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Abstract The repeated expansion of East Asian steppe cultures was a key driver of Eurasian history, forging new social, economic, and biological links across the continent. Climate has been suggested as important driver of these poorly understood cultural expansions, but paleoclimate records from the Mongolian Plateau often suffer from poor age control or ambiguous proxy interpretation. Here, we use a combination of geochemical analyses and comprehensive radiocarbon dating to establish the first robust and detailed record of paleohydrological conditions for Lake Telmen, Mongolia, covering the past ~ 4000 years. Our record shows that humid conditions coincided with solar minima, and hydrological modeling confirms the high sensitivity of the lake to paleoclimate changes. Careful comparisons with archaeological and historical records suggest that in the vast semi-arid grasslands of eastern Eurasia, solar minima led to reduced temperatures, less evaporation, and high biomass production, expanding the power base for pastoral economies and horse cavalry. Our findings suggest a crucial link between temperature dynamics in the Eastern Steppe and key social developments, such as the emergence of pastoral empires, and fuel concerns that global warming enhances water scarcity in the semi-arid regions of interior Eurasia.

Background: The archaeological and ethnographic heritages of Mongolia reflect a multi-millennial continuity of typically mobile-pastoral occupations across sparsely populated, environmentally diverse landscapes, but the threats of modernisation and industrialisation to those heritages are nevertheless present and substantial. The construction of the Erdeneburen Hydroelectric Dam on the Khovd River in western Mongolia is planned to submerge hundreds of archaeological features and jeopardise at least another thousand. Methods: The Mongolian Archaeology Project: Surveying the Steppes, in collaboration with the Mongolian Institute of Archaeology, integrates a variety of digital techniques including GIS (geographic information systems), Machine Learning automated site detection, drone mapping, and Structure-from-Motion LiDAR scanning to document the endangered archaeology. This paper presents the resulting dataset of archaeological features across three different impact zones associated with the dam construction and evaluates the degree of efficacy of the initial data integration strategy through informal partner feedback and self-assessment. Results: While only approximately 20% of the documented sites fall within the planned flood zone, the remaining sites will be subjected to collateral threats such as industrial and infrastructural development that will necessitate extended monitoring, both temporally and spatially. In consideration of these results, this paper argues that a ‘responsive’ mode of heritage disaster intervention can bridge the gap between ‘reactive’ and ‘proactive’ modes, but requires development of an integrated (digital) methodology. Conclusions: The paper concludes by offering a new, more interconnected ‘transmethodology’ that addresses spatiality, sub-sampling, data reuse, and community input across multiple disciplines such as cultural heritage preservation, salvage archaeology, computer vision, and community archaeology. The authors developed this ‘transmethodology’ and the resulting workflows out of a theoretical framework that considers principles of Symmetrical Archaeology, Resilience Humanitarianism, and the CARE standard for inclusive data management (Collective benefit, Authority to control, Responsibility, and Ethics).

Innovations in horse equipment during the early Middle Ages provided advantages to societies from the steppes, reshaping the social landscape of Eurasia. Comparatively little is known about the precise origin of these crucial advances, although the available evidence points to early adoption in East Asia. The authors present new archaeological discoveries from western and northern Mongolia, dating to the fourth and fifth centuries AD, including a wooden frame saddle with horse hide components from Urd Ulaan Uneet and an iron stirrup from Khukh Nuur. Together, these finds suggest that Mongolian groups were early adopters of stirrups and saddles, facilitating the expansion of nomadic hegemony across Eurasia and shaping the conduct of medieval mounted warfare. Keywords: Mongolia, Middle Ages, horse riding, stirrup, saddle, Introduction Across Eurasia, the development of sophisticated cavalry had tremendous consequences for ancient societies. Beginning with the domestication of the horse and the subsequent invention of the chariot in the [...]

The extremely rich palaeontological record of the horse family, also known as equids, has provided many examples of macroevolutionary change over the last ~55 Mya. This family is also one of the most documented at the palaeogenomic level, with hundreds of ancient genomes sequenced. While these data have advanced understanding of the domestication history of horses and donkeys, the palaeogenomic record of other equids remains limited. In this study, we have generated genome‐wide data for 25 ancient equid specimens spanning over 44 Ky and spread across Anatolia, the Caucasus, Central Asia and Mongolia. Our dataset includes the genomes from two extinct species, the European wild ass, Equus hydruntinus, and the sussemione Equus ovodovi. We document, for the first time, the presence of sussemiones in Mongolia and their survival around ~3.9 Kya, a finding that should be considered when discussing the timing of the first arrival of the domestic horse in the region. We also identify strong spatial differentiation within the historical ecological range of Asian wild asses, Equus hemionus, and incomplete reproductive isolation in several groups yet considered as different species. Finally, we find common selection signatures at ANTXR2 gene in European, Asian and African wild asses. This locus, which encodes a receptor for bacterial toxins, shows no selection signal in E. ovodovi, but a 5.4‐kb deletion within intron 7. Whether such genetic modifications played any role in the sussemione extinction remains unknown. [ABSTRACT FROM AUTHOR]

While few places on earth have been as deeply impacted by the human-horse relationship as the steppes of Mongolia and eastern Eurasia, gaps in the archaeological record have made it strikingly difficult to trace when and how the first domestic horses were integrated into ancient societies in this key region of the world. Recently, organic materials preserved in melting mountain ice have emerged as a key source of archaeological insight into the region's deep past. Newly-identified artefacts recovered from melting snow and ice in the Altai Mountain range of western Mongolia (including metal artefacts, skeletal remains, and hoof fragments) provide archaeological evidence for the use of horses at high altitudes from the Bronze Age through the 20th century. Direct radiocarbon dating and genomic sequencing demonstrate the presence of Przewalski's horse in the region during the early second millennium BCE, suggesting that this taxon may have once foraged at high altitudes frequented by human hunters. Importantly, directly-dated remains of horse hoof trimmings provide some of the oldest direct evidence of horse transport in the Eastern Steppe as early as the 14th century BCE, and suggest a role for high-mountain hunting in the innovation of reliable mounted riding. [ABSTRACT FROM AUTHOR]

Across Eurasia, horse transport transformed ancient societies. Although evidence for chariotry is well dated, the origins of horse riding are less clear. Techniques to distinguish chariotry from riding in archaeological samples rely on elements not typically recovered from many steppe contexts. Here, the authors examine horse remains of Mongolia's Deer Stone-Khirigsuur (DSK) Complex, comparing them with ancient and modern East Asian horses used for both types of transport. DSK horses demonstrate unique dentition damage that could result from steppe chariotry, but may also indicate riding with a shallow rein angle at a fast gait. A key role for chariots in Late Bronze Age Mongolia helps explain the trajectory of horse use in early East Asia. Keywords: Eurasian Steppe, Mongolia, Bronze Age, horse domestication, chariots, horseback riding, Introduction Domestic horses (Equus caballus) facilitated the emergence of highly mobile pastoral lifeways and the transcontinental equestrian empires of the steppes of Mongolia and eastern Eurasia. Although historical documents trace [...]

The vast mortuary complexes of the Xiongnu, the world's first nomadic empire (c. 200 BC--AD 100), were important statements of elite power and ritual commemoration in Inner Asia. Very few of the features that accompanied the main tombs, however, have been fully excavated and investigated. This study is one of the first to assess completely the small archaeological features--and associated faunal remains--that surround the more monumental structures, features that intimate substantial investments in, and ritual activities around, these mortuary arenas. This research provides an important contribution to the understanding of the social politics of ritual practices and the development of complex institutions in steppe pastoral societies.Keywords: Inner Asia, Mongolia, Xiongnu, animal sacrifice, ritual, IntroductionDuring the first millennium BC, the Inner Asian steppe exhibited increasingly ostentatious mortuary monuments that displayed the power of nomadic elites and facilitated the interregional consolidation of the Xiongnu steppe [...]

Millet is a highly adaptable plant whose cultivation dramatically altered ancient economies in northern Asia. The adoption of millet is associated with increased subsistence reliability in semi-arid settings and perceived as a cultigen compatible with pastoralism. Here, we examine the pace of millet's transmission and locales of adoption by compiling stable carbon isotope data from humans and fauna, then comparing them to environmental variables. The Bayesian modelling of isotope data allows for the assessment of changes in dietary intake over time and space. Our results suggest variability in the pace of adoption and intensification of millet production across northern Asia. [ABSTRACT FROM AUTHOR]

The adoption of the horse for chariots, wagons and riding had a major impact on human societies, but it has proved difficult to reliably identify early domesticated horses in the archaeological record. This comparative study of equine palaeopathology addresses the problem by analysing wild and domestic horses used for traction or riding. Osteological changes to the skull appear to be the result of mechanical and physiological stress from the use of horses for transport. The results are applied to archaeological examples from the Deer Stone-Khirigsuur Complex of Bronze Age Mongolia (1300-700 BC) and show that those horses were probably bridled and used for transport. Keywords: Mongolia, Bronze Age, Deer Stone-Khirigsuur Complex, equestrianism, zooarchaeology, nasal remodelling, Introduction Horseback riding and the use of chariots has been linked with dramatic changes to the form and scale of social organisation among prehistoric peoples, but the chronology of their [...]

Domesticated yaks endure as iconic symbols of high-altitude frozen landscapes, where herding communities depend on their high-fat milk, transport, dung, and natural fibers. While there is established proteomic evidence for ancient consumption of ruminant and horse milk in the mountains and steppes of northern Eurasia, yak dairy products have yet to be detected. Yak domestication and the species' dispersal from Tibet into the mountainous zones to the north are also poorly resolved due to a paucity of zooarchaeological data. To examine the potential of paleoproteomics to shed light on domesticated yak in Mongolia, we analyzed human dental calculus from Mongol era elite individuals recovered from permafrost burials in Khovsgol province, where people continue to herd yak to this day. We report the first evidence for yak dairy consumption, linked to local resource control. In addition, we confirm a large diversity of recovered whey, curd, tissue, and blood proteins, likely reflecting the excellent preservation conditions found at permafrost sites. Proteomic evidence from permafrost human dental calculus indicates consumption of yak milk in Mongolia as early as 1270 CE. [ABSTRACT FROM AUTHOR]

The initial movement of herders and livestock into the eastern steppe is of great interest, as this region has long been home to pastoralist groups. Due to a paucity of faunal remains, however, it has been difficult to discern the timing of the adoption of domesticated ruminants and horses into the region, though recent research on ancient dairying has started to shed new light on this history. Here we present proteomic evidence for shifts in dairy consumption in the Altai Mountains, drawing on evidence from sites dating from the Early Bronze to the Late Iron Age. We compare these finds with evidence for the rise of social complexity in western Mongolia, as reflected in material remains signaling population growth, the establishment of structured cemeteries, and the erection of large monuments. Our results suggest that the subsistence basis for the development of complex societies began at the dawn of the Bronze Age, with the adoption of ruminant livestock. Investments in pastoralism intensified over time, enabling a food production system that sustained growing populations. While pronounced social changes and monumental constructions occurred in tandem with the first evidence for horse dairying, ~1350 cal BCE, these shifts were fueled by a long-term economic dependence on ruminant livestock. Therefore, the spread into the Mongolian Altai of herds, and then horses, resulted in immediate dietary changes, with subsequent social and demographic transformations occurring later. [ABSTRACT FROM AUTHOR]

Goats were initially managed in the Near East approximately 10,000 years ago and spread across Eurasia as economically productive and environmentally resilient herd animals. While the geographic origins of domesticated goats (Capra hircus) in the Near East have been long-established in the zooarchaeological record and, more recently, further revealed in ancient genomes, the precise pathways by which goats spread across Asia during the early Bronze Age (ca. 3000 to 2500 cal BC) and later remain unclear. We analyzed sequences of hypervariable region 1 and cytochrome b gene in the mitochondrial genome (mtDNA) of goats from archaeological sites along two proposed transmission pathways as well as geographically intermediary sites. Unexpectedly high genetic diversity was present in the Inner Asian Mountain Corridor (IAMC), indicated by mtDNA haplotypes representing common A lineages and rarer C and D lineages. High mtDNA diversity was also present in central Kazakhstan, while only mtDNA haplotypes of lineage A were observed from sites in the Northern Eurasian Steppe (NES). These findings suggest that herding communities living in montane ecosystems were drawing from genetically diverse goat populations, likely sourced from communities in the Iranian Plateau, that were sustained by repeated interaction and exchange. Notably, the mitochondrial genetic diversity associated with goats of the IAMC also extended into the semi-arid region of central Kazakhstan, while NES communities had goats reflecting an isolated founder population, possibly sourced via eastern Europe or the Caucasus region. [ABSTRACT FROM AUTHOR]

In interior Eurasia, high mountain zones are crucial to pastoral subsistence, providing seasonally productive pastures and abundant wild resources. In some areas of northern Mongolia, mountainous tundra zones also support a low-latitude population of domestic reindeer herders–a lifestyle whose origins are poorly characterized in the archaeological record of early Mongolia. Traditionally, reindeer pastoralists make significant seasonal use of munkh mus (eternal ice) for their domestic herds, using these features to cool heat-stressed animals and provide respite from insect harassment. In recent years, many of these features have begun to melt entirely for the first time, producing urgent threats to traditional management techniques, the viability of summer pastures, and reindeer health. The melting ice is also exposing fragile organic archaeological materials that had previously been contained in the patch. We present the results of horseback survey of ice patches in Baruun Taiga special protected area, providing the first archaeological insights from the region. Results reveal new evidence of historic tool production and wild resource use for fishing or other activities, and indicate that ice patches are likely to contain one of the few material records of premodern domestic reindeer use in Mongolia and lower Central Asia. The area's ancient ice appears to be rapidly melting due to changing climate and warming summer temperatures, putting both cultural heritage and traditional reindeer herding at extreme risk in the years to come. [ABSTRACT FROM AUTHOR]

The emergence of mobile herding lifeways in Mongolia and eastern Eurasia was one of the most crucial economic and cultural transitions in human prehistory. Understanding the process by which this played out, however, has been impeded by the absence of a precise chronological framework for the prehistoric era in Mongolia. One rare source of empirically dateable material useful for understanding eastern Eurasia's pastoral tradition comes from the stone burial mounds and monumental constructions that began to appear across the landscape of Mongolia and adjacent regions during the Bronze Age (ca. 3000–700 BCE). Here, along with presenting 28 new radiocarbon dates from Mongolia's earliest pastoral monumental burials, we synthesise, critically analyse, and model existing dates to present the first precision Bayesian radiocarbon model for the emergence and geographic spread of Bronze Age monument and burial forms. Model results demonstrate a cultural succession between ambiguously dated Afanasievo, Chemurchek, and Munkhkhairkhan traditions. Geographic patterning reveals the existence of important cultural frontiers during the second millennium BCE. This work demonstrates the utility of a Bayesian approach for investigating prehistoric cultural dynamics during the emergence of pastoral economies. [ABSTRACT FROM AUTHOR]

Walled sites attributed to the Xiongnu steppe empire (2nd century b.c.–1st century a.d.) constitute the earliest investments in explicitly non-mortuary built environments among pastoralists of the Mongolian Steppe. These permanent constructions are important precedents to the medieval towns of subsequent steppe empires, yet very little of the complete layouts and surrounding remains of the large enclosures are understood. Our investigations of the walled site at Kherlen-Bars in eastern Mongolia are among the first surveys to fully document all aspects above and below ground across an entire Iron Age walled site area. In addition to detailed documentation of the large structures, our geomagnetic prospections and digital elevation modeling brought to light numerous small sub-surface and subtle-surface features that show such early walled sites, often deemed virtually empty, to be far more complex and diachronic in their formation processes and possible functions. [ABSTRACT FROM AUTHOR]

This paper presents the analysis of textiles from the tenth century Mongolian rock burial of Üzüür Gyalan. It is the only undisturbed rock burial discovered to date and presents a unique opportunity to examine a complete set of grave goods, both organic and inorganic, included in a Medieval horse-accompanied burial. The high degree of preservation and the prevalence of woven wool textiles set it apart from other rock burials where silk, sheepskin, and felt garments have been found. Certain features of the assemblage, including the abundance of the woven wools, their technical consistency, and evidence for household rather than specialised production, suggest the work of local weavers. Drawing comparisons with a living nomadic weaving tradition in Ladakh, Tibet, this paper relates aspects of the assemblage to production in a mobile pastoralist context. [ABSTRACT FROM AUTHOR]

Horse domestication revolutionized warfare and accelerated travel, trade, and the geographic expansion of languages. Here, we present the largest DNA time series for a non-human organism to date, including genome-scale data from 149 ancient animals and 129 ancient genomes (≥1-fold coverage), 87 of which are new. This extensive dataset allows us to assess the modern legacy of past equestrian civilizations. We find that two extinct horse lineages existed during early domestication, one at the far western (Iberia) and the other at the far eastern range (Siberia) of Eurasia. None of these contributed significantly to modern diversity. We show that the influence of Persian-related horse lineages increased following the Islamic conquests in Europe and Asia. Multiple alleles associated with elite-racing, including at the MSTN "speed gene," only rose in popularity within the last millennium. Finally, the development of modern breeding impacted genetic diversity more dramatically than the previous millennia of human management. • Two now-extinct horse lineages lived in Iberia and Siberia some 5,000 years ago • Iberian and Siberian horses contributed limited ancestry to modern domesticates • Oriental horses have had a strong genetic influence within the last millennium • Modern breeding practices were accompanied by a significant drop in genetic diversity Genome-wide data from 278 ancient equids provide insights into how ancient equestrian civilizations managed, exchanged, and bred horses and indicate vast loss of genetic diversity as well as the existence of two extinct lineages of horses that failed to contribute to modern domestic animals. [ABSTRACT FROM AUTHOR]

The Xiongnu established the first nomadic imperial power, controlling the Eastern Eurasian steppe from ca. 200 BCE to 100 CE. Recent archaeogenetic studies identified extreme levels of genetic diversity across the empire, corroborating historical records of the Xiongnu Empire being multiethnic. However, it has remained unknown how this diversity was structured at the local community level or by sociopolitical status. To address this, we investigated aristocratic and local elite cemeteries at the western frontier of the empire. Analyzing genome-wide data from 18 individuals, we show that genetic diversity within these communities was comparable to the empire as a whole, and that high diversity was also observed within extended families. Genetic heterogeneity was highest among the lowest-status individuals, implying diverse origins, while higher-status individuals harbored less genetic diversity, suggesting that elite status and power was concentrated within specific subsets of the broader Xiongnu population.

DNA recovery from ancient human remains has revolutionized our ability to reconstruct the genetic landscape of the past. Ancient DNA research has benefited from the identification of skeletal elements, such as the cochlear part of the osseous inner ear, that provides optimal contexts for DNA preservation; however, the rich genetic information obtained from the cochlea must be counterbalanced against the loss of morphological information caused by its sampling. Motivated by similarities in developmental processes and histological properties between the cochlea and auditory ossicles, we evaluate the ossicles as an alternative source of ancient DNA. We show that ossicles perform comparably to the cochlea in terms of DNA recovery, finding no substantial reduction in data quantity and minimal differences in data quality across preservation conditions. Ossicles can be sampled from intact skulls or disarticulated petrous bones without damage to surrounding bone, and we argue that they should be used when available to reduce damage to human remains. Our results identify another optimal skeletal element for ancient DNA analysis and add to a growing toolkit of sampling methods that help to better preserve skeletal remains for future research while maximizing the likelihood that ancient DNA analysis will produce useable results., (© 2020 Sirak et al.; Published by Cold Spring Harbor Laboratory Press.)