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This paper focuses on the anthropic dynamics of environmental transformation and natural resource management in the specific case of the archaeological site of Tossal de la Vila in Castelló, Spain, a hillfort located at the end of the Eastern Iberian Cordillera. It presents two phases of occupation determined by multiple radiocarbon dating analyses: the first phase during the late Bronze Age (8th–7th centuries BCE) and the second at the beginning of the al-Andalus period (8th–10th centuries CE). The results of the comparison of the subsistence strategies and the biotic configuration of the natural environment in the same place at two different times have shown us that in the protohistoric period, the diversity of economic activities represented in the archaeobiological record is certainly heterogeneous, while in the al-Andalus period the complexity of these options is reduced to much less significant limits, possibly due to the different functionality of this space in both periods. [ABSTRACT FROM AUTHOR]