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The family Inversiulidae, and its only genus Inversiula, are unique among cheilostome bryozoans owing to the reverse orientation of the operculum, which hinges distally instead of proximally. Inversiula currently includes four extant and one fossil species almost exclusively distributed in the Southern Hemisphere. A phylogenetic analysis of the Inversiulidae based on morphological characters was performed, showing the existence of two clades with contrasting development of the frontal shield. As a result of this analysis, Parainversiula gen. nov. is erected to contain inversiulid species with an umbonuloid frontal shield structure and fewer than 30 central spoked pores. Two poorly known fossil species from the early Miocene of Patagonia (Argentina), originally assigned to the genus Microporella, are here redescribed as Parainversiula sp. and P. chubutiana (Canu) comb. nov. The historical, time-calibrated biogeography of Inversiulidae was estimated using an event-based, multi-model quantitative analysis. The analyses estimated that Australasia was the ancestral area of Parainversiula and a dispersion of the genus into South America occurred between the late Eocene and Oligocene (c. 23–42 Ma), an event that could have been helped by the origin of the Antarctic Circumpolar Current. [ABSTRACT FROM AUTHOR]

Pelecanus paranensis sp. nov., a new pelican (Aves, Pelecanidae) from the marine Upper Miocene Paraná Formation, which crops out in the Province of Entre Ríos, Argentina, is described. This record constitutes the first report of a fossil pelican from Argentina and the southernmost from South America. The holotypical specimen consists of a very large and nearly complete pelvis, which is characterized by having the cristae iliacae dorsales continuous throughout its entire length and a large foramen acetabuli. The U-shaped morphology of the postacetabular section of the pelvis of the new species as well as the wide incisura sutura iliosynsacralis, allow to infer its phylogenetic position within the New World pelican species clade, showing a close relationship with the clade (P. occidentalis + P. thagus). A probable transAtlantic dispersal route for the ancestor of the New World pelicans is thus inferred. The inland Paranaense Sea, which flooded the South American Chaco-Paraná basin during the mid-Neogene, is proposed as a south-north pathway for ancestral forms of the clade (P. occidentalis + P. thagus). These regressive marine paleoenvironments of the Late Miocene may have acted as the evolutionary driver for the transition of pelican species from brackish or freshwater habitats to those inhabiting strictly marine coastlines. [ABSTRACT FROM AUTHOR]

This study describes two new species of cheilostome bryozoans associated with gastropod shells from the Monte León Formation (Burdigalian, early Miocene), Argentine Patagonia. Burdwoodipora griffini sp. nov. is the second known species and the first fossil representative of the genus, whose geographic distribution is so far restricted to the Magellan region in the southern southwest Atlantic. it differs from the type species, B. paguricola, in lacking suboral adventitious avicularia in most autozooids. Odontoporella miocenica sp. nov. is morphologically very close to the type species, the Recent O. adpressa, from which it differs in having smaller zooids with fewer areolae and relatively wider orifices. The stratigraphic range of Odontoporella is here extended from the Pleistocene to the early Miocene. its geographic distribution includes southern south America, New Zealand, and New caledonia, suggesting an austral origin of the genus in the Paleogene of former Gondwana. The possible association of these two species with paguroid crabs is discussed. [ABSTRACT FROM AUTHOR]

Fossil remains of extinct mylodontine sloths have been discovered in numerous localities throughout the American supercontinent, but knowledge of them is still mainly centered on the Pleistocene forms rather than their Neogene relatives. In this contribution, previously unpublished cranial and postcranial materials herein ascribed to Glossotherium chapadmalense (Kraglievich), coming from the Pliocene Chapadmalal Formation (Province of Buenos Aires, Argentina), are presented. These remains are described and compared with those of other mylodontine species from South and North America, providing an exhaustive characterization of cranial morphology in G. chapadmalense and some preliminary insights into its postcranial anatomy. The results have revealed a unique combination of differences and similarities of this enigmatic species with the larger-sized representatives of Glossotherium. In addition, these materials allowed shedding light on the long-standing uncertainties regarding its taxonomic status, as this form has been alternately considered to belong to the genera Glossotherium or Glossotheridium over the last century. As a result, the genus Glossotherium was present, at least, at the Early–Late Pliocene transition in the Pampean region of Argentina, and is recorded by the Upper Pleistocene throughout the entire South American continent. The radiation of the Pleistocene species is associated with a consistent body mass increase that was probably a response to the cooling temperatures and higher predation pressure in South America during the Late Pliocene–Early Pleistocene. These phenomena were probably driven by the stable emergence of the Panamanian Isthmus around 3 Ma and the successive faunal turnover pulses of the Great American Biotic Interchange. [ABSTRACT FROM AUTHOR]

Bryozoans with calcified frontal shields formed by the fusion of costae, collectively constituting a spinocyst, are traditionally assigned to the family Cribrilinidae. Today, this family is regarded as nonmonophyletic. In the Argentine Cenozoic, cribrilinids were until recently represented by only two fossil species from the Paleocene of Patagonia. This study describes the first fossil representatives of Jolietina and Parafigularia: J. victoria n. sp. and P. pigafettai n. sp., respectively. A fossil species of Figularia, F. elcanoi n. sp., is also described. The material comes from the early Miocene of the Monte León and Chenque formations (Patagonia, Argentina). For comparison, we also provide redescriptions of the remaining extant species of Jolietina: J. latimarginata (Busk, 1884) and J. pulchra Canu and Bassler, 1928a. The systematic position of some species previously assigned to Figularia is here discussed. Costafigularia n. gen. is erected, with Figularia pulcherrima Tilbrook, Hayward, and Gordon, 2001 as type species. Two species previously assigned to Figularia are here transferred to Costafigularia, resulting in C. jucunda n. comb. and C. tahitiensis n. comb. One species of Figularia is reassigned to Vitrimurella, resulting in V. ampla n. comb. The family Vitrimurellidae is here reassigned to the superfamily Cribrilinoidea. The subgenus Juxtacribrilina is elevated to genus rank. Inferusia is regarded as a subjective synonym of Parafigularia. Parafigularia darwini Moyano, 2011 is synonymized with I. taylori Kuklinski and Barnes, 2009, resulting in Parafigularia taylori n. comb. Morphological data suggest that these genera comprise different lineages, and a discussion on the disparities among cribrilinid (sensu lato) spinocysts is provided. UUID: http://zoobank.org/215957d3-064b-47e2-9090-d0309f6c9cd8 [ABSTRACT FROM AUTHOR]

The cheilostome bryozoan genus Taylorius Gordon, 2014 comprises six named living and fossil species from New Zealand and one Recent species from South Africa. Taylorus nom. nov. is proposed as a replacement name for Taylorius Gordon, 2014 (Bryozoa), a secondary homonym of Taylorius Britton, 1960 (Arthropoda: Coleoptera). Here, we describe Taylorus patagonicus sp. nov. from the early Miocene of Argentine Patagonia, and two new living species from New Zealand, Taylorus microperforatus sp. nov. and Taylorus alatus sp. nov. These new finds extend the known range of Taylorus nom. nov. to areas formerly comprising the Gondwanan supercontinent. Cladistic analysis suggests that there is a basal divergence within Taylorus nom. nov. in which an extant lineage consisting of species from South Africa and New Zealand is the sister group to the remaining species. Two New Zealand fossil taxa, T. masoni and T. waiparaensis, were not recovered in an exclusively Zealandian clade. The early Miocene New Zealand species T. masoni nested with three Recent Zealandian species, this subclade being sister to a group comprising the remaining New Zealand species and that from Patagonia. These different lineages suggest that Taylorus nom. nov. originated in the Paleogene, perhaps in the early Oligocene or late Eocene. [ABSTRACT FROM AUTHOR]

We present the first taphonomic analysis on basal sauropodomorph dinosaur remains from the Los Colorados Formation. We analyzed 22 bone samples with X-ray diffractometry, three of these in petrographic sections and one of them in SEM. In this article we recognize the taphonomic processes (biostratinomic and fossil-diagenetic) involved in the genesis of the 'La Esquina Local Fauna'. Regarding fossil diagenesis, it was possible to recognize the different processes that acted on the bone remains. In this sense, we analyzed different types of fracture present in the bones and their relationship with expansion generated by changes in the volume of minerals (for example, carbonates, gypsum and anhydrite). Finally, the importance of more taphonomic studies throughout the Los Colorados Formation is highlighted in order to better understand the vertebrate fossil record it contains. • Sauropodomorph bones were affected by different taphonomic processes. • At least three types of fractures associated with volume changes of minerals. • The fractures analyzed in the bones are of fossil-diagenetic origin. • The fossil record of vertebrate remains is found in well-developed overbank/floodplain deposits and rarely within fluvial channel facies. • Major bone loss can be explained by actual exhumation. [ABSTRACT FROM AUTHOR]

We examine three avian remains from the Última Esperanza cave complex. This site is a Natural Monument situated along the flanks of Cerro Benítez, Magallanes Region of Chile. A tarsometatarsus (MLP 94-VIII-10-12) is assigned to Rhea pennata, and a humerus (MLP 94-VIII-10-111) to Phalacrocorax brasilianus. Both, R. pennata and P. brasilianu s, are currently distributed in the Magallanes Region, and their presence within the cave can be easily explained by the cut marks found on the tarsometatarsus, that evidence human consumption. The absence of traces on the humerus is not decisive in this regard, because the wings are not the most fleshy parts of the body and therefore, not the preferred piece by predators. It can represent the waste. A small bill (MLP 94-VIII-10-140) of a passerine bird belongs to a fresh bone and should be treated as contamination. • This contribution presents the first study of the Cueva del Milodón avian fossil record. • A humerus assigned to Phalacrocorax brasilianus and a tarsometatarsus of Rhea pennata, belong to species that occur in the area today. • The cut marks observed in Rhea pennata evidence anthropic interaction. • A bill from Cueva del Milodón belongs to a recent passerine bird and is considered contamination. [ABSTRACT FROM AUTHOR]

The analysis of the hyoid apparatus of fossil xenarthrans provides insight on the form of the tongue and its function in food intake and intraoral processing. The hyoid apparatus of xenarthrans is notable for fusion among its elements. The presence of a V-bone, a complex consisting of fused basihyal and thyrohyal bones, is a consistent and probably synapomorphic feature of xenarthrans. Fusion of other elements is variable in fossil xenarthrans. Most fossil sloths retain independent elements, as in living dasypodids and mammals generally. Among nothrotheriids, the elements are slender and their articular surfaces indicate considerable mobility, and the relatively long and horizontal orientation of the geniohyoid muscle suggests considerable tongue protrusion. Among mylodontines, such as Paramylodon and Glossotherium, the elements indicate relatively mobile articulations, except between the stylo- and epihyals. The relatively posterior placement of the apparatus and the length and alignment of the geniohyoid muscle indicate considerable capacity for tongue protrusion. Scelidotherium, however, had rigidly articulated stylohyal and epihyal, and the apparatus lies farther anteriorly, which together with the elongated, steeply inclined mandibular symphysis, indicates a relatively shorter geniohyoid muscle and thus more limited capacity for tongue protrusion. A similar situation is indicated for Megatherium, casting doubt on the classical reconstruction of this sloth as having a long prehensile tongue. Among cingulates Prozaedyus resembles living dasypodids, indicating considerable tongue protrusion important in food acquisition and intake. More extensive fusion of hyoid elements occurs in the cingulates Glyptodon and Proeutatus, in which the stylohyal and epihyal at least, are fused into a single element termed the sigmohyal. The presence of this element supports recent proposals of a sister-group relationship between glyptodonts and eutatines. The rigidity of the apparatus suggests limited tongue protrusion, but the tongue, in glyptodonts at least, was a powerful structure important for intraoral manipulation of food.