5). Abdominal tergites and laterotergites without ciliate microtrichia on lines of sculpture, but often with a few large irregular microtrichia on sculpture lines at least on segment VIII, posterior margin of tergites V–VII without microtrichia laterally; tergite II with 4 lateral marginal setae; tergites VI–VIII with S4 setae minute; tergite IX with both anterior and posterior pairs of CPS; tergite X split longitudinally in distal 1/3 (Fig. 3); sternites without discal seta; sternite VII with median pair of setae situated in front of posterior margin. Measurements (holotype female in microns). Body length 1390. Head, length 90; width across eyes 142. Ocellar setae III length 68. Pronotum, length 135; maximum width 162; posteroangular setae length, inner 70, outer 70. Metanotum median setae length 54. Fore wing length 778. Antennal segments III–VIII length 60, 53, 36, 45, 8, 17. Male. Unknown. Specimens examined. Holotype female: CHINA, Yunnan Province, Xuanwei City, Xize County, from Capsicum annuum, 3.x.2016 (Bo Kong), in collection of Yunnan Agricultural University, Kunming. Paratypes: 4 females collected with holotype, same locality and collector. Etymology. In reference to body bicolor. Comments. This species is very closely related to caudibrunneus in Taiwan, with the same coloration of abdomen and fore wing, ocellar setae I present, abdominal tergite II with four lateral marginal setae. However, it is distinguished by posterior margin of tergites V–VII without microtrichia laterally, metascutum with median pair of setae near anterior margin (Wang 1999, Masumoto and Okajima 2006). This species is also similar to consociatus, but differs in bicoloured body colour and fore wing with base pale., Published as part of Li, Yajin & Zhang, Hongrui, 2022, Five new species and two new records of Mycterothrips (Thysanoptera: Thripidae) from Southern China, pp. 544-566 in Zootaxa 5214 (4) on pages 544-566, DOI: 10.11646/zootaxa.5214.4.4, http://zenodo.org/record/7397755, {"references":["Masumoto, M. & Okajima, S. (2006) A revision of and key to the world species of Mycterothrips Trybom (Thysanoptera, Thripidae). Zootaxa, 1261 (1), 1 - 90. https: // doi. org / 10.11646 / zootaxa. 1261.1.1"]}

Helionothrips phragmitesi sp.n. (Figs 5, 10, 22, 38–42) Female macroptera. Body color dark brown, head entirely dark; fore femora brown, tibiae yellowish brown; mid and hind legs dark brown, with small extreme apex of tibiae yellow (Fig. 38); all tarsi yellow; antennal segments I–II dark brown, III–V and basal half of VI yellow, apical half of VI and VII–VIII light brown (Fig. 10); fore wing (Fig. 5) brown, base and apex darkest, with a sub-basal white band, weakly shaded between anterior and posterior margin at distal half; clavus dark brown. Head about 1.4–1.5 times as wide as long, completely sculptured; ocellar hump reticulate obvious with internal wrinkles (Fig. 39); ocelli large; anterior and posterior carina of occipital collar almost parallel, the posteromedian reticles of collar with many thicken dots inside. Antennae 8-segmented, general stout; forked sense cones on III extending to basal third of IV, on IV not surpassing midline of V (Fig. 10); simple dorsal sense cone on VI scarcely beyond apex of VIII, microtrichia present on ventral surface of segments IV–VI. Mouth cone rounded apically; maxillary palps 2–segmented. Pronotum entirely covered by polygonal reticles of a rather uniform size, discal setae moderately long; most median and posterior reticles with wrinkles inside (Fig. 22); mesoscutum with weak wrinkles in median reticles; metascutal triangle lacking wrinkles (Fig. 40), median setae outer and anterior to the CPS; metascutellum about 2.5 times as wide as long. Fore wing (Fig. 5) first vein with 6–7 basal and 2 distal setae, second vein with about 7 setae, posteromarginal cilia distinctive wavy; clavus with 4 veinal setae. Abdominal tergites I and II completely reticulate (Fig. 41); tergites III–VIII with heavy antecostal line divided into arches connected by a fine line, entirely covered with polygonal reticles except for posterior median unsculptured areas; II–VII lateral fourth with wrinkles in reticles (Fig. 41); VIII with comb of microtrichia narrowly interrupted medially by lacking about 4–5 teeth (Fig. 42); three pairs of needle-like setae on apex of tergite IX, S2 the longest, S1 and S3 almost subequal in length; Abdominal segment X short and small, less than half the length of IX; X with median split complete. Sternites entirely polygonally reticulate posterior of antecostal line, with three pairs of long posterior margin setae. Measurements (holotype female in microns): Body length 1562. Head, length 129; width across genae 183. Eye, length 73; width 48. Pronotum, length 154; width 223. Fore wing length 900. Abdominal segments VIII–X median length 101, 118, 55. Antennal segments I–VIII length (width): 23(21), 38(30), 53(22), 44(22), 40(20), 29(17), 9(8), 27(4). Male unknown. Specimens studied. Holotype female, CHINA, Chongqing City, Xiushan County, from leaves of Phragmites communis, 22.v.2017 (Li Yajin & Liu Hui), in collection of Yunnan Agricultural University, Kunming. Paratypes: 19 females collected from same locality and plant as holotype, with 2 females deposited in Australian National Insect Collection, Canberra. Etymology. In reference to the host plant of this species. Comments. This new species is distinguished from other Helionothrips species by the reticles on the posterior half of the pronotum having internal wrinkles, the forked antennal sense cone on IV not surpassing mid-point of V, antennal segments I & II dark brown while VI light brown with basal half yellow, also mid and hind tibiae with a small yellow area at apex. This new species is most similar to communis, ponkikiri and unitatis in having the posterior half of the pronotum with reticles bearing internal wrinkles. However, in communis, ponkikiri and unitatis, the forked sense cone on antennal segment IV at least extends to the apex of V, and segment VI as brown as I & II (ponkikiri and unitatis) or darker than I & II (communis). Moreover, in this new species, wrinkles are present in the mesoscutum reticles but not in the metascutum reticles. In contrast, both the meso- and metascutal reticles have internal wrinkles in communis and unitatis versus both lacking in ponkikiri., Published as part of Xie, Yanlan, Li, Yajin & Zhang, Hongrui, 2022, The genus Helionothrips (Thysanoptera, Panchaetothripinae) in China, with two new species and an identification key, pp. 392-402 in Zootaxa 5194 (3) on pages 399-400, DOI: 10.11646/zootaxa.5194.3.3, http://zenodo.org/record/7154424

Pseudopoda uncata Yang & Zhang sp. nov. (Figs 4–6) Type material. Holotype: ♂, CHINA : Yunnan Province: Mengjiao Township, Cangyuan County (23°16.38′N, 99°11.21′E, 1769 m), 11 May 2021, Z.Z. Yang & Z.M. Li (MHBU). Paratypes: 1♂, 5♀, with same data as for holotype (1♀ in MHBU, 1♂ 4♀ in DUIER); 2♂, 1♀, with same data as for holotype except: 23 May 2017, Z.Z. Yang (DUIER). Etymology. The specific name is a Latin adjective—derived from uncus (=hook), referring to the shape of embolus in ventral view. Diagnosis. According to the morphology of genital organs (for details see Jäger 2001), P. uncata sp. nov. belongs to the diversipunctata species-group, which, up to now, was comprised of six species: P. confusa, P. diversipunctata, P. intermedia, P. lutea, P. marsupia and P. serrata. Both sexes of this new species are most similar to P. confusa Jäger, Pathoumthong and Vedel, 2006 (see Jäger et al. 2006: 220, figs 1–13, 29–32) in: embolus hook-shaped in ventral view, with similar arising point and length; U-shaped median septum; spherical structures of internal duct system. It can be distinguished from the latter species by the following combination of characters: 1. Embolus tip inversely U-shaped in ventral view (embolus bent at right angles in P. confusa); 2. Finger-like dRTA longer than in P. confusa; 3. Median septum narrower than in P. confusa; 4. First windings of internal duct system close to each other (far away from each other in P. confusa). Females of this new species also resemble those of P. lutea (Thorell, 1895) (see Jäger 2002: 57, figs 184–185) by: anterior and median margins of lateral lobes goblet-shaped, internal duct system with distinct lateral loops visible through cuticle in ventral view as oval patches on anterior part of epigyne. It can be distinguished from the latter species by the following combination of characters: 1. Anterior margin trilobate (Figs 5D, 6D; bilobate in P. lutea); 2. Loops of internal duct system bending laterally (Figs 5E, 6E; loops bending medially in P. lutea). Description. Male: Total length 9.39–9.46 (n=4). Holotype total length 9.46: prosoma 4.63 long, 4.36 wide; opisthosoma 4.97 long, 3.10 wide. Two eye rows slightly recurved. Eyes diameters and interdistances: AME 0.23, ALE 0.33, PME 0.24, PLE 0.30, AME–AME 0.13, AME–ALE 0.08, PME–PME 0.29, PME–PLE 0.35. MOA 0.73 long, anterior width 0.59, posterior width 0.76. Clypeus height 0.31. Chelicerae furnished with 3 promarginal teeth and 4 retromarginal teeth, and some denticles between them. Leg measurements: I 23.03 (6.10, 1.97, 5.90, 6.62, 2.44), II 25.38 (7.57, 2.60, 5.95, 6.92, 2.34), III 18.77 (6.60, 1.74, 3.71, 5.03, 1.69), IV 15.44 (6.60, 1.22, 3.05, 2.08, 2.09). Leg formula: 2134. Leg spination: palp 131, 101, 2121; femur I 323, II–III 313, IV 323; patella I–III 101, IV 100; tibia I 4314, II 3124, III 3213, IV 2323; metatarsus I–III 2222, IV 3333. Palp as in diagnosis (Figs 5A–C, 6A–C). Cymbium slender, with distinct retrolateral bulge; subdistal tibia with a retrolaterad hump; RTA arising proximally to medially from tibia, vRTA obviously broader than dRTA; tegulum oval, upper part of tegulum with upfold; retrolateral part of spermophore almost straight, and running submarginally in tegulum; embolus arising from tegulum at 10 to 11 o’clock position; terminal embolus with groove (visible in retrolateral view); subapical part of embolus with two small teeth; conductor arising from tegulum at 1 o’clock position, slender, bent medially and then directed prolaterally. Coloration in ethanol (Figs 4A, B): Carapace yellowish, with some dark spots, two lateral bands inconspicuous, margin with gray black patterns. Cervical groove and radial furrow obvious. Fovea dark brown. Each eye surrounded with circular black patch. Chelicerae, labium, gnathocoxae and sternum yellowish. Legs yellowish, with small spots and slightly larger spine patches. Dorsal abdomen with lots of black dots; venter yellowish, with black patches. Female: Total length 8.97–9.91 (n=6). One paratype: total length 8.97; prosoma 4.12 long, 4.09 wide; opisthosoma 4.86 long, 3.59 wide. Eyes diameters and interdistances: AME 0.21, ALE 0.27, PME 0.22, PLE 0.27; AME–AME 0.18, AME–ALE 0.15, PME–PME 0.28, PME–PLE 0.41. MOA 0.67 long, anterior width 0.56, posterior width 0.70. Clypeus height 0.09. Leg measurements: I 17.64 (4.97, 2.13, 4.72, 4.18, 1.64), II 19.10 (5.92, 2.22, 4.94, 4.37, 1.65), III 13.81 (4.09, 1.84, 3.80, 2.55, 1.53), IV 15.51 (4.76, 1.59, 3.36, 3.64, 2.16). Leg formula: 2143. Leg spination: palp 131, 101, 2121, 2112; femur I–IV 323; patella I–IV 101; tibia I 3423, II 3323, III 3213, IV 3323; metatarsus I–III 2222, IV 3433. Epigyne as in diagnosis (Figs 5 D-E, 6D-E). Epigynal field wider than long, with trilobate anterior margin, anterior bands absent; lateral lobes longer than wide, anterior margin of lateral lobes converged on the central axis, anterior and median margins of lateral lobes goblet-shaped; posterior incision of lateral lobes indistinct, near the meeting point of lateral lobes; laterad loops of internal duct system appearing as oval dark patches in ventral view. Coloration in ethanol: Color and markings of body as in male, but generally lighter (Figs 4C, D). Distribution. China (Yunnan Province)., Published as part of Yang, Zi-Zhong, Wu, Ya-Ying, Li, Zhi-Min & Zhang, Bao-Shi, 2022, Two new species of the genus Pseudopoda Jäger, 2000 and first description of the male of Pseudopoda physematosa (Araneae, Sparassidae) from Yunnan Province China, pp. 347-360 in Zootaxa 5188 (4) on pages 352-354, DOI: 10.11646/zootaxa.5188.4.3, http://zenodo.org/record/7095600, {"references":["Jager, P. (2001) Diversitat der Riesenkrabbenspinnen im Himalaya. Uber eine Radiation zweier Gattungen in den Schneetropen (Araneae: Sparassidae: Heteropodinae). Courier Forschungsinstitut Senckenberg, 232, 1 - 136.","Jager, P., Pathoumthong, B. & Vedel, V. (2006) First record of the genus Pseudopoda Jager 2000 in Laos with description of new species (Arachnida, Araneae, Sparassidae). Senckenbergiana Biologica, 86, 219 - 228.","Thorell, T. (1895) Descriptive catalogue of the spiders of Burma, based upon the collection made by Eugene W. Oates and preserved in the British Museum, Taylor and Francis, London, 406 pp. https: // doi. org / 10.5962 / bhl. title. 17492","Jager, P. (2002) Heteropodinae: transfers and synonymies (Arachnida: Araneae: Sparassidae). Acta Arachnologica, 51, 33 - 61. https: // doi. org / 10.2476 / asjaa. 51.33"]}

Pseudopoda mingshengi Yang & Zhang sp. nov. (Figs 1–3) Type material. Holotype: ♂, CHINA: Yunnan Province: Mengjiao Township (23°16.38′N, 99°11.21′E, 1769 m), Cangyuan County, 23 May 2017, Z.Z. Yang & C.G. Li (MHBU) Paratypes: 4♀, with same data as for holotype (1♀ in MHBU, 3♀ in DUIER). Etymology. The specific name is dedicated to the late Prof. Dr Ming-Sheng Zhu for his efforts on arachnological studies in China; nomen in genitive case. Diagnosis. Males of this new species resemble those of P. contraria Jäger and Vedel, 2007 (see Jäger & Vedel 2007: 31, figs 114–116) in the embolus with a huge prolateral projection, and a broad apical tip of the embolic projection. It can be distinguished from the latter by the following combination of characters: 1. Apical part of embolus very broad (Figs 2B–C, 3B–C; thinner in P. contraria); 2. Embolic projection trapezoid-shaped in ventral view (Figs 2B, 3B; square-shaped in P. contraria); 3. Top of embolic projection with teeth (Figs 2B, 3B; absent in P. contraria). Females of this new species resemble those of P. pingu Jäger, 2015 (see Jäger 2015: 336, figs 21–23) by the lateral windings projecting anteriorly above the median septum and the longitudinally sclerotised internal duct system. It can be distinguished from the latter by the following combination of characters: 1. lateral lobes touching each other (Figs 2D, 3D; distinctly separated in P. pingu); 2. the first winding of internal duct system obviously wide (Figs 2E, 3E; moderately wide in P. pingu). Description. Male (holotype): Total length 10.83: prosoma 5.61 long, 5.24 wide; opisthosoma 5.24 long, 3.42 wide. AER slightly recurved, PER straight. Eyes diameters and interdistances: AME 0.21, ALE 0.45, PME 0.37, PLE 0.42, AME–AME 0.27, AME–ALE 0.08, PME–PME 0.29, PME–PLE 0.46. MOA 1.03 long, anterior width 0.64, posterior width 1.03. Clypeus height 0.26. Chelicerae brown, with 3 promarginal teeth and 4 retromarginal teeth, and some denticles between them. Sternum with dark setae. Legs yellowish brown, with small spots and slightly larger spine patches, metatarsus of leg with dense scopula. Leg measurements: I 23.47 (6.75, 2.80, 5.34, 6.24, 2.34), II 26.04 (7.61, 2.79, 6.62, 6.57, 2.45), III 20.12 (6.49, 2.40, 5.22, 4.36, 1.65), IV 21.18 (6.43, 2.15, 6.01, 4.52, 2.07). Leg formula: 2143. Leg spination: palp 131, 101, 2121; femur I–III 323, IV 321; patella I–IV 101; tibia I–IV 3223; metatarsus I–II 2112, III–IV 2222. Palp as in diagnosis (Figs 2A–C, 3A–C). Cymbium slender; RTA arising proximally to medially from tibia, dRTA long and distally sharp, vRTA indistinct; spermophor running submarginally at retromargin of tegulum, turning into an S-shaped duct at promargin of tegulum to the base of embolus; embolus arising from tegulum at 9 o’clock position, very broadened in its middle part; embolic projection flat and very large, pointing prolaterally, embolic projection distally with two triangular teeth; conductor arising from tegulum at 12 o’clock position, slender, bent ventrad. Coloration in ethanol: Carapace yellowish brown, bearing some small spots. Cervical groove and radial furrow obvious. Longitudinal fovea dark brown. Ocular area darker. Each eye surrounded with circular black patch. Labium, gnathocoxae and sternum yellowish brown, the posterior part of labium dark brown. Legs yellowish brown, with small spots and slightly larger spine patches, metatarsus darker. Dorsal opisthosoma gray black dorsally, with white lateral patterns and a big dark triangular median pattern, the posterior part whitish; venter gray black (Figs 1A–B). Female: Total length 13.65–13.71 (n=4). One paratype: Total length 13.71; prosoma 6.25 long, 5.78 wide; opisthosoma 7.47 long, 5.00 wide. Eyes diameters and interdistances: AME 0.30, ALE 0.44, PME 0.33, PLE 0.42, AME–AME 0.27, AME–ALE 0.11, PME–PME 0.38, PME–PLE 0.54. MOA 1.11 long, anterior width 0.77, posterior width 1.07. Clypeus height 0.50. Leg measurements: I 21.81 (6.27, 2.93, 5.98, 5.11, 1.52), II 23.63 (6.56, 3.43, 6.13, 5.51, 2.00), III 18.74 (5.47, 2.68, 4.57, 4.06, 1.96), IV 21.21 (6.48, 2.52, 5.22, 4.41, 2.58). Leg formula: 2143. Leg spination: palp 131, 101, 2121, 2112; femur I–II 323, III 332, IV 331; patella I–IV 001; tibia I–IV 3223; metatarsus I–II 2112, III–IV 2222. Epigyne as in diagnosis (Figs 2D–E, 3D–E). Epigynal field wider than long, with slightly trilobate anterior margin, anterior bands short, indistinct; lateral lobes longer than wide, slightly converged on the central axis; anterior margins of lateral lobes heart - shaped; loops of internal duct system bending laterally, the posterior parts hidden behind lateral lobes; the anterior margin of longitudinally sclerotised parts beyond copulatory opening. Coloration in ethanol: Color and markings of carapace as in male. Dorsal opisthosoma dark brown dorsally, with some irregular patches laterally and a distinct white transversal patch in the posterior part; venter brown, with small and irregular patches (Figs 1C–D). Distribution. China (Yunnan). Note. The species P. mingshengi cannot be allocated to any of the nine known species groups according to morphological features. Males share the broad prolateral embolic projection with several species, e.g., P. daxing Zhao and Li, 2018 in Jiang et al. 2018, P. digitata Jäger and Vedel, 2007 and P. contraria, but females of this new species are completely different.

dentimarginus Li & Zhang, 2005 Distribution: China (Henan)., Published as part of Cao, Wenjun & Xing, Jichun, 2022, Japananoides, a new leafhopper genus from China (Hemiptera: Cicadellidae: Deltocephalinae), pp. 345-354 in Zootaxa 5154 (3) on page 346, DOI: 10.11646/zootaxa.5154.3.7, http://zenodo.org/record/6644812, {"references":["Li, Z. Z. & Zhang, B (2005) A taxonomic study of the genus Hishimonoides Ishihara from China (Hemiptera: Cicadellidae: Euscelinae). Acta Zootaxon Sinica, 30 (4), 794 - 797. [in Chinese with English summary]"]}

miaolingensis Li & Zhang, 2006 Distribution: China (Guizhou, Guangxi, Zhejiang, Hainan, Jiangxi, Guangdong)., Published as part of Cao, Wenjun & Xing, Jichun, 2022, Japananoides, a new leafhopper genus from China (Hemiptera: Cicadellidae: Deltocephalinae), pp. 345-354 in Zootaxa 5154 (3) on page 346, DOI: 10.11646/zootaxa.5154.3.7, http://zenodo.org/record/6644812, {"references":["Li, Z. Z. & Zhang, B. (2006) A new species of the genus Hishimonoides Ishihara (Hemiptera: Cicadellidae: Euscelinae) from Guizhou Province, China. Entomotaxonomia, 28, 262 - 264. [in Chinese with English summary] https: // doi. org / 10.1016 / S 1872 - 2067 (06) 60034 - X"]}

9. Conocephalus (Anisoptera) dorsalidentatus Li, Zhang & Shi, sp. nov. (Figs. 5, 7 A–B) Description. Male. Body medium for genus. Fastigium verticis narrow, apex blunt, width about 1/3 of scape, with a thin median longitudinal furrow; connected with or slightly apart from frontal verticis. Eyes ovoid, protruding forward and outward. Anterior margin of pronotum slightly arc-shaped, posterior margin rounded; lateral lobe nearly triangular, higher than long, anterior margin with 1 small nick, posterior margin rounded; the area corresponding to thoracic foramen semitransparent, slightly plump; humeral sinus indistinct. Prosternum with a pair of spines. Procoxae with 1 spine; profemora unarmed on ventral surface, apices of genicular lobes with 1–2 spines on inner margin and outer margin unarmed; protibiae with 6 spines on inner margin and 4–7 spines on outer margin of ventral surface; tibial tympana slit-like. Mesofemora unarmed on inner margin of ventral surface, outer margin of ventral surface with 4–5 spines; mesotibiae with 6–7 spines on inner margin and 6–8 spines on outer margin of ventral surface. Postfemora unarmed on inner margin and with 9–12 spines on outer margin of ventral surface, apices of inner and outer genicular lobes with 2 spines separately, dorsal spine long than ventral spine; posttibiae with 24–29 spines on inner margin and 16–23 spines on outer margin of dorsal surface, 1 pair of dorsal apical spurs and 2 pairs of ventral apical spurs. Tegmina long, surpassing apices of postfemora, apices rounded. Stridulatory area of left tegmen trapezoid, Cu2 vein robust, fusiform. Hind wings longer than tegmina. 10 th abdominal tergite broad, posterior margin with a shallow triangular notch, the lateral lobes slightly curved ventrad, apices rounded. Cerci long, slightly incurved, 3/4 basal areas cylindrical, robust, 1/4 apical areas constricted, apices rounded; with two teeth near the middle, the tooth near base short, basal half slightly broad, gradually narrowing, apical half slightly compressed, apex acute, inserted on dorsal surface, directing dorsoinward; the other tooth near apex long, spine-shaped, inserted on inner surface, apex thin and acute, directing ventrad. Subgenital plate roughly rectangular, the middle of posterior margin with a shallow triangular notch. Styli slightly short, cylindrical, apices rounded, inserted on apices of the lobes of subgenital plate. Coloration. Body green. Dorsal surface of head with a broad longitudinal brown stripe, the midline with a thin light stripe. Disc of pronotum with a broad longitudinal blackish brown stripe, outer margins with a light yellow stripe separately, and the midline with a thin light stripe. Dorsal surface of abdomen reddish brown, outer margins with a longitudinal light yellow stripe separately. Spines on ventral surface of mesofemora and postfemora, and spines on dorsal surface of posttibiae brown. Apical areas of two teeth on cercus brown. Costal area of tegmen light, Sc vein light yellow, the other areas light brown. Female. Appearance is similar to male. 10 th abdominal tergite short, the middle of posterior margin with a nearly triangular concavity; epiproct tongue-shaped. Cerci long, conical. Ovipositor straight, basal area slightly narrow, middle area broad, gradually narrowing; dorsal and ventral margins smooth, dorsal valvulae longer than ventral ones, apices of dorsal valvulae blunt while ventral ones acute. Subgenital plate roughly triangular, basal area broad, narrowing, posterior margin straight. Ovipositor yellowish brown. Material examined. Holotype: ♂, Meilinshan, Shenzhen, Guangdong, 15 July, 2018, coll. Tao Zhang. Paratypes: 4♂ 3♀, the other data same as holotype. Measurements (mm). Body: ♂ 13.0–19.0, ♀ 16.0–18.0; pronotum: ♂ 4.0–4.3, ♀ 4.0–4.3; tegmina: ♂ 14.0–15.5, ♀ 15.5–16.0; postfemora: ♂ 9.0–10.5, ♀ 10.0–11.0; ovipositor: 12.0–12.5. Distribution. China (Guangdong). Remarks. The new species is similar to Conocephalus (Conocephalus) bidentatus Shi & Zheng, 1994 and Conocephalus (Conocephalus) differentus Shi & Liang, 1997, but it differs in: tegmina elongate, surpassing apices of postfemora; anterior margin of lateral lobe of pronotum with 1 small incision; 1/4 apical area of male cercus constricted, the tooth near base short, inserted on dorsal surface, the tooth near apex elongate, spine-shaped, apex thin and acute, inserted on inner surface; basal area of ovipositor slightly narrow, middle area slightly broad. Etymology. The name of new species is derived from the tooth near base of male cercus inserted on dorsal surface. Latin dorsal- referring to dorsal, and Latin dent- referring to tooth.

Arctosa tangguoi Wang, Li & Zhang, sp. nov. (Figs 16A–E, 17A–D, 18A–B) Type material. Holotype ♂, Lingshui County, Diaoluo Mountain Nature Reserve, 18°41.445′N, 109°52.410′E, elev. 620 m, 16.IV.2009, G. Tang leg. (IZCAS). Etymology. The specific epithet is taken from the name of Mr. Guo Tang in honour of his great contribution to Chinese arachnology; noun (name) in genitive case. Diagnosis. The new species resembles A. hainan Wang, Li & Zhang, sp. nov. (Figs 10C–F, 11A–F, 12A–B) but differs by the sword-shaped median apophysis (Figs 16B–E, 17A–B, D, 18A–B), which is slightly curved with a hook-shaped tip in the latter (Figs 10C–F, 11A–D, F, 12A–B). Terminal apophysis axe-shaped in the new species (Figs 16B–E, 17A–C, 18A– B), knife-shaped in A. hainan Wang, Li & Zhang, sp. nov. (Figs 10C–F, 11A–E, 12A–B). Embolus crescent-shaped in the new species (Figs 16B–E, 17A–C, 18A–B), longer than wide with broad tip in the latter (Figs 10C–F, 11A–E, 12A–B). Description. Male holotype (Fig. 16A) total length 4.40. Carapace 2.55 long, 1.81 wide; opisthosoma 1.91 long, 1.39 wide. Carapace black-brown. Eye region black-brown. Fovea longitudinal. Cervical groove and radial furrows indistinct. Eye sizes and interdistances: AME 0.08, ALE 0.09, PME 0.21, PLE 0.16; AME–AME 0.06, AME–ALE 0.04, PME–PME 0.13, PME–PLE 0.16. Clypeus height 0.06. Chelicerae brown. Labium yellow-brown. Endites yellow-brown, longer than wide. Sternum yellow-brown, with sparse brown setae. Legs yellow-brown, with brown pigmentation. Leg measurements: I 5.23 (1.65, 1.92, 1.01, 0.65); II 5.14 (1.46, 1.76, 1.13, 0.79); III 4.97 (1.39, 1.55, 1.32, 0.71); IV 7.25 (1.87, 2.27, 2.06, 1.05). Dorsum of opisthosoma black-brown, with 5 pairs of white spots, cardiac mark distinct. Ventrum yellow-brown. Palp (Figs 16B–E, 17A–D, 18A–B). Median apophysis sword-shaped, with an anterior furrow. Terminal apophysis longer than wide, distally axe-shaped. Conductor small, membranous. Embolus crescent-shaped with an outgrowth medially. Female unknown. Distribution. Known only from the type locality, Hainan Island, China., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 25-26, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340

Allotrochosina limu Wang, Li & Zhang, sp. nov. (Figs 6A–C, 7A–B) Type material. Holotype ♂, Qiongzhong County, Limu Mountain Nature Reserve, Zhufeng, 19°10.874′N, 109°45.322′E, elev. 940 m, 4.V.2011, Y.Y. Zhou leg. (IZCAS). Etymology. The specific epithet refers to the type locality; noun in apposition. Diagnosis. The new species resembles A. huangi Wang, Li & Zhang, sp. nov. (Figs 3C–G, 4A–D, 5A–C) but differs by the heart-shaped median apophysis (Figs 6B–C, 7A–B,); the median apophysis is liver-shaped in A. huangi (Figs 3C–F, 4A–B, D, 5A–C,). Embolus rodlike, pointed at the end in the new species (Figs 6B–C, 7A–B), crescent-shaped in A. huangi (Figs 3C–G, 4A–C, 5A–B). In retrolateral view, the terminal apophysis is horn-shaped in the new species (Figs 6B–C, 7A– B) but hook-shaped in A. huangi (Figs 3C–G, 4A–C, 5A–C). Description. Male holotype (Fig. 6A) total length 3.25. Carapace 1.78 long, 1.34 wide; opisthosoma 1.40 long, 1.00 wide. Carapace yellow-brown. Eye region black. Cervical groove and radial furrows indistinct. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.18, PLE 0.15; AME–AME 0.05, AME–ALE 0.03, PME–PME 0.11, PME–PLE 0.12. Clypeus height 0.06. Chelicerae yellow-brown. Labium yellowish brown. Endites yellowish brown. Sternum yellowish brown with sparse brown setae. Legs yellow-brown. Leg measurements: I 4.88 (1.33, 1.66, 1.17, 0.72); II 4.81 (1.39, 1.57, 1.14, 0.71); III 4.55 (1.32, 1.39, 1.18, 0.66); IV 6.07 (1.66, 1.84, 1.70, 0.87). Dorsum of opisthosoma yellow-brown, cardiac mark indistinct. Ventrum yellowish brown. Palp (Figs 6B–C, 7A–B). Terminal apophysis sclerotized, horn-shaped, with pointed end. Embolus long, rodlike, with pointed end. Conductor membranous, wider than long, tongue-shaped, with blunt end. Median apophysis sclerotized, heartshaped. Female unknown. Distribution. Known only from the type locality, Hainan Island, China.

Allotrochosina huangi Wang, Li & Zhang, sp. nov. (Figs 1A–B, 3A–I, 4A–D, 5A–E) Type material. Holotype ♂, Ledong County, Jianfengling Nature Reserve, Mingfeng Valley, 18°44.485′N, 108°50.812′E, elev. 960 m, 28.III.2015, L.Y. Wang & G.Q. Huang leg. (SWUC). Paratypes. 9♂ 13♀, same data as holotype (SWUC); 2♂ 4♀, Jianfengling Nature Reserve, Wufengqu, 18°44.415′N, 108°51.802′E, elev. 900 m, 18 May 2011, Y.Y. Zhou leg. (IZCAS). Etymology. The specific name comes from the family name of the collector, Guiqiang Huang; noun (name) in genitive case. Diagnosis. The new species resembles A. schauinslandi (Simon, 1899) (Vink, 2002: 18, figs 7, 34, 41, 68, 95; Framenau, 2008: 467, fig. 7) but differs by the crescent-shaped embolus (Figs 3C–G, 4A–D, 5A–C); the embolus is needle-shaped, and the embolic division has a small beak-shaped projection in A. schauinslandi. Females can be differentiated by the shape and orientation of the spermathecae (Figs 3H–I, 5D–E). Description. Male. Total length 2.66–2.91. Holotype (Fig. 3A) total length 2.85. Carapace 1.60 long, 1.11 wide; opisthosoma 1.24 long, 0.78 wide. Live specimens with a long stripe of white setae from the base of the median eyes to the base of the spinnerets. Carapace yellow-brown. Eye region black. Cervical groove distinct, radial furrows indistinct. Eye sizes and interdistances: AME 0.06, ALE 0.05, PME 0.23, PLE 0.17; AME–AME 0.05, AME–ALE 0.07, PME–PME 0.14, PME–PLE 0.16. Clypeus 0.08 high. Chelicerae yellow-brown, with 3 promarginal and 2 retromarginal teeth. Labium yellowish brown. Endites yellow-brown. Sternum yellowish brown and scutellate, with sparse, yellow-brown setae. Legs yellowish brown. Leg measurements: I 4.11 (1.03, 1.36, 1.08, 0.64); II 3.70 (1.04, 1.28, 0.87, 0.51); III 3.69 (1.02, 1.15, 1.01, 0.51); IV 5.20 (1.30, 1.66, 1.50, 0.74). Dorsum yellow-brown, with black pigmentation. Ventrum yellowish brown. Palp (Figs 3C–G, 4A–D, 5A–C). Terminal apophysis small, hook-shaped, covered by conductor. Median apophysis liver-shaped, slightly sclerotized. Embolus crescent-shaped. Conductor wide, membranous, tapering distally. Female (paratype, Fig. 3B) total length 5.37. Carapace 2.52 long, 1.70 wide; opisthosoma 2.85 long, 1.71 wide. Eye sizes and interdistances: AME 0.05, ALE 0.06, PME 0.26, PLE 0.20; AME–AME 0.05, AME–ALE 0.10, PME–PME 0.15, PME–PLE 0.21. Clypeus height 0.08. Leg measurements: I 4.97 (1.43, 1.70, 1.12, 0.72); II 4.66 (1.32, 1.56, 1.11, 0.67); III 4.61 (1.30, 1.53, 1.19, 0.59); IV 6.55 (1.74, 2.07, 1.85, 0.89). Leg formula: 4123. Opisthosoma oval. Dorsum of opisthosoma yellow-brown, with black pigmentation. Ventrum yellow-brown. Epigyne (Figs 3H–I, 5D–E). Spermathecal heads round. Spermathecal stalks V-shaped, joined at the base, separated from each other by two times the length of a fertilization duct. Fertilization ducts relatively large, crescent-shaped. Distribution. China (Hainan)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 17-21, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Vink, C. J. 2002. Lycosidae (Arachnida: Araneae). Fauna of New Zealand, 44: 1 - 94.","Framenau, V. W. 2008. A new species in the wolf spider genus Allotrochosina from New South Wales, Australia (Araneae, Lycosidae). Journal of Arachnology, 35: 463 - 469.","Tanaka, H. 1985. Descriptions of new species of the Lycosidae (Araneae) from Japan. Acta Arachnologica, Tokyo, 33: 51 - 87."]}

Zoica hainan Wang, Li & Zhang, sp. nov. (Figs 72A–H, 73A–D, 74A–D) Type material. Holotype ♂, Haikou City, Dongzhaigang Mangrove Nature Reserve, 19°57.113′N, 110°33.983′E, elev. - 2 m, 20.VIII.2007, C.X. Wang leg. (IZCAS); Paratypes (7♂ 8♀). 4♂ 4♀, same data as holotype (IZCAS); 1♂ 3♀, Haikou City, Dongzhaigang Mangrove Nature Reserve, 19°57.113′N, 110°33.983′E, elev. - 2 m, 20.VIII.2007, S. Li leg. (IZCAS); 2♂ 1♀, Ding’an County, Tongren Village, 19°30.953′N, 110°18.904′E, elev. 58 m, 27. VI.2019, L.Y. Wang, T. Yuan, K. Yu & J.X. Zhao leg. (SWUC). Etymology. The specific epithet refers to the type locality; noun in apposition. Diagnosis. The new species resembles Z. unciformis Li, Wang & Zhang, 2013 (Li, Wang & Zhang, 2013: 30, figs 5A– E, 6A–H) but differs by the lateral apophysis with two small projections, the inner projection spinelike, the outer projection longer than wide with a sharp tip (Figs 72C–F, 73A–D, 74A–B); in Z. unciformis, the lateral apophysis is sinuous, slightly curved, with a small outgrowth. Terminal apophysis covers embolus in the new species (Figs 72C–F, 73A–D, 74A–B) but not in Z. unciformis. Spermathecae round in the new species (Figs 72G–H, 74C–D) but oval in Z. unciformis. Description. Male. Total length 0.99–1.39. Holotype (Fig. 72A) total length 1.05. Carapace 0.56 long, 0.40 wide; opisthosoma 0.50 long, 0.33 wide. Carapace uniformly yellow-brown. Eye region black. Eye sizes and interdistances: AME 0.08, ALE 0.08, PME 0.12, PLE 0.09; AME–AME 0.02, AME–ALE 0.02, PME–PME 0.05, PME–PLE 0.04. Clypeus height 0.07. Chelicerae elongate, yellowish brown, with three promarginal and two retromarginal teeth. Labium yellowish brown. Endites yellowish brown. Sternum yellowish brown, with sparse brown setae. Legs yellowish brown. Leg measurements: I 2.57 (0.79, 0.93, 0.39, 0.46); II 2.41 (0.72, 0.82, 0.47, 0.40); III 1.69 (0.50, 0.43, 0.31, 0.45); IV 3.20 (0.91, 1.06, 0.76, 0.47). Opisthosoma brown. Venter yellowish brown. Palp (Figs 72C–F, 73A–D, 74A–B). Lateral apophysis with two small projections, inner projection spinelike, outer projection longer than wide with sharp tip. Terminal apophysis leaf-shaped with pointed tip. Conductor wide, membranous, with a pointed tip. Embolus stout, covered by terminal apophysis. Female (paratype, Fig. 72B) total length 1.40. Carapace 0.68 long, 0.49 wide; opisthosoma 0.65 long, 0.49 wide. Eye sizes and interdistances: AME 0.07, ALE 0.08, PME 0.13, PLE 0.09; AME–AME 0.02, AME–ALE 0.02, PME–PME 0.05, PME–PLE 0.06. Clypeus height 0.05. Leg measurements: I 2.75 (0.84, 0.90, 0.55, 0.46); II 2.58 (0.78, 0.85, 0.53, 0.42); III 2.54 (0.79, 0.80, 0.54, 0.41); IV 3.16 (0.89, 1.13, 0.75, 0.39). Opisthosoma yellow-brown. Epigyne (Figs 72G–H, 74C–D). Copulatory openings located posteriorly, separated by the diameter of the opening. Spermathecal heads round. Spermathecal stalks wide. Fertilization ducts long and hooklike. Distribution. Known only from the type locality, Hainan Island, China. Funding This work was supported by the National Natural Science Foundation of China (31672278, 31702005), the Key Natural Science Foundation of Chongqing (cstc2019jcyj-zdxmX0006) and the Investigation Project of Basic Science and Technology (2018 FY100305) to Zhisheng Zhang and Luyu Wang. Acknowledgements The manuscript benefited greatly from comments by two anonymous referees. Sarah Crews checked the English. Theo Blick checked etymology. We thank Feng Zhang and Tianlang Lv for the loan of specimens and Guiqiang Huang for his assistance during field work and collection., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 70-72, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340

Blepephaeus taomeiae (Hayashi, Nara & Yu, 1995) comb. nov. K��������+ (Figure 6) Uraechoides taomeiae Hayashi et al., 1995: 149 (type locality: ���Luen Din, Chiai, Taiwan, China ���); Chou 2008: 297, figs (male and female); L��bl & Smetana 2010: 288 (catalogue); Lin & Yang 2019: 324 (catalogue). As stated above, Uraechoides Breuning, 1981 is treated as a junior synonym of Paracyriothasastes Breuning, 1978, therefore, Uraechoides taomeiae Hayashi, Nara & Yu, 1995 should be treated in Paracyriothasastes. Paracyriothasastes can be distinguished from Blepephaeopsis by having the prothorax with lateral tubercles near the base (prothorax with lateral tubercles near the middle in Blepephaeopsis), elytron with an acute marginal angle at the apex (elytron with a weak marginal angle at the apex in Blepephaeopsis). Paracyriothasastes taomeiae (Hayashi, Nara & Yu, 1995) has characters of Blepephaeus, besides, examining the holotype photographs of U. taomeiae (Fig. 6), Blepephaeus fulvus (Pic, 1933) (Fig. 7) and B. nigrostigma Wang & Chiang, 1998 (Fig. 8), we found that U. taomeiae is extremely similar to Blepephaeus fulvus and B. nigrostigma in following characters: elytra densely covered with brown hairs forming four large maculae (an inverted triangular macula basally, a pair of arcuate maculae laterally and a sub-triangular macula apically), with a large X-shaped grayish white haired band among above four maculae, sparsely with sub-rounded small black haired maculae on above four maculae, prothorax with lateral tubercles near middle, marginal angles weak. We thus transfer U. taomeiae to the genus Blepephaeus Pascoe, 1866. Type material examined. Holotype (MSME), Luen Din, Chiai, Taiwan, China, 13 May 1991, Taomei Li leg. Male. Examined from a modified photograph (Fig. 6) from original paper of Hayashi et al. (1995). Distribution. China (Taiwan)., Published as part of Huang, Gui-Qiang, Zhao, Xin-Xia, Yan, Kai, Li, Song & Zhang, Gui-Mei, 2020, Notes on the genus Paracyriothasastes Breuning, 1978 (Coleoptera: Cerambycidae Lamiinae: Lamiini), pp. 142-146 in Zootaxa 4868 (1) on page 145, DOI: 10.11646/zootaxa.4868.1.9, http://zenodo.org/record/4417449, {"references":["Hayashi, M., Nara, H. & Yu, S. - K. (1995) A new species of genus Uraechoides Breuning (1981) from Taiwan (Cerambycidae, Lamiinae). Chinese Journal of Entomology, Tao-Yuan, 15, 149 - 151.","Chou, W. I. (2008) The Atlas of Taiwanese Cerambycidae (second edition). Owl Publishing House, Taipei, 408 pp.","Lobl, I. & Smetana, A. (2010) Catalogue of Palaearctic Coleoptera. Vol. 6. Chrysomeloidea. Apollo Books, Stenstrup, 924 pp. https: // doi. org / 10.1163 / 9789004260917","Lin, M. & Yang, X. (2019) Catalogue of Chinese Coleoptera. Vol. 9. Chrysomeloidea: Vesperidae, Disteniidae, Cerambycidae. Science Press, Beijing, 575 pp.","Breuning, S. (1981) Nouvelles especes de Lamiinae (Col. Cerambycidae) de la coll. Vives. Nouvelle Revue d' Entomologie, Paris, 11 (1), 73 - 75."]}

Paragnia tiani Huang sp. nov. moeḦĦK+ Figures 27–44 Description. Male. Body length: 12.0 mm, humeral width: 4.5–5.0 mm (2 individuals); holotype (Figs 27–43), body length: 12.0 mm, humeral width: 5.0 mm. Body reddish brown, antennomeres V–X (except for apex) and antennomere XI yellowish brown; antennomeres III–X dark brown apically; prothorax (except for prosternum), most parts of elytra, sides of meso- and metaventrite and urosternite I (except for base) black; urosternites IV–V yellowish brown (Figs 27–29). Frons sparsely covered with short grayish-white and black setae (Fig. 30). Scape, pedicel and middle of antennomere XI sparsely covered with short black setae; apical half of antennomere IV with moderately dense short black setae, antennomeres IV–IX with several long black setae ventrally, scape, pedicel and base of antennomere III with sparse short grayish-white setae, basal half of antennomere IV with moderately dense short light-yellow setae (Figs 27–29). Prothorax with a yellow setal band at laterobasal half (Figs 27, 29). Each elytron with a longitudinal yellow setal band on the basal 1/3 close to the suture and an irregular yellow setal spot on inner margin of apex (Fig. 27). Prosternum and mesoventrite sparsely covered with short grayish-white setae (Fig. 28). Antennae 1.4 times as long as body, antennal formula (ratio) based on length of antennomere III: scape= 0.60, pedicel= 0.10, IV= 0.87, V= 0.53, VI= 0.47, VII= 0.40, VIII= 0.35, IX= 0.32, X= 0.28, XI= 0.38. Inner side of antennomeres V–IX slightly projected near base (Figs 27–28). Elytra 2 times as long as wide at base. MP 3+4 and MP 4 veins of hind wing far from each other (Fig. 32). Margin of urosternite V slightly notched medially (Fig. 28). Male terminalia. Tergite VIII (Figs 33–34) sub-semicircle, with a feebly triangular projection at apical middle. Stem of spiculum gastrale more than 2 times as long as branches (Figs 35–37), curved near basal 1/ 3 in lateral view (Fig. 36). Tegmen (Figs 38–40) shorter than penis, dorsum of parameres sparsely covered with short fine brown setae and several long thick brown setae near base and apex, phallobase expanded near the apical 1/3 and gradually tapered to base, then curved near the basal 1/ 3 in lateral view (Fig. 39), tegminal struts distinctly expanded towards inside near apex (Figs 38, 40). Venter of dorsal struts of penis distinctly constricted near apical 1/ 4 in lateral view (Fig. 42). Female. Unknown. Diagnosis. Males of this new species can be distinguished from Paragnia fulvomaculata by the prothorax with a yellow setal band on the laterobasal half (without yellow setal band in P. fulvomaculata); the long yellow setal bands on basal middle and basal margin of elytra (short in P. fulvomaculata); each elytron with a yellow setal band on the basal 1/3 close to suture (with several yellow setal spots in P. fulvomaculata), with a yellow setal spot on inner margin of apex (with sparse short light yellow hairs in P. fulvomaculata); antennae short (long in P. fulvomaculata), antennomeres III–IV thicker (thin in P. fulvomaculata); CuA 2 vein of hind wing short (long in P. fulvomaculata), the distance between MP 3+4 and MP 4 veins long (short in P. fulvomaculata), sr vein missing (with sr vein in P. fulvomaculata); ventrites slightly notched at apical middle (sub-truncated in P. fulvomaculata); tergite VIII with a feebly triangular projection at the apical middle (truncated in P. fulvomaculata); stem of spiculum gastrale narrow (wide in P. fulvomaculata), membrane connecting both parameres wide (narrow in P. fulvomaculata); tegminal struts distinctly expanded towards inside near apex (not expanded in P. fulvomaculata); venter of dorsal struts of penis distinctly constricted near apical 1/ 4 in lateral view (not constricted in P. fulvomaculata). Type material. Holotype: male (LPSNU), VIETNAM. Dak Lak, IV.2019, local collector leg. Paratype: 1 male (LPSNU), same data as the holotype, but III.2018. Etymology. This new species is dedicated to Prof. Ying-Zhou Tian (mLJNJ), former president of Liupanshui Normal University, for his generous supports to this study. Distribution. Only known from type locality of Dak Lak province, Vietnam., Published as part of Huang, Gui-Qiang, Li, Song & Zhang, Gui-Mei, 2020, A review of the genus Paragnia Gahan, 1893 (Coleoptera: Cerambycidae: Lamiinae: Lamiini), pp. 159-170 in Zootaxa 4778 (1) on pages 166-168, DOI: 10.11646/zootaxa.4778.1.6, http://zenodo.org/record/3823784

Allothrips yunnanensis sp.n. (Figs 1���6) Female aptera. Body legs and antennae brown with head and antennal segment III paler than the abdomen; abdominal segment IX yellow (Fig. 4), tube golden with grey terminal ring; major setae pale. Head projecting in front of eyes, with genae weakly concave, bearing small fine setae (Fig. 1); postocular setae long and capitate; postocellar setae shorter and capitate; mid-dorsal setae small and finely pointed, arising just posterior to a line between the bases of the postoculars; ocelli usually small; vertex without sculpture except reticulation on posterior third and in ocellar region; eyes ventrally with less than 10 facets, dorsally with at least 15 of which three posterolaterally are unusually large; maxillary stylets broad, retracted almost to level of eyes and less than one third of head width apart medially; maxillary palp segment II long with prominent terminal sensorium. Antennae 7-segmented (Fig. 2); setae pointed, except 2 pairs on II with apices blunt; 2 slender sense cones on each of III���V. Pronotum with 5 pairs of weakly capitate major setae; surface strongly reticulate only near posterior margin; notopleural sutures complete. Fore tarsus with no tooth. Prosternal basantra not developed although several chitinous islets are sometimes close together but not forming a discrete sclerite; ferna almost meeting medially; mesopresternum slender medially. Mesonotum reticulate, strongly transverse (Fig. 3), with 2 pairs of capitate setae laterally; marginal setae usually pointed, median marginal setae rarely weakly capitate. Metanotum reticulate, median setal pair wide apart, weakly capitate. Pelta almost complete across anterior margin of tergite II (Fig. 3), anterior third reticulate but less strongly so than posterior area. Tergites II���VII reticulate on anterior half, with transverse row of small fine setae, posterior margins with 2 pairs of capitate major setae; tergite IX setae S1 capitate, S2 pointed; tube almost twice as long as IX. Sternites II���VII with median transverse row of small fine setae, posteromarginal median setae long and pointed. Measurements (holotype in microns). Body length 2250. Head, length 250; maximum width 215; postocular setae 65; postocellar setae 30. Pronotum, length 150; width 280; major setae, anteromarginal 40, anteroangular 35, midlateral 50, epimeral 50, posteroangular 45. Tergite IX setae S1 80; setae S2 90. Tube length 180. Antennal segments III���VII length 70, 50, 50, 50, 70. Female macroptera [only de-alates available]. Similar to aptera in colour and structure, including the number of compound eye facets, but differing in: ocelli well developed; meso and metanotum less transverse (Fig. 5); fore wing with 3 capitate sub-basal setae arranged in a triangle; tergites IV���VII (Fig. 6) each with one pair of weakly sigmoid wing-retaining setae (II���III with this pair of setae minute and straight). Male aptera. Similar to female aptera except: compound eyes with about 8 facets dorsally and 4 ventrally; ocelli absent; fore tarsal tooth present; large male with fore femora swollen and tarsal tooth almost as long as tarsal width; sternites without pore plates. Measurements (large paratype male in microns). Body length 2100. Head length 230. Tube length 175. Specimens studied. Holotype female aptera, CHINA, Yunnan, Kunming City, Arboretum, (N 25�� 08��� E 102�� 43���, 2006m asl), from leaf litter of Pinus yunnanensis, 10.iii.2011 (Jun Sun & Tao Li) in Yunnan Agricultural University, Kunming, China. Paratypes, all from leaf litter and dead leaves at same site as holotype, between March and November 2011: 5 female apterae, 2 female macropterae collected with holotype; 3.iii.2011, 2 female apterae, 2 female macropterae from Magnolia denudata, 2 female apterae from Cinnamomum camphora; 21.iii.2011, 6 female apterae from Pinus yunnanensis; 25.iii.2011, 2 female apterae from Sabina chinensis; 14.iv.2011, 1 male aptera from Eucalyptus; 20.v.2011, 1 female aptera from Cinnamomum camphora, 2 female macropterae, 4 female apterae, 1 male aptera from Pinus yunnanensis; 14.vii.2011, 1 female macroptera from Pinus yunnanensis; 11.viii.2011, 1 female aptera, 1 female macroptera from Eucalyptus robusta; 2.x.2011, 25 female apterae from dead leaves; 25.xi.2011, 3 female apterae from Sabina chinensis. [all deposited in Yunnan Agricultural University Collection, Kunming, with 11 slides in Australian National Insect Collection, Canberra]. Comments. This new species shares with brasilianus the anterior position of the mid-dorsal setae on the head. However, in addition to the differences noted in the key above the number of facets in the compound eyes of female apterae is not smaller than in macropterae (cf. Okajima 2006: 51)., Published as part of Mound, Laurence, Li, Yajin & Zhang, Hongrui, 2020, Species of the genus Allothrips (Thysanoptera, Idolothripinae) from Southeast Asia, with one new species from Yunnan, China, pp. 593-595 in Zootaxa 4732 (4) on pages 594-595, DOI: 10.11646/zootaxa.4732.4.10, http://zenodo.org/record/3667312, {"references":["Okajima, S. (2006) The Insects of Japan. Volume 2. The suborder Tubulifera (Thysanoptera). Touka Shobo Co Ltd., Fukuoka, 720 pp."]}

As a little girl, Li Zhang was brisk and outgoing, and always spoke cheerfully which made her families believe she was especially congenial to be a doctor. Followed the advices of the elders who were doctors as well, the innocent Li Zhang chose the road of Medicine.

The ancient Greek aphorism “know thyself” engraved on the forecourt of the Temple of Apollo epitomizes the human desire for the exploration into the unknown. In fact, never has mankind dragged their feet in understanding themselves, as in the achievements scientists have made all these years in the studies of disease and pathology.

Understanding China’s Urbanization is a massive compendium of facts and knowledge regarding China’s post-1980s rapid urbanization. It reflects the research findings of faculty at Tongji University,...

Bracon (Glabrobracon) planitibiae Yang, Cao et Gould, sp. nov. (Figs 7–15) Type materials. Holotype, 1♀, China, Beijing City, Haidian Distinct, Minzhuang Road, N39°58′15″, E116°12′45″, 2016.VIII.4, reared from a cocoon collected from the gallery of ALB first instar larva in the sentinel logs hanging in willow forest for one month, Yang Zhongqi & Li Fei leg. Paratypes, 6♀♀, same data as holotype. Female. Body length 2.79–3.33 mm, fore wing length 3.02–3.15 mm. Color. Body black. Two spots under antennal toruli yellow (Fig. 9); sides of first tergite white; malar suture and adjacent area, basal mandible, tibial spurs, basal tibia and apical tarsus yellow to dark yellow. Wings hyaline, pterostigma brown, veins light brown. Setae of head, mesopleuron, mesoscutum, propodeum golden. Head (Figs 7–10, 12). Median length 2/3 of its width in dorsal view; vertex smooth, with sparse long setae, but setae on occiput shorter; eyes large, transverse diameter 1.63 × length of temple in lateral view; face with dense long setae and reticulate; front clypeal suture bent intensely, anterior tentorial pit large bilaterally; clypeus narrow, oral cavity deeply depressed; malar suture 1/6 of height of eye and 1/5 basal width of maxillary palp; mandible strong with long thick setae, upper margin with one big tooth; antenna 28 antennomeres (Figs 7, 8, 10), scape twice of pedicel length, antenna about 0.75 × body length, and 0.79 × fore wing length. Mesosoma (Figs 7, 8, 11, 12). In lateral view, length 1.48 × maximum height; pronotum depressed in the middle, smooth and shining; propleuron with long setae marginally; mesopleuron smooth, broad and swollen downwards, with sparse long setae medially; in dorsal view, mesoscutum smooth with sparse long setae; notauli depressed without any sculpture, mid pit lacking; scutellar sulcus narrow and deep, with 12–15 round sculptures; apical and lateral scutellum with dense long golden setae; length of metanotum 0.29 × propodeum length; propodeum smooth, most central part with dense long setae bilaterally. Legs (Figs 7, 8, 14). Fore trochantellus 0.4 × length fore femur, the latter as long as fore tibia and about 0.6 × length fore tarsus; apical spur of fore tibia 1/2 of basitarsus, each tarsomere with thick setae apically, length ratio of fore tarsomeres I–V = 2:1:0.8:0.5:1. Mid tibia slightly longer than femur, apical two spurs 1/3 length basitarsus, length ratio of mid tarsomeres I–V = 1.5:0.7:0.6:0.5:0.7. Hind femur depressed, 2.8 × as long as broad, with inner side very flattened and outer side only evenly convex, 2.4 × as wide as thick; hind tibia very flattened and 1.3 × the femur, 6.4 × as long as broad, and 1.8 × as wide as thick; hind tarsomere 1–4 considerably flattened bilaterally. Wings (Fig. 13). Fore wing 2.84 × as long as its maximum width; pterostigma 2.6 × as long as its width; 1- R 1 about 1.40 × length of pterostigma; radial vein arising from middle pterostigma; SR1 about 2.0 × length of 3-SR; radial vein about 0.65 × length of 2-SR, and 0.58 × length of 3-SR; cu-a vein straight, 1-CU1 vein invisible; m-cu vein enters into first submarginal cell; 2-SR+M and basal 2-M weakly sclerotized; 1-SR+M vein straight; 3-M and CU1a vein reaching apex of fore wing; 3-CU1 vein bent. Hind wing 3.5 × as long as its maximum width; cu-a vein 0.63 × length of M+CU, and 0.3 × length of 1-M; 2-M and SR intensely sclerotized. Metasoma (Figs 7, 8, 15). Oblong, first tergite quadrate, with two rows of oblique punctations, spiracles located at 1/3 basally, triangular broadening side as wide as first tergite; tergites 2–7 smooth, with even long golden setae; suture between tergites 2–3 deep, with granulose punctations. Ovipositor sheath short, as long as hind tibia; ovipositor golden and sharp. Male. Unknown. Distribution. China (Beijing). Etymology. The new species name is derived from its flattened hind tibia. Diagnosis. The new species is similar to B. (Glabrobracon) nigriventris Wesmael, which is also a parasitoid of cerambycids, and the two species can be separated with the key below. 1 Face with two yellow spots under antennal toruli (Fig 9); propodeum completely polished and shining (Fig 11); hind femur depressed, only 2.8 ×as long as broad; hind tibia and hind tarsomere 1–4 considerable flattened; ovipositor sheath short, as long as hind tibia (Fig 8)............................................................... Bracon planitibiae sp. nov. - Face without yellow spots under antennal toruli (Papp, 2012); propodeum having short rugae above lunule (Papp, 2012 p62, Fig 28C); hind femur 3.0 × as long as broad; tibia and tarsomere 1–4 normal and not flattened; ovipositor sheath long, as long as hind tibia + basitarsus combined (Papp, 2012)..................................... Bracon nigriventris Wesmael Biological observations. The new species is a solitary ectoparasitoid of first instar larvae of Anoplophora glabripennis. Percentage parasitism averaged 4.0% during the three year study, with a maximum of 5.9% in 2017. The newly emerged larva of the parasitoid is attached to the host body. In only seven days the host was completely consumed and the mature parasitoid larva made a cocoon beside the mummy of the host. The pupal stage of the parasitoid lasted 18–20 days at a temperature of approximately 25°C.

Schizocyrtillus fuzhouensis sp. nov. Diagnosis. Adult female. Dorsum oval, truncate anteriorly and broadly rounded posteriorly; smooth, covered with more than 100 simple setae and numerous irregularly placed small circular pores; sternal plate with three pairs of simple setae, two pairs of pores and network of sculpture lines; metasternal plates narrow, lying between coxae II and III, bearing one pair of setae; ventral, anal, parapodal and metapodal shields fused, forming genito-ventrianal shield; anterior margin of this shield convex, with notched protuberance medially; its surface smooth, with 28 pairs of simple setae and numerous irregularly placed circular pores; post-anal plate more than six times as wide as long, with one pair of setae. Type material. Holotype Female, China, Lianjiang county, Fuzhou city, Fujian province, 27 October 2016, by Yun Xu, from rotten wood and bark of Pinus massoniana Lamb. Paratypes: 1 female, same data as holotype. The holotype and paratype are deposited in the Department of Plant Protection, Fujian Agriculture and Forestry University, China. Female (n=2). Idiosomal dorsum (Fig. 5): Shape oval, truncate anteriorly and broadly rounded posteriorly, 540 length, 395 at width level of coxae IV; smooth, covered by a single shield with more than 100 simple setae; all setae subequal in length (25���35); surface with numerous irregularly placed small circular pores. Marginal plate with a single row of simple setae. Idiosomal venter (Fig. 6). Tritosternal base adjacent to anterior margin of sternal plate, laciniae free and pilose. Sternal shield wider than long, both anterior and posterior margins concave. Sternal plate with three pairs of simple setae (st 1 ��� st 3), st 1 placed anteriorly and st 2 ��� st 3 inserted at the same level posteriorly, shield with two pairs of pores, and a network of sculpture lines. Metasternal plates narrow, lying between coxae II and III, bearing setae st 4. Pore III slit-shaped, located anterior to setae st 4. Posterior margins of metasternal plates sinuous, with heavily sclerotised indentations to accommodate heads of vaginal sclerites. Ventral, anal, parapodal, peritremal and metapodal shields fused, forming genito-ventrianal shield. Anterior margin of this shield convex, with notched protuberance medially. Shield surface smooth, with 28 pairs of simple setae and numerous irregularly placed circular pores. Stigmata lateral to coxae IV. Peritremes straight, extending to anterior margin of coxae II. Ventrimarginal shields border ventrianal shield from level of coxae II to posterior margin; shields posteriorly separated from one another by post-anal plate. Each ventrimarginal shield with three simple setae posteriorly. Postanal plate (16 length, 105 width) more than six times as wide as long, with one pair of setae. Legs (Fig. 7). All legs with simple setae. Leg I without claws. Leg II much stouter than others. Genua III and IV each with one dorsal macroseta. Leg I���IV chaetotaxy (from coxa to tarsus): I 2-5-9-9-12 -31; II 1-4-12-9-7 -19; III 1- 4-7-8-7-18; IV 0-5- 4-8-7-21. Gnathosoma (Figs 8���10). Tectum as in Fig. 9. Fixed digit of chelicera with two teeth on the cutting edge and 13���14 teeth on the lateral face. Movable digit with eight teeth, last tooth much larger than others; with two branched excrescences ventrally. Hypostome with four pairs of setae; corniculi deeply divided and trifurcate. Male and immatures unknown. Etymology. The species is named after the type locality, Fuzhou. Remarks. The adult female of Schizocyrtillus fuzhouensis sp. nov. is easily distinguishable from those of the other three species by the following: dorsal dorsum smooth, lacking big pores (dorsal dorsum smooth, covered with three to four pairs of big pores in Schizocyrtillus lathrius Kinn, 1970 and Schizocyrtillus rarus Khaustov, 1999, dorsal dorsum covered with reticular ornamentation, lacking big pores in Schizocyrtillus josefinae Gwiazdowicz, 2002); genito-ventrianal shield smooth, with 28 pairs of simple setae (shield smooth, with 17 pairs of setae in S. lathrius and 31 setae in S. rarus, shield with reticular ornamentation and 14 pairs of setae in S. josefinae (according to the Fig. 2 in Gwiazdowicz, 2002); post-anal plate more than six times as wide as long (post-anal plate about 1.5 times as wide as long in S. lathrius, four times as wide as long in S. josefinae and three times as wide as long in S. rarus)., Published as part of Xu, Yun, Li, Xiang & Zhang, Fei-Ping, 2019, Two new records of the Family Celaenopsidae (Acari: Mesostigmata) from China, with description of a new species, pp. 326-334 in Zootaxa 4604 (2) on pages 327-333, DOI: 10.11646/zootaxa.4604.2.5, http://zenodo.org/record/2807230, {"references":["Kinn, D. N. (1970) New genus of Celaenopsidae from California with a key to the genera (Acari: Mesostigmata). Pan Pacific Entomologist 46 (2), 91 - 95.","Khaustov, A. A. (1999) A new species of the genus Schizocyrtillus (Acarina: Mesostigmata: Celaenopsidae) from Crimea. Acarina, 7, 107 - 109.","Gwiazdowicz, D. J. (2002) Schizocyrtillus josefinae n. sp. of the family Celaenopsidae (Acari, Antennophorina) from Poland. Acarologia, 42, 21 - 27."]}

Stegana (Steganina) flavivittata sp. nov. (Figs 2J, 13) Diagnosis. This species resembles S. flavipalpata Chen & Chen in Lu et al. 2011a in the yellow clypeus and palpus, and colour patterns of body (Fig. 2J), but can be distinguished by the aedeagus lacking serrated, minute processes basally in lateral view, and with round opening in ventral view (Fig. 13B, C). The minimum interspecific genetic distance to S. flavipalpata is 6.9 % (Table 2). ������continued on the next page Min. intra., minimum intraspecific distance; Max. intra., maximum intraspecific distance; Mean intra. Vari., mean intraspecific variability (standard deviation, SD); Min. inter., minimum interspecific distance; Max. inter., maximum interspecific distance; Mean inter. Vari., mean interspecific variability (SD); NA, not applicable. Description. Male and female. Frons black, with orange yellow stripe medially and band submedially. Frontoorbital plate orange yellow. First flagellomere medially grayish yellow on basal 2/3, black on distal 1/3. Face black on upper 2/3, yellow on lower 1/3. Mesonotum medially brown and with yellow, thin longitudinal stripe, laterally dark brown, with yellow longitudinal stripes and a few patches (Fig. 2J). Katepisternum entirely white (Fig. 2J). Scutellum brownish yellow medially, dark brown to balck laterally (Fig. 2J). Veins R 4+5 and M 1 lacking setae on basal section of dorsal surface. Legs white, brownish knee articulations of fore and hindlegs, and black on distal half of hindleg, distal half of femur and basal half of tibia of midleg. Fore femur distally with 6���8 setae on posteroventral surface. Abdominal 1st to 5th tergites dark brown to black medially, yellow laterally; 6th tergite nearly black. Sternites nearly yellow. Male terminalia (Fig. 13 A���F): Distal part of 10th sternite diamond-shaped, about 1.85 times as wide as long. Hypandrium anteromedially elongated, as long as width of epandrium and with a few paramedian setae posteroventrally. Gonopods slightly rounded apicoomedially. Aedeagus about 0.5 times as long as aedeagal apodeme. Female terminalia (Fig. 13 G���I): Epiproct, cercus and hypoproct with dense setae and pubescence. Eighth sternite mostly glabrous, with several setae and pubescence on posterolateral corners. Measurements. BL = 4.87 mm in holotype (mm in 5♂ and 5♀ paratypes: 3.20���4.67 in ♂, 4.13���4.53 in ♀), ThL = 2.53 mm (1.73���2.40 in ♂, 2.07���2.27 in ♀), WL = 3.47 mm (2.53���3.67 in ♂, 3.00��� 3.47 in ♀), WW = 1.73 mm (1.20���1.80 in ♂, 1.33���1.87 in ♀), arb = 10/6 (8���10/4���6), avd = 0.67 (0.67���0.83), adf = 1.80 (1.50���2.00), flw = 1.50 (1.30���2.13), FW/HW = 0.42 (0.36���0.48), ch/o = 0.21 (0.20���0.36), prorb = 0.65 (0.67���1.00), rcorb = 0.87 (0.60���1.10), orbito = 2.14 (1.50���2.33), vb = 0.40 (0.18���0.45), sctl = 1.69 (1.29���2.50), sterno = 1.00 (0.50���0.94), sctlp = 1.25 (1.67���2.50), C = 2.80 (1.59���2.92), 4c = 0.83 (1.00���1.38), 4v = 1.92 (1.77���2.50), 5x = 1.75 (1.50��� 2.50), ac = 5.00 (5.50���9.00), M = 0.58 (0.58���1.00), C3F = 0.80 (0.67���0.83). Specimens examined. Holotype ♂ (SCAU, No. 111489), China: Hesong, Menghai, Yunnan, 21��49��� N, 100��06��� E, altitude 1800m, 7.v.2012, tree trunk, HW Chen. Paratypes: China: 6♂, 5♀ (SCAU, Nos 111490 ���500), same data as holotype; 1♂, 4♀ (SCAU, Nos 111501 ���05), altitude 1600m, 12.v.2012, same data as holotype; 1♂, 1♀ (SCAU, Nos 111506, 07), Mengyuan, Mengla, Yunnan, 21��43'56''N, 101��23'15''E altitude 876m, 18.xii.2017, tree trunks, HW Chen; 1♀ (SCAU, No. 111508), Wangtianshu Forest Park, Mengla, Yunnan, 21��36'9''N, 101��35'6''E, altitude 640m, 16.iv.2010, tree trunk, HW Chen; 4♀ (SCAU, Nos 111509 ���12), Menglai Grand Canyon, Cangyuan, Yunnan, 23��13'20''N, 99��15'4''E, altitude 1240m, 30.x.2017, tree trunks, HW Chen, L Gong, BX Li; 3♂, 1♀ (KIZ, Nos 0090665���68), Longtan Forest Park, Ximeng, Yunnan, 22��38'44''N, 99��35'39''E, altitude 1145m, 27.x.2017, tree trunks, HW Chen, L Gong, BX Li; 1♂ (SCAU, No. 111513), Moli Forest Park, Ruili, Yunnan, 24��7'5''N, 97��58'51''E, altitude 920m, 5.xi.2017, tree trunk, L Gong; 1♂, 8♀ (SCAU, Nos 111514 ���22), Yixiang, Pu'er, Yunnan, 22��42'N, 101��09'E, altitude 1400m, 13.v.2012, tree trunks, HW Chen; 1♀ (SCAU, No. 111539), Tongbiguan, Yingjiang, Yunnan, altitude 1310m, 19.v.2016, tree trunks, J Huang; 2♀ (SCAU, Nos 111523, 24), Husa, Longchuan, Yunnan, 24��25'19''N, 97��46'38''E, altitude 1227m, 20,21. viii.2016, tree trunks, HW Chen, L Gong; 2♂, 5♀ (SCAU, Nos 111525 ���1), Bangda, Mangshi, Yunnan, 24��7'55''N, 98��13'42''E, altitude 1780m, 8.xi.2017, tree trunks, HW Chen, L Gong, BX Li; 1♀ (SCAU, No. 111532), Botanic Garden, Ruili, Yunnan, 24��4'19''N, 97��49'8''E, altitude 1174m, 22.v.2016, tussock, J Huang; 1♂ (SCAU, No. 111542), Mengdong, Cangyuan, Yunnan, 23��10'8''N, 99��13'52''E, altitude 1320m, tussock, 6.V.2016, J Huang. Etymology. A combination of the Latin words: flavus and vittatus, referring to the mesonotum with yellow longitudinal stripes. Distribution. China (Yunnan)., Published as part of Li, Bingxue, Zhang, Yuan & Chen, Hongwei, 2019, Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae), pp. 81-108 in Zootaxa 4576 (1) on pages 103-107, DOI: 10.11646/zootaxa.4576.1.4, http://zenodo.org/record/2624686, {"references":["Lu, J. M., Gao, J. J., Chen, X. P. & Chen, H. W. (2011 a). The Stegana undulata species group (Diptera, Drosophilidae), with molecular phylogenetic analysis of the Chinese species. European Journal of Entomology, 108, 139 - 152. https: // doi. org / 10.14411 / eje. 2011.017"]}

Stegana (Steganina) hirtifoliacea sp. nov. (Figs 2G, 10) Diagnosis. This species resembles S. unguiculata sp. nov. in the colour patterns of body (Fig. 2G) and male terminalia (Fig. 10), but can be distinguished by the aedeagus with 4 short and thick processes per side (Fig. 10C); sublateral processes of gonopods shorter and broadly separated than in S. unguiculata sp. nov. (Fig. 10D). The minimum interspecific genetic distance to S. unguiculata is 3.3%, which is smaller than to other species (Table 2); see description of S. unguiculata for more detail. Description. Male: Pedicel yellow medially, black laterally; first flagellomere nearly black. Palpus yellow basally, brown on distal 1/ 3 in male. Mesonotum brown (Fig. 2G). Katepisternum dark brown on upper part, whiteyellow on lower part (Fig. 2G). Scutellum black (Fig. 2G). Legs dark brown to black, yellow on basal part of fore femora and 2nd to 5th tarsomeres. Fore femur with 3 long setae on distal half of posteroventral surface. All abdominal tergites brown (Fig. 2G); all sternites yellowish brown. Male terminalia (Fig. 10): Epandrium with ca. 20 setae near posterior portion per side. Surstylus broad in lateral view. Cercus lacking pubescence. Tenth sternite with 1 pair of asymmetric projections sublaterally. Paramere with ca. 2 sensilla. Aedeagus with pubescence. Gonopods with 1 suberect lobe-like process bearing several pubescence. Measurements. BL = 2.80 mm in holotype (1♂ paratype: 2.75), ThL = 1.13 mm (1.26), WL = 2.00 mm (2.65), WW = 1.00 mm (1.21), arb = 6/5 (5/4), avd = 0.60 (0.84), adf = 1.43 (1.02), flw = 1.57 (1.90), FW/HW = 0.31 (0.40), ch/o = 0.13 (0.11), prorb = 1.07 (0.90), rcorb = 0.60 (0.62), vb = 0.25 (0.27), dc1 = damaged (damaged), presct1 = 0.50 (damaged), sct1 = 1.59 (damaged), sterno = damaged (0.87), orbito = 2.25 (1.91), dcp = 0.24 (0.25), sct1p = 1.08 (1.15), C = 2.00 (2.19), 4c = 1.13 (1.42), 4v = 1.57 (1.82), 5x = 1.33 (1.13), ac = 9.00 (7.47), M = 0.50 (0.62), C3F = 0.71 (0.58). Specimens examined. Holotype ♂ (SCAU, No. 124706), China: Dashuigou, Lüchun, Yunnan, 22°57'19''N, 101°57'4''E, altitude 1310m, 2.viii.2016, tussock, HW Chen. Paratype: China: 1♂ (SCAU, No. 124709), Mengma, Menglian, Yunnan, 22°12'59''N, 99°22'51''E, altitude 1070m, 15.iv.2018, tree trunk, HW Chen. Etymology. A combination of the Latin words: hirtus + foliaceus, referring to the aedeagus with several setae. Distribution. China (Yunnan).

Stegana (Steganina) spatulata sp. nov. (Figs 2E, 8) Diagnosis. This species resembles S. nigrithorax Strobl, 1898 in some characters of the colour patterns of body (Fig. 2E) and semicircular gonopods (Fig. 8E), but can be distinguished by the median flaps of hypandrium (Hu & Toda, 1994; fused gonopod in Bächli et al., 2004) concave, spoon-like (Fig. 8E, F); aedeagus with numerous, serrated, minute processes medially (Fig. 8C, D). However, the minimum interspecific genetic distance is 3.1% to S. pililobosa (Table 2). Description. Male: Frons brown. First flagellomere yellow on basal 1/4, brown on distal 3/4. Clypeus yellow, slightly brownish medially. Palpus yellow. Katepisternum entirely white. Scutellum brown (Fig. 2E). Femur of foreleg with 5 long erect setae near distal part of posteroventral surface. Abdominal tergites entirely dark brown medially, yellowish brown laterally (Fig. 2E). Male terminalia (Fig. 8): Epandrium protruded ventrally, with ca. 20 setae on posterior portion per side. Surstylus distally with numerous setulae on inner surface. Cercus lacking pubescence. Tenth sternite with 1 pair of processes submedially. Paramere lacking setula. Measurements. BL = 3.47 mm, ThL = 1.46 mm, WL = 2.90 mm, WW = 1.14 mm, arb = 6/5, avd = 0.77, adf = 1.31, flw = 1.82, FW/HW = 0.28, ch/o = 0.23, prorb = 0.30, rcorb = 0.79, vb = 0.66, dc1 = 0.33, presct1 = 0.35, sct1 = damaged, sterno = 0.88, orbito = 1.45, dcp = 0.30, sct1p = 1.59, C = 2.00, 4c = 1.14, 4v = 1.36, 5x = 1.41, ac = 8.42, M = 0.54, C3F = 0.63. Specimens examined. Holotype ♂ (SCAU, No. 110902), China: Guojiapuzi, Kuandian, Liaoning, 40°46'46''N, 124°45'07''E, altitude 342m, 26.vii.2018, tree trunk, NN Wang. Etymology. From the Latin word: spatulate, referring to the median flaps of hypandrium spoon-like. Distribution. China (Liaoning).

Stegana (Steganina) unguiculata sp. nov. (Figs 2H, 11) Diagnosis. This species resembles S. linnanensis Cheng, Gao & Chen, 2009 from southern China in the colour patterns of body (Fig. 2H) and male terminalia (Fig. 11), but can be distinguished from it by the aedeagus nearly bilobed, with 7 pair of slender processes and thick, long pubescence laterally (Fig. 11C); gonopods with 1 suberect lobe-like process bearing several pubescence (Fig. 11D). However, the minimum interspecific genetic distance is 3.3% to S. hirtifoliacea (Table 2). Description. Male and Female: Pedicel brown medially, black laterally; first flagellomere nearly black. Palpus yellow basally, brown on distal 1/ 3 in male, brown on distal 2/ 3 in female. Mesonotum dark brown medially, brown laterally (Fig. 2H). Katepisternum dark brown on upper part, white-yellow on lower part (Fig. 2H). Scutellum black (Fig. 2H). Legs dark brown to black, yellow on basal part of fore femora and 2nd to 5th tarsomeres. Fore femur with 3 long setae on distal half of posteroventral surface. All abdominal tergites brown (Fig. 2H); all sternites yellowish brown. Male terminalia (Fig. 11 A���E): Epandrium with approximately 22 setae near posterior portion per side. Surstylus broad in lateral view. Cercus lacking pubescence. Tenth sternite with 1 pair of asymmetric projection sublaterally. Plate between hypandrium and aedeagus with 4 sensilla. Aedeagus composed of 1 bilobed medial process with pubescence. Female terminalia (Fig. 11 F���H): Epiproct and cercus with setae, lacking pubescence. Hypoproct pubescent medially, with dense setae along lateral and posterior margins. Eighth sternite glabrous, with a pair of small setae near posterior margin. Measurements. BL = 2.83 mm in holotype (mm in five ♂ and five ♀ paratypes: 2.37���2.83 in ♂, 2.87���3.27 in ♀), ThL = 1.17 mm (1.20���1.30 in ♂, 1.20���1.47 in ♀), WL = 1.83 mm (1.93���2.17 in ♂, 2.00��� 2.23 in ♀), WW = 1.00 mm (1.00��� 1.10 in ♂, 1.07���1.17 in ♀), arb = 7/5 (7���9/5���6), avd = 0.90 (0.60���0.92), adf = 1.67 (1.11���1.83), flw = 2.17 (1.57���2.50), FW/HW = 0.46 (0.34���0.44), ch/o = 0.10 (0.09���0.16), prorb = 0.94 (1.00���1.27), rcorb = 0.59 (0.60���0.91), vb = 0.29 (0.18���0.31), dc1 = damaged (0.31���0.48), presctl = damaged (0.55���0.71), sct1 = 2.00 (1.40��� 1.90), sterno = 1.00 (0.85���1.25), orbito = 2.25 (1.67���2.50), dcp = 0.29 (0.17���0.36), sct1p = 1.30 (1.09���1.42), C = 1.89 (1.71���2.06), 4c = 1.20 (1.00���1.29), 4v = 1.60 (1.50���1.81), 5x = 1.50 (1.18���1.54), ac = 9.00 (7.00���10.05), M = 0.60 (0.43���0.67), C3F = 0.39 (0.50���0.74). Specimens examined. Holotype: ♂ (SCAU, No. 111477), China: Qimaba, L��chun, Yunnan, 22��49'8''N, 102��17'25''E, altitude 1280m, 3,4. viii.2016, tree trunk, HW Chen. Paratypes: China: 3♂, 7♀ (SCAU, Nos 111478 ��� 7), same data as holotype, HW Chen, L Gong, YQ Liu; 1♀ (SCAU, No. 111488), Dashuigou, L��hun, Yunnan, 22��57'19''N, 101��57'4''E, altitude 1310m, 2.viii.2016, tree trunk, HW Chen; 1♂, 1♀ (SCAU, Nos 111540, 41), Jinshan Forest Park, Menglian, Yunnan, 22��20'6''N, 99��34'34''E, altitude 1049m, 24.x.2017, tree trunks, HW Chen, L Gong; 1♂, 1♀ (KIZ, Nos 0 0 90663, 64), Moli Forest Park, Ruili, Yunnan, 24��7'5''N, 97��58'51''E, altitude 920m, 5.xi.2017, tree trunks, HW Chen, L Gong; 1♀ (SCAU, No. 124146), Beibeng, Motuo, Xizang, 29��19'N, 95��20'E, altitude 800m, 29.ix.2010, tree trunk, L Wang. Etymology. From the Latin word: unguiculus, referring to the aedeagus with claw-shape processes. Distribution. China (Yunnan, Xizang)., Published as part of Li, Bingxue, Zhang, Yuan & Chen, Hongwei, 2019, Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae), pp. 81-108 in Zootaxa 4576 (1) on pages 100-102, DOI: 10.11646/zootaxa.4576.1.4, http://zenodo.org/record/2624686, {"references":["Cheng, Y., Gao, J. J. & Chen, H. W. (2009) The Stegana ornatipes species group from the Oriental Region (Diptera, Drosophilidae). Zootaxa, 2216, 37 - 48."]}

Stegana (Steganina) pinguifoliacea sp. nov. (Figs 2D, 7) Diagnosis. This species resembles S. longifibula Takada, 1968 in some characters of the body colour patterns (Fig. 2D), but can be distinguished by the male terminalia, in this species, the surstylus without minute projections, except acute apically (Fig. 7A); aedeagus medially expanded in lateral view, strongly narrowed in ventrally (Fig. 7B, C); gonopods nearly vaulted in ventral view (Fig. 7D). The minimum interspecific genetic distance to S. longifibula is 4.3%, which is smaller than to other species (Table 2). Description. Male: Frons black. First flagellomere greish yellow. Clypeus and palpus yellow. Mesonotum brown to dark brown, with 1 thin, unconspicuous, yellow longitudinal stripe medially, yellowish brown laterally. Katepisternum entirely white. Scutellum black. Femur of foreleg with 4 long erect setae near distal part of posteroventral surface. Abdominal tergites nearly black. Male terminalia (Fig. 7): Epandrium slightly protruded ventrally, with ca. 15 setae on posterior portion per side. Surstylus with numerous setulae distally. Cercus lacking pubescence. Paramere with 2 setulae. Tenth sternite flat dorsolaterally. Measurements. BL = 3.01 mm in holotype, ThL = 1.47 mm, WL = 2.95 mm, WW = 1.26 mm, arb = 6/4, avd = 0.62, adf = 1.36, flw = 1.87, FW/HW = 0.41, ch/o = 0.22, prorb = 1.30, rcorb = 0.73, vb = 0.52, dc1 = damaged, presct1 = damaged, sct1 = damaged, sterno = damaged, orbito = 1.73, dcp = 0.35, sct1p = 1.68, C = 2.08, 4c = 1.02, 4v = 1.55, 5x = 1.26, ac = 9.25, M = 0.45, C3F = 0.65. Specimens examined. Holotype ♂ (SCAU, No. 110944), China: Pangquangou, Jiaocheng, Shanxi, 37��04'47''N, 111��30'20''E, altitude 2200m, 30.vii.2016, tree trunk, L Zhu. Etymology. A combination of the Latin words: pinguis and foliaceous, reference to the aedeagus medially expanded in lateral view., Published as part of Li, Bingxue, Zhang, Yuan & Chen, Hongwei, 2019, Nine new species of the subgenus Stegana (Steganina) from China, with DNA barcoding information (Diptera: Drosophilidae), pp. 81-108 in Zootaxa 4576 (1) on pages 94-95, DOI: 10.11646/zootaxa.4576.1.4, http://zenodo.org/record/2624686, {"references":["Takada, H. (1968) Drosophila survey of Hokkaido, XXVI. Descriptions of three new species of Drosophilidae from Japan. Journal of the Faculty of General Education, Sapporo University, 1, 119 - 129."]}

Stegana (Steganina) biflava sp. nov. (Figs 2I, 12) Diagnosis. This species resembles S. flavichypeata Chen & Chen in Lu et al. 2011a in the yellow clypeus and palpus, and colour patterns of body (Fig. 2I), but can be distinguished by having the face black on upper 2/3, yellow on lower 1/3; aedeagus with triangular opening in ventral view (Fig. 12B). In S. flavichypeata: Face black, with yellow band medially; aedeagus with rhombic opening in ventral view (fig. 24 in Lu et al. 2011a). The minimum interspecific genetic distance to P. flavichypeata is 7.1%, which is smaller than to other species in this group (Table 2). Description. Male and female. Frons black, with orange yellow stripe medially and band submedially. First flagellomere medially grayish yellow on basal half, black on distal half. Mesonotum black, with thin, yellow longitudinal stripes sublaterally (Fig. 2I). Katepisternum entirely white (Fig. 2I). Scutellum black, yellow at tip (Fig. 2I). Veins R 4+5 and M 1 lacking setae on basal section of dorsal surface. Legs white on basal half of all femora, brown on distal 1/3 of foreleg femur, black on distal half of femora and basal half of tibiae of mid and hindleg, yellow on the rest. Fore femur distally with 3 or 4 setae on posteroventral surface. Abdominal tergites 1st to 5th medially brown, laterally yellowish brown, nearly black on 6th tergite in male; all tergites black in female. Sternites yellow to brownish yellow. Male terminalia (Fig. 12 A–F): Hypandrium anteromedially slightly elongated, about 1.2 times as long as width of epandrium, with a few paramedian setae posteroventrally. Gonopods slightly triangular. Aedeagus slightly expanded distally in ventral view, about 3/5 times as long as aedeagal apodeme. Female terminalia (Fig. 12 G–I): Epiproct, cercus and hypoproct with dense setae and pubescence. Eighth sternite glabrous on anterior 2/3 region, with dense setae and pubescence on posterior 1/3 region. Measurements. BL = 3.60 mm in holotype (mm in 2♂ and 3♀ paratypes: 3.67–4.13 in ♂, 4.00– 4.13 in ♀), ThL = 1.73 mm (1.53–2.33 in ♂, 1.87–2.07 in ♀), WL = 3.00 mm (2.80–3.47 in ♂, 3.00– 3.87 in ♀), WW = 1.53 mm (1.40–1.87 in ♂, 1.33–1.60 in ♀), arb = 4/2 (7–8/3–5), avd = 0.50 (0.33–0.89), adf = 2.40 (1.50–1.88), flw = 1.60 (1.40–2.40), FW/HW = 0.51 (0.40–0.50), ch/o = 0.17 (0.16–0.33), prorb = 0.93 (0.82–1.42), rcorb = 0.71 (0.70–1.00), orbito = 1.80 (2.20–3.00), vb = 0.75 (0.18–0.80), sctl = 1.60 (1.25–1.60), sterno = 0.67 (0.50–0.89), sctlp = 1.45 (1.27–1.83), C = 1.92 (1.25–2.00), 4c = 1.50 (1.08–1.40), 4v = 2.38 (1.10–2.00), 5x = 1.50 (1.25– 2.00), ac = 6.00 (6.50–12.00), M = 0.75 (0.50–0.80), C3F = 0.67 (0.67–0.79). Specimens examined. Holotype ♂ (SCAU, No. 111533), China: Mulun, Huanjiang, Guangxi, 25°07'N, 108°02'E, altitude 480m, 16.ix.2015, tree trunk, YQ Liu. Paratypes: China: 2♀ (SCAU, Nos 111534, 35), same data as holotype; 1♂ (KIZ, No. 0090669), Qimaba, Lühun, Yunnan, 22°49'8''N, 102°17'25''E, altitude 1280m, 3.viii.2016, tree trunk, HW Chen; 1♂ (KIZ, No. 0090670), Yakou, Huanglianshan, Lühun, Yunnan, 22°53'40''N, 102°18'17''E, altitude 1900m, 31.x.2016, tree trunk, HW Chen; 2♂, 1♀ (SCAU, Nos 111536 –38), Wengang, Libo, Guizhou, 25°13'N, 107°56'E, altitude 754m, 19.ix.2015, tree trunks, YQ Liu, Y Zhang, L Zhu. Etymology. A combination of the Latin words: bi- + flavus, referring to the yellow clypeus and palpus. Distribution. China (Yunnan, Guizhou).

10. Conocephalus (Anisoptera) nanlingensis Li, Xin & Shi sp. nov. (Figs. 6, 7 C���D) Description. Male. Body medium-sized for genus. Fastigium verticis slightly narrow, apex blunt, width about 1/2 of scape; in frontal view, lateral margins paralleled, ventral margin connected with frontal verticis. Eyes subsphaeroidal, protruding forward and outward. Anterior margin of pronotum straight, posterior margin rounded; lateral lobe nearly triangular, higher than deep, anterior margin arc-shaped, posterior margin nearly straight, oblique; the area corresponding to thoracic foramen semitransparent, small, slightly tumid; humeral sinus absent. Prosternum with a pair of spines. Procoxae with 1 spine; profemora unarmed on ventral surface, apices of genicular lobes with 1 spine on inner margin and outer margin unarmed; protibiae with 6 spines on inner and outer margins of ventral surface separately; tibial tympana slit-like. Mesofemora unarmed on inner margin of ventral surface, outer margin of ventral surface with 0���2 spines; mesotibiae with 6 spines on inner and outer margins of ventral surface separately. Postfemora with 0���1 spine on inner margin and 6���8 spines on outer margin of ventral surface, apices of inner and outer genicular lobes with 2 spines separately, dorsal spine long than ventral spine; posttibiae with 22���30 spines on inner and outer margins of dorsal surface separately, with 2 pairs of ventral apical spurs, without dorsal apical spurs. Tegmina not reaching apex of abdomen, apices slightly rounded. Stridulatory area of left tegmen trapezoid, Cu2 vein robust, cylindrical. Hind wings slightly longer than tegmina. 10 th abdominal tergite elongated, basal half broad, apical half narrowing, near apex split, lateral lobes rectangular, curved ventrad. Cerci long, roughly cylindrical, apical areas conical, apices blunt; with two inner teeth near the middle, and bases of two teeth far apart; the tooth near base short, slightly compressed, apex acute, directing inward; the other tooth near the middle conical, robust, apical half slightly compressed, hook-like, apex slightly acute, curved ventrad. Subgenital plate rectangular, the midline of basal half with a shallow longitudinal groove, posterior margin with a V-shaped notch. Styli cylindrical, apices rounded, inserted on apices of posterior margin of the lobes of subgenital plate. Coloration. Body yellowish green (green when alive). Dorsal surface of head with a broad longitudinal reddish brown stripe. Disc of pronotum with a broad reddish brown stripe, outer margins with a light stripe separately, which blue when alive. Apices of postfemora, spines of posttibiae on dorsal surface brown. Middle area of dorsal surface of 10 th abdominal tergite blackish brown. Dorsal surface of apical half of cercus and two inner teeth brown. Costal and subcostal areas of tegmen light, the other areas brown. Female. Appearance is similar to male. Posttibiae with 1 apical spur on outer margin of dorsal surface and 2 pairs of ventral apical spurs. Tegmina reaching end of abdomen. 10 th abdominal tergite short, the middle of posterior margin split, the lobes curved ventrad. Cerci long, conical. Ovipositor rather long, narrow, moderately curved dorsad, basal area slightly narrow; dorsal and ventral margins smooth, dorsal valvulae longer than ventral ones, apices of dorsal and ventral valvulae acute. Subgenital plate broad, posterior margin rounded. Ovipositor light brown. Material examined. Holotype: ♂, Mangshan, Yizhang, Hunan, 23 August, 2018, coll. Yanqing Li and Yarui Xin. Paratype: 1♀, Xingzi, Lianzhou, Guangdong, 18 August, 2018, coll. Yanqing Li and Yarui Xin. Measurements (mm). Body: ♂ 17.0, ♀ 18.5; pronotum: ♂ 3.8, ♀ 4.0; tegmina: ♂ 11.5, ♀ 13.0; postfemora: ♂ 11.5, ♀ 12.5; ovipositor: 14.0. Distribution. China (Hunan, Guangdong). Remarks. The new species is similar to Conocephalus (Anisoptera) guangdongensis Shi & Liang, 1997 and Conocephalus (Conocephalus) bidentatus Shi & Zheng, 1994, but it is distinguished from the former in: the tooth near middle of male cercus obviously long, and bases of two teeth far apart, without overlapping arrangement; and from the latter in: male cerci long, roughly cylindrical, apical areas conical, apices blunt; bases of two teeth on cercus far apart, inserted near the middle; ovipositor slightly long and narrow, moderately curved dorsad. Discussion. Males of the species have comparatively short tegmina and hind wings when alive, which do not reach apices of abdomen, but tegmina and hind wings are near the abdominal posterior margin in the dry specimens because abdomen shrinks a bit. Etymology. The name of new species is derived from the type locality, Nanling., Published as part of Li, Yan-Qing, Zhang, Tao, Xin, Ya-Rui & Shi, Fu-Ming, 2019, The genus Conocephalus (Orthoptera: Tettigoniidae: Conocephalinae) from Guangdong and Nanling region, China with description of two new species, pp. 590-600 in Zootaxa 4565 (4) on pages 597-599, DOI: 10.11646/zootaxa.4565.4.12, http://zenodo.org/record/2591291, {"references":["Shi, F. M. & Liang, G. Q. (1997) Descriptions of two new species of the genus Conocephalus Thunberg (Orthoptera: Conocephalidae). Entomologia Sinica, 4 (3), 211 - 214.","Shi, F. M. & Zheng, Z. M. (1994) Two new species of katydids from Sichuan (Orthoptera: Tettigonioidea). Journal of Shanxi Teacher's University, 8 (1), 44 - 46."]}

Trichromothrips antidesmae sp.n. (Figs 10, 27���30) Female macroptera. Body and legs yellow(Fig. 10), head and pronotum with dark lateral margin(Fig. 27); antennal segments I���II brown, III���V yellow with the apex brown, VI���VII brown, VIII pale brown(Fig. 30); fore wing brown with apex pale, clavus brown(Fig. 29). Head (Fig. 27) without constriction behind eyes; ocellar setae III situated at tangent of hind ocelli posterior margin; five pairs of postocular setae present, I longest, III minute. Antennae (Fig. 30) segments II���VI with microtrichia, VIII slightly longer than VII. Pronotum (Fig. 27) smooth with some transverse striae on lateral anterior margin and near posterior margin; two pairs of posteroangular setae and two pairs of posteromarginal setae present; prosternal ferna divided medially. Mesonotum (Fig. 27) anterior smooth with some transverse striate sculpture medially, CPS absent anteromedially; median pair of setae near posterior margin. Metascutum smooth medially but sculptured longitudinally at each side; CPS absent; median pair of setae close to anterior margin (Fig. 27). Metaepimeron with two setae. Mesosternum with sternopleural sutures; only mesothoracic endofurca present. Fore wing first vein usually have 6 basal setae and 2 distal setae, second vein with 14 setae; clavus usually with 4 marginal setae and 1 discal seta (Fig. 29). Abdominal tergites with transverse striae laterally, tergites II���VII with three setae (S3 to S5 setae) arranged in a straight line along lateral margin; tergite IX without CPS; sternites without discal setae, II with two pairs of posteromarginal setae, III���VII each with 3 pairs of posteromarginal setae, all marginal setae inserted on posterior margin. Measurements (holotype female in microns). Distended body length 1046. Head length 86, width across cheeks 128; width across eyes129; ocellar setae III length 20. Pronotum median length 89, width 161; posteroangular setae length, inner 52, outer 44; posteromarginal setae I length 24. Metascutum median setae length 18. Fore wings 673. Antennal segments I to VIII length as follows: 19, 29, 38, 55, 40, 49, 7, 14. Male macroptera. General structure and colour similar to female. Abdominal tergite IX with a pair of long posteromarginal drepanae; sternites without pore plates. Measurements (paratype male in microns). Distended body length 1034. Head length 130, width across cheeks 125; width across eyes123; ocellar setae III length 31. Pronotum median length 96, width 150; posteroangular setae length, inner 47, outer 43; posteromarginal setae I length 22. Metascutum median setae length 20. Fore wings 600. Antennal segments I to VIII length as follows: 20, 28, 46, 52, 37, 52, 9, 20. Specimens studied. Holotype female, CHINA, Yunnan, Menglun Town, Xishuangbanna Tropical Botanic Garden, from Antidesma fordii, 24.x.2017 (Hui Liu), in collection of Yunnan Agricultural University, Kunming. Paratype: 1male, same data as holotype. Etymology. In reference to the host plant of this species. Comments. The head and pronotum of this species with dark lateral margin is similar to bilongilineatus, priesneri and trachelospermi, but the meso-, metanotum and abdominal tergite IX without CPS, marginal setae situated on posterior margin on sternite VII are different from bilongilineatus. The two pairs of posteromarginal setae on sternite II, and the absence in males of sternal pore plates also distinguishes this species from the Indian species priesneri and Chinese species trachelospermi (Bhatti 2000, Zhang &Tong 1996). In addition, the presence of two microtrichial rows on the dorsal surface of antennal segment II is unusual among Trichromothrips species., Published as part of Li, Yajin, Li, Zhengyue & Zhang, Hongrui, 2019, Trichromothrips genus-group (Thysanoptera, Thripidae) from China, with descriptions of three new species and ten new records, pp. 79-92 in Zootaxa 4544 (1) on pages 81-82, DOI: 10.11646/zootaxa.4544.1.3, http://zenodo.org/record/2618149, {"references":["Bhatti, J. S. (2000) Revision of Trichromothrips and related genera (Terebrantia: Thripidae), Oriental Insects, 34 (1), 1 - 65. https: // doi. org / 10.1080 / 00305316.2000.10417252","Zhang, W. Q. & Tong, X. L. (1996) A new species and some new records of Thripinae (Thysanoptera: Thripidae) from China. Entomotaxonomia, 18 (4), 253 - 256."]}

Trichromoithrips moundi sp.n. (Figs 1,2, 13–21) Female macroptera. Body bicolored (Fig. 1); head, metanotum, abdominal segments II–III &VII brown; mesonotum and abdominal segment I light brown; pronotum and abdominal tergites IV–VI and VIII–X yellow; antennae brown, III pale, the apex of IV–VI also VII–VIII light brown (Fig. 20); fore wing brown with sub-basal, median and apex white, clavus brown (Fig. 21); all legs pale. Head longer than wide, slightly constricted just behind compound eyes; ocellar setae I absent, ocellar setae II sub-equal to setae III or sometimes slightly shorter than setae III, setae III situated at tangent of posterior margin of hind ocelli (Fig. 13); four pairs of postocular setae present. Compound eyes without pigmented ommatidia. Mouthcone short and rounded at apex with 3-segmented maxillary palpi. Antennae (Fig. 20) 8-segmented, segment I with two dorsal apical setae, III and IV each with forked sense cone, III–VI with microtrichia, VIII longer than VII. Pronotum (Fig. 13) distinctly sculptured with transverse striae, two small circular striae on posterior third; two pairs of posteroangular setae and two pairs of posteromarginal setae developed; Prosternal ferna not divided at middle. Mesonotum (Fig. 17) smooth at anterior, with transverse striate sculpture on posterior 2/3; median pair of setae near posterior margin. Metascutum (Fig. 17) transversely reticulate medially; campaniform sensilla absent; median pair of setae behind anterior margin. Both meso- and metasternal furca with spinula, slightly weak on metasternum; sternopleural sutures present. Fore wing first and second veins each with complete row of setae; clavus usually with 4–5marginal setae and 1–2discal setae; posteromarginal cilia wavy (Fig. 21). Tarsi 2- segmented. Abdominal tergites smooth, with a few striae on anterior half, posteromarginal craspeda and ctenidia absent; lateral area of tergites II–VIII with about 4–6 setae in a regular or irregular transverse row; tergites II–VII with three setae (S3 to S5 setae) arranged in a straight line along lateral margin (Figs 16, 18); tergite VIII without posteromarginal comb; tergite IX with posterior pair of CPS and SB2 setae absent; tergite X without median split; sternites without posteromarginal craspeda, sternite VII with two discal setae laterally in front of setae S2 and S3 (Fig. 19); sternites II–VII each with 3 pairs of posteromarginal setae in front of post margin(Fig. 19); laterotergites without discal setae. Measurements (holotype female in microns). Distended body length 1697. Head length 110, width across cheeks 149; width across eyes 127; ocellar setae III length 21. Pronotum median length 149, width 204; posteroangular setae length, inner 45, outer 43; posteromarginal setae I length 40. Metascutum median setae length 23. Fore wings 1054. Antennal segments I to VIII length as follows: 20, 34, 56, 51, 46, 64, 14, 20. Male macroptera. General structure and colour similar to female (Fig. 2). Abdominal tergite IX with a pair of long posteromarginal drepanae (Fig. 15); sternites III–VIII each with some scattered small pore plates, all posteromarginal setae situated on post margin, no discal setae present (Fig. 14). Measurements (paratype male in microns). Distended body length 1394. Head length 88, width across cheeks 134; width across eyes 105; ocellar setae III length 29. Pronotum median length 118, width, 175; posteroangular setae length, inner 42, outer 42; posteromarginal setae I length 40. Metascutum median setae length 15. Fore wings 881. Antennal segments I to VIII length as follows: 21, 28, 47, 44, 40, 56, 15, 18. Specimens studied. Holotype female, CHINA, Guizhou, Tongren City, Fanjingshan National Nature Reserve, from bamboo, 11.viii.2017 (YajinLi), in collection of Yunnan Agricultural University, Kunming; Paratypes: 1 female, 1 male, same data as holotype. 1 male and 1 female deposited in Australian National Insect Collection, Canberra. Etymology. The species is in honor of Dr. Laurence Mound for his assistance and his excellent work on taxonomy of thrips. Comments. This new species differs from the other Trichromothrips species in having the fore wing with two complete rows of setae, and abdominal tergites II–VIII with about 4–6 setae in a regular or irregular transverse row. Otherwise, the female of T. moundi with two discal setae laterally on sternite VII can also be distinguished from caespitis section.

Trichromothrips guizhouensis sp.n. (Figs 4, 22–26) Female macroptera. Body bicolored (Fig. 4); head uniformly dark brown (Fig. 22); antennal segments I–II dark brown, III–V yellow with the apical third brown, VI–VIII pale brown, VIII with apex slightly paler(Fig. 26); pronotum with dark brown lateral stripes, median triangular pale area not reaching anterior margin(Fig. 22); fore wing including clavus brown with a small spot near the base(Fig. 25); all legs yellowish white; abdominal tergite II with a small pale brown patch in middle, III–VI yellowish white each with an inverted trapezoidal-shaped area in the middle, VII–IX dark brown, X yellowish brown with apex pale(Figs 4, 25); body setae brown. Head (Fig. 22) longer than wide, with a fairly long neck-like constriction and transverse striae behind compound eyes; ocellar setae III situated at tangent of posterior margin of hind ocelli; six pairs of postocular setae present, V far from compound eyes. Antennal segments III–VI with microtrichia (Fig. 26), VIII longer than VII. Pronotum (Fig. 22) wider than long, smooth with about 16–18 discal setae, inner posteroangular setae longer than outer, two pairs of posteromarginal setae present. Mesonotum (Fig. 24) anterior smooth with some transverse striate sculpture near posterior margin, CPS absent anteromedially; median pair of setae situated in front of submedian pairs. Metascutum (Fig. 24) smooth medially but sculptured longitudinally at each side, CPS absent; median pair of setae close to anterior margin. Prosternal ferna undivided. Metaepimeron without setae. Mesosternum with sternopleural sutures; only mesothoracic endofurca present. Fore wing first vein usually with 7 basal and 2 discal setae, second vein with 17 setae; clavus usually with 3–5 veinal and 1 discal setae. Abdominal tergites with transverse striae laterally and a few striae at anterior margin; tergites II–VII (Fig. 23) with three setae (S3 to S5 setae) arranged in a straight line along lateral margin; S4 minute on tergites V–VIII; tergite IX without CPS; sternites without discal setae, II–VII each with 3 pairs of posteromarginal setae, VII with all three pairs of setae inserted ahead of posterior margin. Measurements (holotype female in microns). Distended body length 1836. Head length 149, width across cheeks 124; width across eyes 162; ocellar setae III length 38. Pronotum median length 130, width 210; posteroangular setae length, inner 96, outer 61; posteromarginal setae I length 25. Metascutum median setae length 34. Fore wings 1132. Antennal segments I to VIII length as follows: 20, 43, 57, 75, 70, 88, 20, 27. Male. Unknown. Specimens studied. Holotype female, CHINA, Guizhou, Tongren City, Fanjingshan National Nature Reserve, from bamboo, 11.viii.2017 (Yajin Li), in collection of Yunnan Agricultural University, Kunming. Paratypes: 6 females, same data as holotype. Etymology. The species is named for the collection place. Comments. This new species runs to bellus in the key provided by Bhatti (2000), but is distinguished from bellus by the presence of the mesothoracic endofurca, S4 minute on tergites V–VIII and abdominal tergites VII–IX dark brown. It is closely related to dorsalis (Masumoto & Okajima 2011), but can be distinguished by head with a long neck-like constriction, abdominal tergites III–VI each with an inverted trapezoidal brown patch in the middle and tergites VII–IX dark brown (in dorsalis head slightly concave behind compound eyes, abdominal tergites I–VII yellowish white with III–VII each with a transverse diamond-shaped dark area in middle, VIII–IX dark brown). According to Masumoto and Okajima (2011) this species may belong to bellus -section together with alis, arorai, dorsalis, elegans and taiwanus.

Araliacothrips daweishanensis sp.n. (Figs 1–15) Female. With the characters in the generic diagnosis above. Body entirely blackish brown (Fig. 1); legs brown except for extreme apex of tibiae and tarsi yellow; antennal segment I yellowish brown, II brown, III–IV and basal half of V yellow; apical half of V and VI–VII entirely brown, VIII light brown (Fig. 7); fore wing brown, white sub-basally and with diffuse paler area sub-apically, clavus brown with apex pale (Fig. 6). Body setae clear in color. Head with ocellar hump slightly elevated (Fig. 8); reticles with a few scattered dot-like thickening area medially on posterior margin; anteocellar, interocellar and four pairs of postocellar setae well-developed; mouth cone broadly rounded. Pronotum with slightly rounded reticles, with semicircular hind margin with setae about equal in length to postocellar setae (Fig. 9). Fore wing first vein with 7–8 basal setae and 2 distal setae, second vein with 6–8 widelyspaced setae; anterior fringe cilia longer than costal setae; clavus with 4–5 veinal setae but no discal setae (Fig. 6). Abdominal tergite I with median pair of minute setae near the heavy antecostal line (Fig. 10); tergite II with antecostal line thick and completely covered with polygonal reticulations, but smooth posteriorly (Fig. 11); III–VII with arched sculpture (Fig. 12), sub-median setae becoming stronger gradually on V–VIII; sternites II–VII reticulate medially on anterior half. Measurements (holotype female in microns): Body length 1567. Head, length 130; width 166. Pronotum, length 140; width 226. Fore wing length 1038. Tergite IX setae length, S1 115; S2 139. Antennal segments III–VIII length 75, 63, 45, 29, 10, 40. Male. Similar to female in colour, sculpture and chaetotaxy but a little smaller (Fig. 2). Abdominal tergite IX with three pairs of stout thorn-like setae, anterior pair slightly stouter, each posterior pair slightly more slender and lateral to preceding one (Fig. 3). Sternites IV–VII each with a broad transverse pore plate near anterior margin (Fig. 4). Measurements (paratype male in microns): Body length 1428. Head, length 120; width 155. Pronotum, length 127; width 187. Fore wing length 891. Antennal segments III–VIII length 72, 56, 37, 23, 9, 38. Specimens examined. Holotype: female, CHINA, Yunnan Province, Pingbian County, Dawei Mountain Nature Reserve, from Eleutherococcus nodiflorus leaves [Araliaceae], 3.viii.2017 (Kong Bo & Wu Can), in collection of Yunnan Agricultural University, Kunming. Paratypes: 3 females, collected with holotype; same locality, 1 female from fern, 2.viii.2017; 1 male from Malachium aquaticum, 13.v.2017; 3 females, 3 males from E. nodiflorus leaves, 3.x. 2017 (with about 10 specimens preserved in alcohol); 1 male and 1 female deposited in Australian National Insect Collection, Canberra. Etymology. The genus is named after the host plant, and the specific epithet is taken from the collecting place. Comments. All specimens were collected from the same location. Initially, one male was collected from fern, and then one female was collected from Malachium aquaticum, but subsequently this thrips species was collected twice from the medicinal plant Eleutherococcus nodiflorus, although no larvae were collected.

Araliacothrips gen.n. Macropterous, body slender, dark brown, with strongly reticulate surface. Antennae 8-segmented, III and IV with long apical neck (Figs. 5, 7), III and IV each with sense cone long and forked, IV with one simple sense cone laterally (Fig. 5); segment VI constricted at base, VIII much longer than VII. Head completely reticulate, sculpture on dorsal surface not raised; slightly longer than wide, cheeks parallel, constricted behind eyes and at basal neck; compound eyes with 6 weakly pigmented facets; occipital region without apodeme or collar; ocellar setae pair I present, pair III on anterolateral margins of ocellar triangle; maxillary palps 2-segmented. Pronotum reticulate, with convex posterior margin, setae small and equal in length (Fig. 9). Mesoscutum with short posteromedian cleft; metascutum reticulate without median strong triangle, median pair of setae far from anterior margin, campaniform sensilla present (Fig. 13); metathoracic endofurca simple and not sinuate, with short Y-shaped arms (Fig. 15). Femora and tibiae reticulate, tarsi 1-segmented. Fore wing with apex acute and posterior fringe cilia wavy, first vein with setal row widely interrupted, second vein with scattered setae (Fig. 6). Abdominal tergites reticulate, tergal median setae small; posterior margin of tergite I with dentate craspedum (Fig. 10); lateral thirds of tergites with reticles; tergite II not constricted and without strong microtrichia (Fig. 11); tergites III–VIII with arched antecostal ridge, median tergal setae small and well separated from each other, sub-median setae slightly curved or sigmoid (Figs 12, 14); both tergites and sternites with broad craspedum on segments II–VIII; tergite VIII with posteromarginal comb present laterally (Fig. 14); tergite X symmetric, with median split complete; setae on tergite IX longer than tergite X (Fig. 14); sternites with setae small. Male sternites IV–VII with transverse pore plate anterior to antecostal ridge (Fig. 4); tergite IX with paired thorn-like setae (Fig. 3). Type species Araliacothrips daweishanensis sp.n.

Indonemoura furcoloba Li & Yang, sp. nov. (Figs. 1, 3) Diagnosis. Laterally, the epiproct is nearly parallel but sinuous for most its length with a rounded apex, distinctly tapering apically in lateral view; the median lobe of paraproct is divided into two branches at basal fifth, inner branch longer and stouter with a blunt tip, outer branch slender and spine-like with acute apical tip. The female is characterized by sternum 7 forming a large trapezoid shaped pregenital plate barely covering sternum 8. Adult habitus (Figs. 3 a, 3b). Holotype male: Forewing length ca. 4.7 mm, hindwing length ca. 3.9 mm. Female forewing length ca. 6.2 mm, hindwing length ca. 4.9 mm, apex of both wings of one female slightly damaged. General color brown, teneral females yellow brown. Head and antennae dark brown; compound eyes dark; mouthparts brownish. Thorax including pronotum dark brown; pronotum (Fig. 3 b) nearly quadrate with obtuse corners, lateral margins light brown, medial portion with weak markings; legs uniformly dark brown. Wings subhyaline with dark veins. Abdominal segments brownish but terminalia darker with hairs pale brown. Male terminalia. Terga 7–8 with sclerotized anterior margins. Tergum 9 (Fig. 1 a) weakly sclerotized, anterior margin with shallow indentation. Sternum 9 with convex anterior margin except a rounded median notch before a claviform vesicle; vesicle with stalk-like narrow base with slight medial constriction, length> 4X width (Fig. 1 b); hypoproct broad and subquadrate at basal half, then gradually tapering toward tip, apical fifth tubular and the extreme apex nipple-like. Tergum 10 sclerotized with distinctly sclerotized lateral margins and a pair of bands obliquely across each side of the tergum, a deep median concavity present beneath epiproct, surrounded lateral areas slightly elevated and bearing several spinules along anterolateral margins of the concavity. Cercus membranous with many clothing hairs, generally cylindrical but gently narrowing toward tip, length ca. 2.5X width. Epiproct (Figs. 1 a, 1c–1d) slender and recurved, with sinuous lateral margins; basal half roughly parallelsided, apical half slightly enlarged with circular apex; in dorsal aspect each side with a thin lateral sclerite, in lateral view, extending along the lowest face; ventral sclerite with darkly sclerotized gourd-like apex, in dorsal aspect, base broad and gradually narrowing, medial portion narrowest until the swollen gourd-like apex. In lateral view, medial portion with weakly developed ventral ridge bearing a row of ca. 10 tiny spines before fusing with dorsal sclerite. Paraproct (Fig. 1 e) divided into 3 lobes: inner lobe slightly sclerotized, triangular, adhering to median lobe-typical of the genus, nearly as long as outer lobe; median lobe mostly sclerotized, half width of base of paraproct, divided into 2 branches at basal fifth, inner branch mostly parallel-sided, nearly straight but apex slightly out-curved; in ventral view with blunt tip in dorsal view blade-shaped, outer branch slender than inner one, darkly sclerotized and out-curved contacting apex of outer lobe, generally spine-like with sharp tip; outer lobe strongly sclerotized, stripe shaped and adhering to outer margin of the median lobe. Female (Figs. 3 c–3e). Sternum 7 mostly membranous, mid-posterior margin greatly produced forming a large trapezoid pregenital plate barely covering sternum 8, as wide as half of width of sternum 7. Sternum 8 moderately sclerotized but subgenital plate not fully sclerotized. Sternum 9 sclerotized with anterior margin slightly extended backward, the mid-anterior portion touching posterior margin of pregenital plate. Sternum 10 and paraproct sclerotized. Pregenital plate with posterior margin in dorsal view nearly truncate with gentle indistinct median notch; in lateral view, slightly downcurved, an inner crescent sclerite underlying the extreme anterior portion of pregenital plate, followed by a pair of medially touching archaic structures together with two triangular brown sclerites beneath anterolateral margins of M-shaped pregenital plate. Type Material. Holotype: male (HIST), China: Guangxi Zhuang Autonomous Region, Jinxiu County, Dayaoshan National Nature Reserve, Shengtang Mount., 1170 m, 2008. IV.28, leg. Gang Yao. Paratypes: 2 females (HIST & CAUC), same data as holotype. Etymology. The specific name refers to the bifurcate median lobe of paraproct. Distribution. China (Guangxi Zhuang Autonomous Region). Remarks. The new species seems to be most similar to I. bilobata Sivec and Stark, 2010 recently described from Thailand in having a comparable epiproct in dorsal view and similar paraprocts, especially the shape of outer lobe in caudal aspect. However, the new species may be separated from I. bilobata by its ventral sclerite of epiproct medially enlarged, and by the median lobe divided at basal fifth with two elongate lobes. In I. bilobata, the ventral sclerite of epiproct lacks the median ridge and bears a small subapical bulging in lateral view (figs. 12 & 14 in Sivec & Stark 2010); the median lobe is bilobed at apex with two short lobes in comparison with I. furcoloba n. sp. (comparing fig. 15 in Sivec & Stark 2010 and Fig. 1 e). The female of I. bilobata is still unknown. The new species also shares a similar bifurcate median paraproctal lobe with I. trichotoma Li & Yang, 2008b from Yunnan Province, China but it is easily separated from that species by the spineless hypoproctal apex and the stouter outer paraproctal lobe with gently curved apex (comparing figs. 7 & 10 in Li & Yang 2008b and Figs. 1 b, 1e).

Indonemoura nigrihamita concava Li & Yao, ssp. nov. (Figs. 2, 4) Diagnosis. Tergum 9 typical and gradually constricted medially. Epiproct bears rounded but incised apex in dorsal view; ventral sclerite extended apically forward dorsal sclerite forming a short finger-like flagellum. Outer lobe of paraproct forms stout strip around basal margin of cercus at its basal and median portions, with membranous and triangular apical portion. Adult habitus (Figs. 4 a, 4b). Males forewing length 6.1���6.2 mm, hindwing length 5.0��� 5.1 mm, females forewing length 8.0��� 8.2 mm, hindwing length 7.0��� 7.2 mm. Head dark brown to black and antennae a little lighter; mouthparts dark brown, compound eyes black. Thorax including pronotum dark brown to black, legs mostly brown, basal portion of femora and tibiae lighter, distal portion of femora darker, hindleg femora with distinct yellow band, band and color pattern on foreleg and midleg less distinct; wings subhyaline with darker veins; Abdominal segments mostly brown except terminalia darker; hairs on abdomen mostly pale brown. Male Terminalia (Figs. 2 a���2f). Tergum 9 weakly sclerotized, gradually constricted medially, with two irregular rows of tiny spines but lacking protruding lobes along mid-posterior margin (Fig. 2 a). Sternum 9 with claviform vesicle, length ca. 4X width, base slightly narrower and dorsolateral aspect sclerotized (Figs. 2 b, 2c), hypoproct broad and parallel-sided at base, gradually tapering toward tubular tip, covered by dense hairs, overlapping inner lobe of paraproct. Tergum 10 sclerotized, with narrow longitudinal concavity beneath epiproct. Cercus membranous with distinct hairs, length ca. 3X width. Epiproct (Figs. 2 b���2d) elongate and recurved; basal half nearly parallel-sided, apical half slightly enlarged, apex circular with triangular incision, in dorsal aspect each side with a thin lateral sclerite extended from entire basal sclerite; ventral sclerite sclerotized but medial portion heavier so, broad at base and distinctly narrower toward apex after inserting in dorsal sclerite, extended apically forward dorsal sclerite forming a short finger-like flagellum; in lateral aspect with weakly developed ventral ridge but bearing a row of tiny spines, mostly forward of a median constriction. Paraproct (Fig. 2 f) divided into 3 lobes: triangular inner lobe sclerotized, mostly hidden by hypoproct; median lobe forming a sclerotized spine-like structure with hook-like apex; outer lobe forming sclerotized strip around basal margin of cercus at its basal and median portions, with membranous and triangular apical portion. Female (Figs. 2 g, 4c���4e). Sternum 7 membranous with sclerotized pregenital plate near mid-posterior margin, the plate forming an elevated conical hump followed by a small sclerite until the posterior margin of the segment. Sternum 8 with large sclerotized subgenital plate covering the entire length, the plate generally trapezoidal but with distinct anterior and lateral incisions; the anterior incision deep and bowl-shaped, nearly semicircular and the lateral one shallow and crescent-shaped; the posterolateral corners slightly upraised, connecting to the triangular paragenital plate. Sternum 9 sclerotized with anterior fourth forming a trapezoidal extension, slightly overlapping posterior margin of preceding segment, with a wide shallow mid-anterior incision. Sternum 10 and paraproct typical. Pregenital and subgenital plates covered with dense short setae (Fig. 4 c). Type material. Holotype male (HIST), 1 paratype male (CAUC), 9 females (5 in HIST, 4 in CAUC), China: Sichuan Province, Aba Zang and Qiang Autonomous Region, Songpan County, Huanglong Scenic Area, Huanglong Temple to Wucai Lake, 3550 m, E 103.83140, N 32.72666, 2011. VIII.2, W.H. Li. Etymology. The specific name refers to the concave posterior margin of tergum 9 of the male. Distribution. China (Sichuan Province, Songpan County). Remarks. The above two males are similar to the holotype male of I. nigrihamita nigrihamita Li & Yang, 2008c from Kangding, western Sichuan Province in most aspects. The two populations are separated> 300 km and we did not collect the new subspecies from nearby Jiuzhaigou County. The major differences are in the shape of tergum 9 and the epiproct apex. The posterior margin of the new subspecies is unmodified and concave but that of I. nigrihamita nigrihamita bears a pair of paramedial protruding lobes at posterior margin; the apex of epiproct has a triangular incision but epiproct apex of I. nigrihamita nigrihamita is entire. Additionally, the median hook of median paraproctal lobe and the apical bulb of the outer paraproctal lobe are directed differently (compare Figs. 2 b, 2f and figs. 37 & 40 in Li & Yang 2008c). the sclerotized lateral margin of the new subspecies ends medially in a swollen apex whereas that of I. n. nigrihamita is forked medially (figs. 37 & 40 in Li & Yang 2008c)., Published as part of Li, Weihai, Zhang, Qin, Yang, Ding & Yao, Gang, 2017, A new Chinese species of Indonemoura (Plecoptera: Nemouridae) and a new subspecies of I. nigrihamita Li & Yang, pp. 255-262 in Zootaxa 4311 (2) on page 259, DOI: 10.11646/zootaxa.4311.2.6, http://zenodo.org/record/847544, {"references":["Li, W. H. & Yang, D. (2008 c) New species of Nemouridae (Plecoptera) from China. Aquatic Insects, 30, 205 - 221."]}

The article reviews the books "In Search of Paradise: Middle-Class Living in a Chinese Metropolis," by Li Zhang and "Marginalization in Urban China: Comparative Perspectives," edited by Fulong Wu and Chris Webster.

Epicephala daltonii Li, sp. nov. Figs. 3, 8, 13, 18 Type material. CHINA: Holotype ♂, Sichuan Province: Pingdi Town (26��12'N, 101��50'E), 1800 m, Renhe District, Panzhihua City, 28.iv.2013, collected from leaves of Glochidion daltonii by Z.G. Zhang, genitalia slide no. YXF13033. Paratypes (9 ♂, 7 ♀): Sichuan Province: 8 ♂, 6 ♀, same locality and host-plant as holotype, 1845 m, 26.iv ���01.v.2013, collected under light or reared from fruits of host-plant by Z.G. Zhang. Yunnan Province: 1 ♀, Jingdong Yi Autonomous county (24��28'N, 100��48'E), 1320 m, Puer ' City, 15.v.2014, collected from leaves of G. daltonii by Z.B. Wang; 1 ♂, same locality, 26.iv.2015, reared from fruits of G. daltonii by Z.G. Zhang. Description. Adult (Fig. 3). Forewing expanse 9.5���12.5 mm. Head snowy white; vertex tinged with pale yellowish white, with a few tufted grey scales laterally. Labial palpus with basal palpomere snowy white; second palpomere blackish brown on outer surface, white on inner surface; third palpomere blackish brown, pointed. Antenna with ventral surface dim yellow; dorsal surface dark brown, indistinctly greyish yellow at apex of each flagellomere; scape snowy white anteriorly. Thorax snowy white; tegula pale yellowish brown. Forewing deep greyish brown; costal margin with two white spots placed between distal 1/4 and 1/8, with three white streaks: basal streak from about 1/4, first extending along costal margin, then obliquely outward to 3/5 of wing, ending in a spot or streak; median streak from about middle slightly oblique to anterior 1/4, then parallel with costal margin to about distal 1/8, distally discontinuous in female; distal streak from beyond distal 1/4 extending slightly downward, then parallel with costal margin to distal 1/8, in some individuals discontinuous or represented by white spots or short streaks; ventral margin with a creamy band from base to end of fold, apically produced to a narrow slightly arched band extending to distal 1/8 at middle, another narrower streak from beyond it and parallel with it to distal 1/8, together forming a distinct pattern of four parallel and almost evenly spaced streaks; narrow greyish silvery fascia with metallic reflection at distal 1/8, edged with yellowish brown; distal 1/8 ochre brown, with a white dot at costal margin and a triangular spot along ventral margin, with an ill-defined central black dot beyond greyish silvery fascia, apex with a silvery dot; cilia greyish white along costal margin and apex except black distally and with a distinct black basal line, grey along dorsal margin. Hindwing and cilia greyish brown. Male genitalia (Fig. 8). Tegumen tongue-shaped, sclerotized laterally. Valva with costal part about same length as tegumen, subrectangular, bluntly rounded at apex; costa wide basally, tapering to a point, apically reaching apex of valva; ventral margin broadly and shallowly concave basally, slightly protruded and dentate at distal 1/3; transtilla slender, acute apically. Valva with sacculus elongate elliptical, about 3/4 length of costal part, approximately 1.5 times of costal part at maximum width; dorsal margin arched, distinctly curved in distal 1/3, in some individuals distinctly curved dorso-apically; ventral margin arched; apex narrowly rounded; with dense denticles distally, dentate along distal 2/5 of ventral margin. Vinculum broad V - shaped; saccus almost same length as arm of vinculum, narrow, less than 1/5 width of sacculus, distally narrowed to pointed apex. Phallus about same length as valva, straight, basal 2/5 slightly thicker, distal 3/5 uniform; cornutus an elongate ovate plate bearing numerous denticles, about 1/4 length of phallus. Female genitalia (Figs. 13, 18). Ovipositor cone-shaped, dentate laterally, acute apically. Apophysis posterioris longer than apophysis anterioris. Lamella postvaginalis torch-like; lateral margin arched, dentate sparsely; posterior margin concave in broad U shape, dentate, posterolaterally produced in triangle, pointed apically. Antrum as thick as ductus bursae, about 2/3 length of ductus bursae. Ductus bursae with sparsely toothed pleats; ductus seminalis arising from between antrum and ductus bursae. Corpus bursae oval, shorter than ductus bursae, full of reticular veins; signum a small rounded or subovate plate, wrinkled. Diagnosis. Epicephala daltonii sp. nov. is similar to E. ancylopa Meyrick, 1918 in appearance. It can be distinguished from the latter in the male genitalia by the sacculus about 3/4 length of the costal part and distinctly incurved in distal 1/3 dorsally (Fig. 8); in the female genitalia by the significantly large lamella postvaginalis concave in broad U shape on the posterior margin (Fig. 18) and the antrum about 2/3 the length of the ductus bursae. In E. ancylopa Meyrick, the sacculus is about 3/5 the length of the costal part and its dorsal margin is roundly arched in the male genitalia; the obviously small lamella postvaginalis is concave in broad V or U shape on the posterior margin (Fig. 16) and the antrum is about 1/3 the length of the ductus bursae in the female genitalia. Biology. Host-plant: Glochidion daltonii (Muell. Arg.) Kurz (Phyllanthaceae). Larvae feed on seeds in the fruits. Distribution. China (Sichuan, Yunnan). Etymology. The specific name is derived from the name of the host-plant Glochidion daltonii (Muell. Arg.) Kurz., Published as part of Li, Houhun & Zhang, Zhenguo, 2016, Five species of the genus Epicephala Meyrick, 1880 (Lepidoptera: Gracillariidae) from China, pp. 391-405 in Zootaxa 4084 (3) on pages 401-402, DOI: 10.11646/zootaxa.4084.3.5, http://zenodo.org/record/1052189, {"references":["Meyrick, E. (1918) Exotic Microlepidoptera, 2 (6), 161 - 192. [Marlborough]."]}

Epicephala assamica Li, sp. nov. Figs. 5, 10, 15 Type material. CHINA: Holotype ♂, Yunnan Province: Xishuangbanna Tropical Botanical Garden (21��41'N, 101��25'E), Chinese Academy of Sciences, Mengla, 550 m, 13.ix.2013, reared from fruit of Glochidion assamicum by Z.G. Zhang. Paratypes (530 ♂, 720 ♀): Yunnan Province: 55 ♂, 32 ♀, same data as holotype except 11���27.ix.2013; 112 ♂, 103 ♀, same data as holotype except 19���24.iii.2014; 104 ♂, 69 ♀, same data as holotype except 26.ix ���02.x.2014; 219 ♂, 493 ♀, same locality as holotype, 19���29.iv.2015, collected from leaves or reared from fruits of G. assamicum by Z.G. Zhang and K.J. Teng; 7 ♂, 2 ♀, Puer ' Sun River National Park (22��36'N, 101��07'E), 1200 m, 13���15.iv.2014, reared from fruits of G. assamicum by Z.G. Zhang. Hainan Province: 17 ♂, 8 ♀, Yinggeling Mountain Nature Reserves (19��01'N, 109��33'E), 450 m, 14���25.ix.2010, collected from leaves or reared from fruits of G. assamicum by B.B. Hu; 16 ♂, 13 ♀, same locality, 21-28.ix.2013, reared from fruits of G. assamicum by Z.B. Wang. Description. Adult (Fig. 5). Forewing expanse 7.5���10.0 mm. Head white, vertex tinged with pale yellow, with a few tufted greyish brown scales laterally. Labial palpus with basal palpomere white; second and third palpomeres white on inner surface, brown on outer surface. Antenna with ventral surface pale greyish yellow, dorsal surface deep greyish brown, scape snowy white anteriorly. Thorax snowy white; tegula deep greyish brown. Forewing deep greyish brown to dark brown; costal margin with three white streaks: basal streak short, from about 1/4 extending outward to 1/3 of wing at anterior 1/5 to 2/5; median streak from about middle slightly oblique downward to anterior 1/3, then parallel with costal margin to about distal 1/8 discontinuously, indistinct in some individuals; distal streak from distal 1/5, extending below and subparallel with costal margin to distal 1/8 discontinuously, in some individual represented by three short streaks or irregular spots; short white streak between median and distal streaks placed below costal margin, slightly oblique; ventral margin with a broad creamy band from base to end of fold, dorsally sinuate, indistinctly produced to a dorsal strip at 2/3 which obliquely extends to middle of wing at posterior 1/4 to 1/3 or at middle, interrupted distally, narrow white line from beyond end of ventral band extending to distal 1/8; two white streaks from above end of fold subparallelly to distal 1/8, joined distally, dorsal streak stronger and longer; narrow silvery white fascia with metallic reflection at distal 1/8, edged with yellowish brown; distal 1/8 ochre brown, with a yellowish white spot at costal margin and a large white triangular spot along ventral margin, with a central black dot or short transverse strip at middle beyond silvery fascia, apex with a silvery strip; cilia greyish white along costal margin and apex except black distally and a distinct black basal line, grey along dorsal margin. Hindwing and cilia greyish brown. Male genitalia (Fig. 10). Tegumen elongate, broadly tongue-shaped, sclerotized marginally. Valva with costal part about same length as tegumen; costa protruded before middle; ventral margin shallowly concave near base and before apex, protruded and dentate medially; apex bluntly rounded; transtilla narrow triangular, tapering distally. Valva with sacculus slightly shorter than 2/3 length of valva, subrectangular, slightly widened to before apex; apex rounded and dentate, slightly wider than costal part at maximum width. Vinculum broadly V-shaped; saccus almost same length as arm of vinculum, broad, about 4/5 width of sacculus, subparallel from base to about 3/5, then narrowed to rounded apex. Phallus about same length as valva, slightly dilated basally, curved and almost uniform distally, apically with two amalgamated but occasionally separated denticles (Figs. 10c, 10d); cornutus a row of clustered short spines. Female genitalia (Fig. 15). Ovipositor narrow, cone-shaped, dentate laterally, acute apically. Apophysis posterioris stronger and longer than apophysis anterioris. Lamella postvaginalis inverted trapezoidal; lateral margin obliquely straight, finely dentate in some individuals; posterior margin weakly sinuate, posterolaterally produced to a long thorn-like process, sharp apically. Antrum slightly shorter than ductus bursae, anteriorly narrowed. Ductus bursae with smooth sclerotized pleats; ductus seminalis arising from between antrum and ductus bursae. Corpus bursae oval, about same length as ductus bursae; signum a small overlapped subtriangular plate. Diagnosis. Epicephala assamica sp. nov. is similar to E. frenata Meyrick, 1908 in appearance and male genitalia. It can be separated from the latter in the male genitalia by the sacculus narrower than 1.5 times of costal part at maximum width, and the phallus with two denticles at apex. In E. frenata Meyrick, the sacculus is wider than 2 times of costal part at maximum width, and the phallus has a small apical thorn in the male genitalia. Epicephala assamica sp. nov. is also similar to E. ancylopa, and the differences between them can be found under E. ancylopa Meyrick. Biology. Larvae feed on seeds in the fruits of G. assamicum (M��ll. Arg.) Kook. f. (Phyllanthaceae). Distribution. China (Hainan, Yunnan). Etymology. The specific name is derived from the name of the host-plant Glochidion assamicum (Muell. Arg.) Hook., Published as part of Li, Houhun & Zhang, Zhenguo, 2016, Five species of the genus Epicephala Meyrick, 1880 (Lepidoptera: Gracillariidae) from China, pp. 391-405 in Zootaxa 4084 (3) on pages 403-404, DOI: 10.11646/zootaxa.4084.3.5, http://zenodo.org/record/1052189

Epicephala jianfenglingina Li, sp. nov. Figs. 4, 9, 14, 19 Type material. CHINA: Holotype ♂, Hainan Province: Jianfengling Nature Reserves (18°44'N, 108°51'E), 950 m, 26.ix.2010, reared from fruits of Glochidion sp. by B.B. Hu, genitalia slide no. HBB10039. Paratypes (2 ♀ f): Hainan Province: 1 ♀, same data as holotype except 24.ix.2010; 1 ♀, same locality and reared from fruits of same host-plant as holotype by J. Zhang on 15.iv.2011. Description. Adult (Fig. 4). Forewing expanse 9.5−10.5 mm. Head white, vertex tinged with a few greyish brown scales laterally. Labial palpus with basal palpomere white; second and third palpomeres white on inner surface, blackish brown on outer surface; third palpomere white tipped, pointed. Antenna with ventral surface pale greyish yellow, dorsal surface deep greyish brown, scape snowy white anteriorly. Thorax snowy white; tegula deep greyish brown. Forewing deep greyish brown; costal margin with three white streaks: basal streak short, from about 1/4, first along costal margin for a short distance, then extending discontinuously and weakening to 1/3 of wing at anterior 1/4; median streak from beyond middle slightly oblique downward to anterior 1/4, then weakening and parallel with costal margin to about distal 1/8 discontinuously; distal streak from distal 1/4, extending below and subparallel with costal margin to distal 1/8, interrupted; short white streak between median and distal streaks placed below costal margin, slightly oblique; ventral margin with a broad creamy band from base to end of fold, dorsally produced to a strip at 2/3 obliquely extending to 3/5 of wing at middle; two white streaks from above end of fold subparallel extending to distal 1/8, stronger in female than in male; narrow greyish silvery fascia with metallic reflection at distal 1/8, edged with yellowish brown, with a yellowish white dot at costal margins in male; distal 1/8 ochre brown, with a small white spot at costal margin and a narrow triangular spot along ventral margin, with a distinct central black dot, apex with a silvery strip; cilia greyish white along costal margin and apex except black distally and with a distinct black basal line, grey along dorsal margin. Hindwing and cilia greyish brown. Male genitalia (Fig. 9). Tegumen elongate, broad tongue shaped, sclerotized marginally. Valva with costal part about same length as tegumen, widened medially, obliquely rounded apically; costa slightly protruded medially; ventral margin with a dentate semicircular protuberance medially, concave basally and before apex; transtilla narrow triangular, tapering distally. Valva with sacculus somewhat fusiform, slightly shorter than 2/3 length of costal part, nearly as wide as costal part at maximum width; dorsal margin roundly arched, ventral margin almost straight, apex narrowly rounded; distal 1/4 with dense denticles. Vinculum in broad V shape; saccus almost same length as arm of vinculum, broad, slightly less than half width of sacculus, parallel except distally narrowed to rounded apex. Phallus about same length as valva, subovately dilated basally, uniform distally; cornutus a row of clustered long spines. Female genitalia (Figs. 14, 19). Ovipositor cone-shaped, dentate laterally, acute apically. Apophysis posterioris longer than apophysis anterioris. Lamella postvaginalis somewhat H shaped; lateral margin subparallel, sinuate; posterior margin deeply concave, broadly V-shaped, posterolaterally produced to a triangular process narrowly pointed at apex, anterolaterally produced to a triangular process with sharp apex. Antrum about 1/3 length of ductus bursae. Ductus bursae thicker than antrum, with sparsely dentate pleats; ductus seminalis arising from between antrum and ductus bursae. Corpus bursae oval, granulated, about half length of ductus bursae; signum a small subtriangular plate. Diagnosis. This new species is similar to E. duoplantaria sp. nov. in appearance. It can be further separated from the latter in the male genitalia by the phallus subovately dilated basally; in the female genitalia by the lamella postvaginalis with lateral margins subparallel and posterior margin deeply concave in broad V shape. In E. duoplantaria sp. nov., the phallus is slightly expanded basally in the male genitalia; the lamella postvaginalis is arched laterally and shallowly concave on the posterior margin in the female genitalia. Biology. Host-plant: Glochidion sp., close to Glochidion rubrum Bl. (Phyllanthaceae). Larvae feed on seeds in the fruits. Distribution. China (Hainan). Etymology. The specific name is derived from the name of the type locality.