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The foregoing investigations of the dynamics of the system (1), the results of which are largely similar to those obtained in our numerical analysis of Rössler's system [13] and to many other published data [1, 3-5, 11, 14, 15], indicate that quasihyperbolic attractors with a small fractional dimension in the domain above the critical state are typified by soft and hard bifurcations of the periodic trajectories. A bifurcation generating a stable two-dimensional torus from a cycle in nonautonomous three-dimensional systems in which the degree of contraction of the phase volume depends on the coordinates is clearly not realized [15], and for this effect to exist it is necessary to increase the dimension of the phase space to N=4 by introducing, for example, an external control signal. An analysis of systems with a forward Andronov-Hopf bifurcation shows that the typical mechanism of the evolution of stochasticity here is the mechanism of Feigenbaum doublings, against the background of which the mixing properties of hard bifurcations occur in the supercritical zone. [ABSTRACT FROM AUTHOR]

Root development is critical to successful establishment after seedlings are outplanted on a forest restoration site. There is an array of practices nursery practitioners and foresters use to ensure seedlings develop quality root systems to aid in outplanting success. To this end, a select number of topics are examined on how they effect root system quality and the linkage between these culturing practices (i.e., nursery root culturing practices), good root morphological and physiological quality and successful field establishment. The following discussion first describes why root development is critical for seedlings to avoid planting stress and successfully transition into field establishment. This is followed by reviewing the seasonality of root growth patterns, and how nursery cultural practices that create optimum levels of nutrients and non-structural carbohydrates affect root growth. Bareroot and container-grown stocktypes are discussed in relation to how their ability to grow roots affects seedling establishment. Lastly, nursery practices related to bareroot root culturing and container type selection for container-grown seedlings are examined on how they can affect seedling root form and field establishment. By focussing on these topics, the intent was to gain a renewed perspective on the importance of root system quality to ensure seedling establishment after outplanting on a forest restoration site. [ABSTRACT FROM AUTHOR]

Ryegrass (Lolium perenne L.), a high-quality forage grass, is a good nutrient source for herbivorous livestock. However, improving nitrogen use efficiency and avoiding nitrate toxicity caused by excessive nitrogen are continual challenges in ryegrass production. The molecular mechanism underlying the response of ryegrass to nitrogen, especially excessive nitrogen, remains unclear. In this study, the transcriptomic changes under different nitrogen levels were investigated in perennial ryegrass by high-throughput next-generation RNA sequencing. Phenotypic characterization showed that treatment with half of the standard N concentration (N0.5) led to a better growth state than the other three treatments. The treatments with the standard N concentration (N1) and treatments with ten times higher than the standard N concentration (N10) contained excessive nitrogen, which placed stress on plant growth. Analysis of differentially expressed genes indicated that 345 and 104 genes are involved in the regulation of nitrogen utilization and excessive nitrogen stress, respectively. KEGG enrichment analysis suggested that "photosynthesis-antenna proteins" may respond positively to appropriate nitrogen conditions, whereas "steroid biosynthesis", "carotenoid biosynthesis" and "C5-branched dibasic acid metabolism" were identified as the top significantly enriched pathways in response to excessive nitrogen. Additionally, 21 transcription factors (TFs) related to nitrogen utilization were classified into 10 families, especially the AP2-EREBP and MYB TF families. Four TFs related to excessive nitrogen stress were identified, including LOBs, NACs, AP2-EREBPs and HBs. The expression patterns of these selected genes were also analyzed. These results provide new insight into the regulatory mechanism of ryegrass in response to nitrogen utilization and excessive nitrogen stress. [ABSTRACT FROM AUTHOR]

Nitrogen (N) and light are critical determinants of plant growth and productivity. The present study attempts to quantify the underlying mechanisms and effects of light, N levels, and seed source on growth, biomass, nutrient allocations, and nutrient use efficiencies (NUEs) in seedlings of Pongamia pinnata. The differential and fixed hypothesis and isometric and allometric relationships were also examined. Six morphologically superior seed sources of P. pinnata from three contrasting agro-climatic conditions of India were evaluated with application of two levels of both N (low dose @ 5g plant−1 and 15 g plant−1) and light regimes (full light-100% and low light-50%) in mother nursery. Among the seed sources, the native Raipur seed source was found outstanding in growth, biomass, and nutrient use efficiency. N was more critical than light in promoting growth in seedlings of P. pinnata. High N and high light regimes increased shoot/root ratios. The components showed allometric growth and followed a differential allocation pattern. The exploitation of suitable genetic resources combined with managerial interventions is necessary for development of quality planting stock of P. pinnata. Nutrient use efficiencies, biomass production, and their allocation were important indicators and criteria for selection. The study suggests that Raipur and Jabalpur seed sources having high NUE should be grown along with adequate N application (15 g plant−1) and light conditions. [ABSTRACT FROM AUTHOR]

Background: The oak tree (Quercus aegilops) comprises ~ 70% of the oak forests in the Kurdistan Region of Iraq (KRI). Besides its ecological importance as the residence for various endemic and migratory species, Q. aegilops forest also has socio-economic values—for example, as fodder for livestock, building material, medicine, charcoal, and firewood. In the KRI, Q. aegilops has been degrading due to anthropogenic threats (e.g., shifting cultivation, land use/land cover changes, civil war, and inadequate forest management policy) and these threats could increase as climate changes. In the KRI and Iraq as a whole, information on current and potential future geographical distributions of Q. aegilops is minimal or not existent. The objectives of this study were to (i) predict the current and future habitat suitability distributions of the species in relation to environmental variables and future climate change scenarios (Representative Concentration Pathway (RCP) 2.6 2070 and RCP8.5 2070); and (ii) determine the most important environmental variables controlling the distribution of the species in the KRI. The objectives were achieved by using the MaxEnt (maximum entropy) algorithm, available records of Q. aegilops, and environmental variables. Results: The model demonstrated that, under the RCP2.6 2070 and RCP8.5 2070 climate change scenarios, the distribution ranges of Q. aegilops would be reduced by 3.6% (1849.7 km2) and 3.16% (1627.1 km2), respectively. By contrast, the species ranges would expand by 1.5% (777.0 km2) and 1.7% (848.0 km2), respectively. The distribution of the species was mainly controlled by annual precipitation. Under future climate change scenarios, the centroid of the distribution would shift toward higher altitudes. Conclusions: The results suggest (i) a significant suitable habitat range of the species will be lost in the KRI due to climate change by 2070 and (ii) the preference of the species for cooler areas (high altitude) with high annual precipitation. Conservation actions should focus on the mountainous areas (e.g., by establishment of national parks and protected areas) of the KRI as climate changes. These findings provide useful benchmarking guidance for the future investigation of the ecology of the oak forest, and the categorical current and potential habitat suitability maps can effectively be used to improve biodiversity conservation plans and management actions in the KRI and Iraq as a whole. [ABSTRACT FROM AUTHOR]

The biomass, carbon and nitrogen storage in the single tree components (foliage, branch, crown, bark, stem and total aboveground) of the grey poplar (Populus × canescens) in its distribution in Eastern Anatolia (Van, Turkey) were determined and modelled. The biomass, carbon and nitrogen storages were not estimated at a stand level but were based on single trees. Regression models based on the tree diameter at breast height (DBH) and total tree height (H) were developed to estimate the biomass, carbon (C) storage and nitrogen (N) storage of the different tree components of a total of 28 grey poplar trees. The two main regression models in the power function were developed based only on the DBH (Model 1) and the combination of the DBH and height (D2H) (Model 2). All regression models, except for those of the foliage components, developed to estimate the biomass and C and N storages of the tree components were found to be statistically significant (p < 0.001). The partitioning of the total aboveground biomass in the bark, foliage, branches and stems was 0.7, 9, 17 and 73%, respectively. The average C concentrations of the tree components ranged from 48 (foliage) to 50% (bark, branch and stem), while the N concentrations ranged from 0.35 (stem) to 1.32% (foliage). Higher biomass and lower nitrogen concentrations of foliage compared with cultivated poplars were likely related to the natural site conditions, low soil nitrogen and/or characteristics of single tree growth. [ABSTRACT FROM AUTHOR]

The morphological, biochemical, and physiological characteristics of silver birch ( Betula pendula Roth), curly birch ( B. pendula Roth var. carelica), and downy birch ( B. pubescens Ehrh.) at early stages of ontogenesis in natural conditions were investigated. Some intra- and interspecific peculiarities of the morphophysiological properties were determined. Priority development of the underground mass for the seedlings of curly birch and that of the aboveground mass for the seedlings of silver birch and downy birch were found. The leaf of curly birch is developed more actively as compared with those in the other two species under equal potential possibility of the growth of their leaves by elongation. A high similarity of the mineral composition (C, N, P, K) of different forms and species of birches was demonstrated. For silver birch the maximum values of the stomatal conductance, rate of photosynthesis, and transpiration under similar atmospheric and soil conditions were obtained. In downy birch we found a higher efficiency of water use. The highest values of the maximum rate of ribulose 1,5-bisphosphate carboxylase/oxygenase carboxylation were found in curly birch. This may serve as an indicator of its shade tolerance, as compared with silver birch and downy birch. [ABSTRACT FROM AUTHOR]

The aims of this study were to investigate the effects of water deficit and recovery on growth, photosynthesis and water relations in four Medicago laciniata populations from saharian (Ml-90), inferior arid (Ml-204), superior arid (Ml-306) and semi-arid (Ml-376) Tunisian regions. After 28 d of sowing with ample irrigation, the plants were subjected to 4 water regimes: optimal irrigation (100% of field capacity, FC), moderate drought (75% FC), severe drought (35% FC) and rewatering (plants submitted to 35% FC during 7 d, afterwards the plants were rewatered to 100% FC). Harvest was carried out after 28 d of treatments. The drought tolerance in M. laciniata populations was found to be increased particularly with increasing temperatures of collection site of the population. The Ml-204 and Ml-90 populations used mainly physiological strategies for survival under moderate water shortage. Higher severe drought tolerance in both signaled populations would be related to their lower photosynthesis metabolic impairment, relatively higher leaf RWC and greater osmotic potential decrease. The results suggest that plants with low values of leaf features are likely to maintain higher leaf RWC under sever drought. The largest decrease of osmotic potential was found associated with the solute accumulations such as proline and K. [ABSTRACT FROM AUTHOR]

Background and aims: Plant growth and photosynthetic ability have been frequently demonstrated to increase with nitrogen (N) supply. N can also promote changes in root water absorption and shoot water status via mechanisms that remain poorly understood. This study aims at investigating the effects of N supply on water absorption. Methods: A hydroponic experiment with two independent rice varieties (cv. 'Shangyou63' hybrid indica and cv. 'Yangdao6' conventional indica, China) supplied by three distinct N levels was performed in a greenhouse. Physiological characteristics were analyzed after a few weeks. Results: Compared to low N supply (20 mg·L), exposure to high N supply (100 mg·L) increased the light-saturated photosynthetic rate ( A) and water use efficiency (WUE) by 17 % and 22 %, respectively, in Shanyou63 and by 43 % and 26 %, respectively, in Yangdao6. The leaf water potential was significantly decreased in Shanyou63 but not in Yangdao6. There were increases in the rate of water uptake and the root hydraulic conductance ( L) under high N supply in both rice cultivars; these changes were accompanied by increased transcription levels of aquaporins (AQPs), decreased aerenchyma formation and root porosity, and decreased root lignin content. Under high N supply, Yangdao6 also exhibited much higher AQP activity, lower aerenchyma and root porosity compared with those of Shanyou63, indicating that Yangdao6 had an increased ability to absorb water compared with that of Shanyou63. Conclusions: The enhanced expression of AQPs and decreased root aerenchyma and lignin contributed to increased water absorption ability under high N supply. In addition, the responses of each of the two rice cultivars (hybrid and conventional) to N supply is related to their water uptake ability, resulting from root porosity and an increase in AQP activity. [ABSTRACT FROM AUTHOR]

Although light is a key factor in forestry, it is surprisingly seldom measured in day-to-day management of European forests. The spherical convex densiometer is a simple instrument that allows to evaluate the canopy openness (CO) by counting the number of 'canopy' dots on a grid lying on a convex mirror reflecting the canopy. In this contribution, we compared the performances of this instrument in mixed oak-beech hardwood forests spanning the lower end of the light gradient [1-17 % above canopy photosynthetically active radiation (PAR)] to two reference techniques: PAR measurements and fish-eye photography, based on a detailed analysis of its functioning. Discrepancies between the densiometer and the fish-eye estimates of CO were due to a combination of differences in dot resolution, dot counting and portion of the hemisphere considered. By contrast, the various effects of operator on densiometer estimates, including the influence of conformation on the angle of view, were found to be relatively minor. Densiometer readings were closely related to the relative light intensity assessed by PAR sensors in overcast conditions, which suggests that the use of this inexpensive tool should be expanded. [ABSTRACT FROM AUTHOR]

Remote sensing classifications of wetland ecosystems can be difficult because of temporal variability of plant cover and hydrological conditions. We developed a multitemporal classification scheme to control for these factors in forested wetlands by classifying pixels within a principal-components ordination of multitemporal reflectance, using data from three Landsat bands from each of seven scenes across a 3-year period in each of two phenological conditions. The principal components of reflectance that defined the classification were statistically related to vegetative conditions of the Louisiana field sites in interpretable ways, so the resulting classification represents the desired ecological gradients more robustly than single-image classifications. [ABSTRACT FROM AUTHOR]

Light availability strongly influences stream primary production, water temperatures and resource availability at the base of stream food webs. In headwater streams, light is regulated primarily by the riparian forest, but few studies have evaluated the influence of riparian forest stand age and associated structural differences on light availability. In this study, we evaluated canopy cover and streambed light exposure in four second-order streams within paired reaches of primary old-growth versus second-growth mature riparian forests. Stand age class was used as a proxy here for canopy complexity. We estimated stream canopy cover using a spherical densiometer. Local streambed light exposure was quantified and compared within and between reaches using fluorescein dye photodegradation. Reaches with complex old-growth riparian forests had frequent canopy gaps which lead to greater stream light availability compared to adjacent reaches with simpler second-growth riparian forests. We quantified light exposure at relatively high resolution (every 5 m) and also found greater variability in stream light along the reaches with old-growth riparian forests in three of the four streams. Canopy gaps were particularly important in creating variable light within and between reaches. This work demonstrates the importance of the age, developmental stage, and structure of riparian forests in controlling stream light. The highly variable nature of light on the stream benthos also highlights the value of multiple measurements of light or canopy structure when quantifying stream light. [ABSTRACT FROM AUTHOR]

Estimates of above-ground biomass are required for better planning, sustainable management and monitoring of changes in carbon stocks in agroforestry systems. The objective of this study was to develop and compare biomass equations for Markhamia lutea, Casuarina equisetifolia, Maesopsis eminii and Grevillea robusta grown in a linear simultaneous agroforestry system in Uganda. These species were established in single rows in the middle of fields in 1995 from four-month old seedlings. A total of 57 trees were sampled for this study, 13 for M. lutea, 12 for C. equisetifolia, 16 for M. eminii and 16 for G. robusta. Biomass values of the various tree components (stem, branches and foliage) as well as the total above-ground biomass were fitted to linear and non-linear allometric models using total height, diameter-at-breast height (DBH), crown width as predictor variables. Although both DBH and height are typically used as independent variables for predicting above-ground biomass, the addition of height in biomass equations did not significantly improve model performance for M. eminii, M. lutea and G. robusta. However, addition of height significantly increased the proportion of variation explained in above-ground biomass for C. equisetifolia, while DBH did not significantly improve the prediction of biomass. The study confirmed the need for developing species-specific biomass equations. [ABSTRACT FROM AUTHOR]

Routine monitoring of above ground biomass within purpose-grown willow biomass energy production systems is important for timing harvest and other operations to maximize profit and increase plantation productivity. The objective of this study was to assess the efficacy of an elegant nondestructive mensurative technique for providing reliable estimates of harvestable biomass for six willow varieties during a 3-year rotation. The LAI-2000 Plant Canopy Analyser was used to measure the stem area index of growing willow and relate it to harvestable biomass at four locations within Saskatchewan, Canada over a 3-year period. Given the highly significant relationship ( R = 0.95; P < 0.0001) between measured stem area index and harvestable willow biomass, independent of variety, age, or location, this simple mensurative technique is a promising alternative for estimating above ground biomass in short-rotation willow plantations. [ABSTRACT FROM AUTHOR]

In even-aged, 120-year-old Norway spruce stands with underplanted beech (in 1995) four permanent research plots were established, each 100 × 100 m in size. Twelve subplots were selected on each plot along a light gradient from complete canopy closure to open light conditions. On each plot, photon flux density (PFD) was measured continuously, and during same time interval (August 2008, 2009), color digital hemispherical photographs were taken. Cumulative PFD values for measuring points were compared with potential PFD radiation obtained from hemispherical photograph analysis for different angle of hemisphere and different parameters from hemispherical image analysis (gap fraction, total openness and direct, indirect and total amount of radiation). Cumulative and average daily values for the plots were compared; 120° hemispherical photograph angle, gap fraction and total openness were the variables that explained the largest proportion of variance in light transmittance. Determination coefficients between direct and total light component were highest for the total potential radiation and lowest for the diffuse light component. Comparison between potential and instantaneous light measurements for radial, height increment and SLA of young beech showed that instantaneous radiation measurements explained height increment best; 120° of hemisphere proved to be the best explaining angle. [ABSTRACT FROM AUTHOR]

Forest restoration projects have become increasingly common around the world and planting trees is almost always a key component. Low seedling survival and growth may result in restoration failures and various mechanical site preparation techniques for treatment of soils and vegetation are important tools used to help counteract this. In this article, we synthesize the current state-of-knowledge concerning mechanical site preparation for improved tree establishment when carried out in different forest restoration situations, point out critical research gaps and provide some recommendations for future directions. Mechanical site preparation often results in improved seedling survival and growth. However, if not intensive methods with much soil disturbance are used, it is a rather ineffective tool for controlling competing vegetation. Methods such as scarification, mounding and subsoiling also lead to multiple interactions among soil physical and chemical properties that affect plant survival and growth, and it may be difficult to determine the actual cause-effect relationship of any positive seedling responses. Most research to date on mechanical site preparation and plantation performance has been conducted using a few conifer tree species. Seedling responses differ among tree species and alternative species are often used during restoration compared to production forestry indicating a need for additional research for improved understanding. Several management objectives such as soil protection and increased biodiversity are many times relevant during forest restoration, and mechanical site preparation methods should be implemented carefully because they can have large impacts on the environment. [ABSTRACT FROM AUTHOR]

Seedling survival and successful forest restoration involves many silvicultural practices. One important aspect of a successful forest restoration program is planting quality seedlings with high survival capability. Thus the nursery needs to create seedlings with plant attributes that allow for the best chance of success once a seedling is field planted. Since the mid-twentieth century, research foresters have critically examined plant attributes that confer improved seedling survival to field site conditions. This review describes the value of commonly measured seedling quality material (i.e. shoot height, stem diameter, root mass, shoot to root ratio, drought resistance, mineral nutrient status) and performance (i.e. freezing tolerance and root growth) plant attributes defined as important in answering the question of why seedlings survive after planting. Desirable levels of these plant attributes can increase the speed with which seedlings overcome planting stress, become 'coupled' to the forest restoration site, thereby ensuring successful seedling establishment. Although planting seedlings with these desirable plant attributes does not guarantee high survival rates; planting seedlings with desirable plant attributes increases chances for survival after field planting. [ABSTRACT FROM AUTHOR]

We examined the effects of various mechanical site preparation methods and windrow burning on container-grown planted lodgepole pine ( Pinus contorta var . latifolia) survival and growth for 20 years after treatment at a sub-boreal site in north-central British Columbia, Canada. Survival was uniformly high (≥80%) regardless of treatment, indicating that site preparation was not necessary to establish pine on this site. Significant treatment effects on height, diameter, and stem volume were present at all assessment dates, but only the windrow burning treatment was associated with growth gains over the untreated control after two decades. Pine planted at the disk trench hinge were significantly larger than control pine only until year five. Of the mechanical treatments, only coarse mixing (by bedding plow) continued to have a significant effect on pine growth for as many as 9 years after treatment. Despite the disappearance of significant differences between mechanical treatments and the untreated control by year 20, the magnitude of stand volume increases suggests the potential for mechanical site preparation to have a beneficial effect on future timber supply. Repeated measures analysis confirmed that trends in early diameter growth differed between the untreated control and the windrow burning or coarse mixing treatments. These data are also potentially valuable for verifying growth and yield or carbon budgeting modelling tools. [ABSTRACT FROM AUTHOR]

We show the potential of a new method combining tree-ring analyses on stems and on coarse roots of individual trees in order to advance the understanding of growth dynamics in forest trees. To this end, we studied the root-shoot allometry of trees and its dependence on site conditions. Along a gradient in water supply in Southern Germany from dry to moist sites we selected 43 Norway spruce trees ( Picea abies [L.] H. Karst.) aged 65-100 years. Increment cores were taken from stem and main roots revealing aboveground and belowground growth course over the last 34 years. Annual growth rates in roots and stems and their allometric relationships were applied as surrogate variables for tree resource allocation to aboveground and belowground organs. The mean sensitivities of both stem and root chronologies were found to be site-specific, and increased from the moist through the dry sites. No temporal offset between aboveground and belowground growth reactions to climate conditions was found in Norway spruce at any of the sites. These results suggest that the root-shoot allometry depends on the specific site conditions only at the driest site, following the optimal biomass partitioning theory (the more restricted the water supply, the more organic matter allocation into the belowground organs). [ABSTRACT FROM AUTHOR]

Short-day treatment has been used for some conifer species to improve planting stock quality. The aim of this paper was to study the effect of short-day treatment on performance and quality of young seedlings of Italian cypress ( Cupressus sempervirens L.), a conifer species abundant in the Mediterranean area. Seeds of Italian cypress were sown in mini-plug cavities. After 5 weeks of growth, seedlings were subjected to four different photoperiods: (a) 14 h (control), (b) 8 h, (c) 6 h, and (d) 4 h for a period of 1-3 weeks, before they were transplanted to bigger size containers. Reducing photoperiod did not result in bud formation or shoot inhibition although it increased the number of lateral shoots. Seedlings subjected to 6-h photoperiod for 2 weeks during pre-cultivation in mini-plugs had improved root characteristics, giving them a higher potential for outplanting success, especially in Mediterranean areas. [ABSTRACT FROM AUTHOR]

Chlorophyll fluorescence, chlorophyll content, growth, and mortality of white spruce ( Picea glauca [Moench] Voss) seedlings were monitored for 2 years after planting under three scenarios of artificial shade: no-shade (control), shade in summer only, and shade all year. The shade frames allowed 50-60% light transmission, with limited effects on air temperature, relative humidity, soil temperature, and soil moisture around seedlings. Based on fluorescence yield and chlorophyll content measurements, summer-only shade reduced photoinhibition and photooxidation, especially in summer and fall; extending to all year shading did not further reduce either photoinhibition or photooxidation. Shade tended to reduce seedling diameter and mortality, but after 2 years the cumulative effect on mortality was not statistically significant. Study results support the establishment of white spruce seedlings under partial forest canopy, especially on sites with harsh environmental conditions. [ABSTRACT FROM AUTHOR]

In a reforestation field experiment undertaken in southern Sweden and planted with pedunculate oak, three site preparation treatments (herbicide application (H), mounding site preparation (MSP) and combined herbicide and mounding treatment (H + MSP)) were compared to an undisturbed control (C). We analysed root spatial and biomass distribution in 48 harvested seedlings. Compared to the control, both MSP and H treatments resulted in a significantly higher root system biomass (means C: 16.4 g, H: 45.7 g, MSP: 41.4 g, MSP + H: 102.2 g). The proportion of lateral root biomass increased from 38% (C) to 62% (MSP + H), while the two remaining treatments (H, MSP) attained percentages of 48% and 51%, respectively. The treatments did not alter seedlings’ allometry and root:shoot ratio significantly. However, biomass distribution of different root system parts varied among treatments, so that lateral root biomass increased relative to leaf biomass from C to MSP + H variant. Site preparation treatments (MSP, MSP + H) increased the horizontal extension of root biomass, compared to C and H variants. We conclude that mounding site preparation is an efficient method for the reforestation of pedunculate oak stands. Due to the comparative positive effects on root system development, MSP presents an alternative to herbicide treatment, particularly at sites where the use of chemicals is restricted. [ABSTRACT FROM AUTHOR]

Previous studies have indicated that recruitment of Abies sachalinensis, a representative conifer species of northern Japan, decreased following single-tree selection harvesting in stands with dense dwarf bamboo understory. We tested the hypothesis that growth and survival of A. sachalinensis seedlings are reduced by canopy opening in that type of stand. A 0.75 ha study plot was examined, and all the seedlings (defined as trees with height 0.5–2 m) were identified and their shoot extensions measured for three years after single-tree selection harvesting (26% intensity in terms of basal area). The leader extensions of A. sachalinensis seedlings that experienced canopy opening were greatly improved. However, a negative effect on survival was also apparent; nearly 40% of seedlings died at the sites that experienced canopy opening. These results were supported by generalized linear models that examined variations of local harvesting intensity for individual seedlings. Despite the ability of A. sachalinensis to respond rapidly to exposure, some physiological stresses may have appeared, and presumably were amplified by co-occurring dwarf bamboos. With regard to the low seedling density (156 stems ha−1), common in this type of stands, local harvesting intensity in the selection system should be reduced to maintain survival of advanced regeneration, thus sustaining stand structure and composition. [ABSTRACT FROM AUTHOR]

Tree growth and carbon dynamics are important issues especially in the context of climate change. However, we essentially lack knowledge about the effects on carbon dynamics especially in mixed stands. Thus, the objective of this study was to test the effects of climatic changes on the above and below ground carbon dynamics of a mixed stand of Norway spruce ( Picea abies [L.] Karst.) and European beech ( Fagus sylvatica L.) by means of scenario simulations. To account for the typical tree interactions in a mixed-species stand a spatial explicit tree growth model based on eco-physiological processes was applied. Three different climate scenarios considering altered precipitation, temperature, and radiation were calculated for an unthinned and a thinned stand. The results showed significant changes of above and belowground biomass over time, especially when temperature and radiation were increased additionally to decreased precipitation. The reduction in biomass increments of Norway spruce were more attenuated above than below ground. In contrast, the results for beech were the opposite: The belowground increments were reduced more. These results suggest a shift in the species contribution to above and belowground biomass under dryer and warmer conditions. Distinct effects were also found when thinned and unthinned stands were compared. A reduced stand density changed the proportions of above and below ground carbon allocation. As a main reason for the changed growth reactions the water balance of trees was identified which lead to changed biomass allocation pattern. [ABSTRACT FROM AUTHOR]

Seedling growth response of two tropical tree species ( Schima superba and Cryptocarya concinna) to simulated N deposition was studied during a period of 11 months. One-year-old seedlings were grown in forest soil treated with N as NH4NO3 at Control–no N addition, N5–5, N10–10, N15–15, and N30–30 g N m−2 year−1. The objective was to examine the effects of N addition on seedling growth and compare this effect between the two tropical tree species of different species-N-requirement. Results showed that both species responded significantly to N addition and exhibited positive effect to lower rate of N addition and negative effect to higher rate of N addition on growth parameters (height and stem base diameter, biomass production, and net photosynthetic rate). The highest values were observed in the N10 plots for S. superba and in the N15 plots for C. concinna, but the lowest values were observed in the N30 plots for both species. However, the reduction in the N30 plots was more pronounced for S. superba than for C. concinna relative to the control plots. Our findings suggest that response of seedling growth of tropical tree species to atmospheric N deposition may vary depending on rate of N deposition and species-N-requirement. [ABSTRACT FROM AUTHOR]

Impacts of nursery short-day treatments on the survival, growth and needle damage of about 5,000 1 + 0 container white spruce (Picea glauca [Moench] Voss) seedlings from a single seedlot were studied for two growing seasons following planting on July 22, 1999 at four boreal reforestation sites in Northern Alberta, Canada of varying soil texture, drainage, aspect, slope, and slope position. Each site was separated into two areas that were site-prepared by either ripping or mounding. When seedlings reached a height of about 20 cm under normal greenhouse growth conditions, the seedlings from different germination dates over a 7-week period were exposed to one of five different conditioning treatments (T), mainly through varying the duration of 12-h short-day exposure to 0 (T0), 3 (T3), 7 (T7), 10 (T10), or 15 (T15) days followed by different periods of reduced N supply. N-reduction produced few differences in needle nutrient concentrations and so was not considered a likely cause of differences in field performance. The treatments progressively (from T0 to T15) increased tolerance to drought and frost, and resulted in a similar seedling size for T3, T7 and T10 (planting height of 21 cm and ground diameter of 2.9 mm) although T0 seedlings were smaller (20 cm) and T15 taller (24 cm). The weather in 1999 was dry, particularly in the weeks immediately before and after planting, but relatively moist and favorable in 2000 and 2001 apart from one major frost event (-7°C) in May 2000. Survival, growth and needle damage varied substantially among sites and short-day treatments, and the treatment differences were largely consistent across the four sites. In general the growth was better on the ripped than on the mounded areas. Seedlings in T7 (intermediate tolerance) survived and grew best in the first year but T0 (actively growing) did best during the second year. After 2 years, no differences were observed among T0, T3 and T7 in mortality (18%) and total height growth (15 cm). However, T10 and T15 had higher mortality (24 and 43%), and lower height growth (12 and 10 cm, respectively). The percentage of seedlings showing very severe needle damage after 2 years increased from T0 through T15 (14-33%). New root egress in the field also differed significantly among treatments and was positively and highly correlated with survival rate and growth. These results suggest that a longer short-day regime, as in T10 or T15, may be counterproductive and a shorter regime may be more effective in improving the performance of summerplanted white spruce seedlings. [ABSTRACT FROM AUTHOR]

Manipulation of the canopy cover and site preparation are the most important silvicultural measures to enhance the conditions for natural regeneration of Norway spruce (Picea abies (L.) Karst.). During the early regeneration phase however, seedling mortality may be high, so it is important to study how different combinations of stand-level treatments and site preparation methods affect seedling establishment. We studied emergence, 1st winter and 2nd summer mortality for naturally regenerated spruce seedlings in a field experiment that combined four harvest treatments (shelterwoods of high (SH), medium (SM) and low (SL) residual basal area, and a 50 x 50 m clear-cut (CC)) and two site preparation methods (patch scarification and inverting). The CC had significantly fewer seedlings the 1st fall than the SL and SH (p = 0.0377), and in all harvest treatments, fewer seedlings emerged in inverted than in patch scarified spots (p = 0.0351). Mortality was also lower with patch scarification than inverting (1st winter: p = 0.0565, 2nd summer: p = 0.0377), but was not affected by harvest treatment (1st winter: p = 0.9211, 2nd summer: p = 0.1896). On average, mortality from 1st to 2nd fall reached 38% and 27% after inverting and patch scarification, respectively. First winter mortality accounted for approximately two thirds of the accumulated mortality, regardless of the harvest treatment and site preparation method. [ABSTRACT FROM AUTHOR]

Leaf area index (LAI) is a key parameter controlling plant productivity and biogeochemical fluxes between vegetation and the atmosphere. Tropical forests are thought to have comparably high LAIs; however, precise data are scarce and environmental controls of leaf area in tropical forests are not understood. We studied LAI and stand leaf biomass by optical and leaf mass-related approaches in five tropical montane forests along an elevational transect (1,050–3,060 m a.s.l.) in South Ecuador, and conducted a meta-analysis of LAI and leaf biomass data from tropical montane forests around the globe. Study aims were (1) to assess the applicability of indirect and direct approaches of LAI determination in tropical montane forests, (2) to analyze elevation effects on leaf area, leaf mass, SLA, and leaf lifespan, and (3) to assess the possible consequences of leaf area change with elevation for montane forest productivity. Indirect optical methods of LAI determination appeared to be less reliable in the complex canopies than direct leaf mass-related approaches based on litter trapping and a thorough analysis of leaf lifespan. LAI decreased by 40–60% between 1,000 and 3,000 m in the Ecuador transect and also in the pan-tropical data set. This decrease indicates that canopy carbon gain, that is, carbon source strength, decreases with elevation in tropical montane forests. Average SLA decreased from 88 to 61 cm2 g−1 whereas leaf lifespan increased from 16 to 25 mo between 1,050 and 3,060 m in the Ecuador transect. In contrast, stand leaf biomass was much less influenced by elevation. We conclude that elevation has a large influence not only on the leaf traits of trees but also on the LAI of tropical montane forests with soil N (nitrogen) supply presumably being the main controlling factor. [ABSTRACT FROM AUTHOR]

Reporting carbon (C) stocks in tree biomass (above- and belowground) to the United Nations Framework Convention on Climate Change (UNFCCC) should be transparent and verifiable. The development of nationally specific data is considered ‘good practice’ to assist in meeting these reporting requirements. From this study, biomass functions were developed for estimating above- and belowground C stock in a 19-year-old stand of Sitka spruce ( Picea sitchensis (Bong) Carr.). Our estimates were then tested against current default values used for reporting in Ireland and literature equations. Ten trees were destructively sampled to develop aboveground and tree component biomass equations. The roots were excavated and a root:shoot ( R) ratio developed to estimate belowground biomass. Application of the total aboveground biomass function yielded a C stock estimate for the stand of 74 tonnes C ha−1, with an uncertainty of 7%. The R ratio was determined to be 0.23, with an uncertainty of 10%. The C stock estimate of the belowground biomass component was then calculated to be 17 tonnes C ha−1, with an uncertainty of 12%. The equivalent C stock estimate from the biomass expansion factor (BEF) method, applying Ireland’s currently reported default values for BEF (inclusive of belowground biomass), wood density and C concentration and methods for estimating volume, was found to be 60 tonnes C ha−1, with an uncertainty of 26%. We found that volume tables, currently used for determining merchantable timber volume in Irish forestry conditions, underestimated volume since they did not extend to the yield of the forest under investigation. Mean stock values for belowground biomass compared well with that generated using published models. [ABSTRACT FROM AUTHOR]

Growth and yield modeling has a long history in forestry. The methods of measuring the growth of stand basal area have evolved from those developed in the U.S.A. and Germany during the last century. Stand basal area modeling has progressed rapidly since the first widely used model was published by the U.S. Forest Service. Over the years, a variety of models have been developed for predicting the growth and yield of uneven/even-aged stands using stand-level approaches. The modeling methodology has not only moved from an empirical approach to a more ecological process-based approach but also accommodated a variety of techniques such as: 1) simultaneous equation methods, 2) difference models, 3) artificial neural network techniques, 4) linear/nonlinear regression models, and 5) matrix models. Empirical models using statistical methods were developed to reproduce accurately and precisely field observations. In contrast, process models have a shorter history, developed originally as research and education tools with the aim of increasing the understanding of cause and effect relationships. Empirical and process models can be married into hybrid models in which the shortcomings of both component approaches can, to some extent, be overcome. Algebraic difference forms of stand basal area models which consist of stand age, stand density and site quality can fully describe stand growth dynamics. This paper reviews the current literature regarding stand basal area models, discusses the basic types of models and their merits and outlines recent progress in modeling growth and dynamics of stand basal area. Future trends involving algebraic difference forms, good fitting variables and model types into stand basal area modeling strategies are discussed. [ABSTRACT FROM AUTHOR]

Impacts of nursery cultural treatments (T) on stress tolerance of greenhouse-grown 1 + 0 container white spruce (Picea glauca [Moench] Voss) seedlings (mean height 24 cm, root collar diameter 3.1 mm) for summer planting were studied. Seedlings were subjected to 12-h short-day treatments of 0 (T0), 3 (T3), 7 (T7), 10 (T10), or 15 (T15) days, followed by 0, 7, 17, 40, or 46 days of reduced N supply, respectively. Relevant physiological and morphological factors were examined concurrently. Foliar N concentrations exceeded optimal levels and differed little among treatments, suggesting a minor confounding role for N reduction. Both frost and drought tolerance increased incrementally from T0 through T15. Electrolyte leakage index decreased steadily from T0 (25% for roots, 17% for needles) to T15 (1% for roots, 2% for needles) after 2-h exposure of fine roots to -2°C and of needles to -8°C. Withholding soil watering for 19 days caused 80% mortality among seedlings in T0, 50% in T3, and < 10% in T7-T15. The transpiration decline curve suggested that enhanced drought tolerance was largely attributable to quicker stomatal closure during water stress and lower cuticular transpiration rate. The treatments increased root growth capacity on a per-seedling, but not per-root-mass, basis. Needle primordia were developed in all T7-T15 seedlings but not in T0 and T3 treatments, suggesting that nurseries may need no more than 7 days of blackout application for conditioning spruce seedlings for summer planting. Shoot dry weight fraction increased gradually from T0 through T15 and was linearly correlated with needle specific weight and frost tolerance, and may thus be useful in monitoring progress of conditioning treatments. [ABSTRACT FROM AUTHOR]

Abstract. Seedlings grown under different N supply were examined for relationships between root system size attributes and its hydraulic properties. These relationships were also studied on seedlings of different stock types (grown in different container types). Measurements with root pressure probes were taken at various times after germination, under applied hydrostatic pressure and non-limiting soil moisture. Different N-treatments and stock types were used solely to produce seedling of different sizes, especially root system sizes. Specific root hydraulic conductivity (Lpr) typically declined with an increasing root system size and correlations between Lpr and the root system size attributes were often negative. The flow of water through the root system correlated well with root system size attributes only in young (3–4 month old) seedlings but the correlations were inconsistent among different N treatments and stock types. Neither the root system surface area nor dry weight reliably reflected its ability to absorb and conduct water. Generally, the amount of water delivered through the root system and available for transpiration per unit or leaf surface area or unit of leaf dry weight correlated poorly or negatively with the root system size. Practical and scientific implications of the findings are discussed. [ABSTRACT FROM AUTHOR]

Effects of water availability on seedling growth were analysed in eight Mediterranean species naturally occurring in the Balearic Islands. Seedlings were grown outdoors during summer under two irrigation treatments: field capacity and 35% of field capacity. The relative growth rate (RGR) strongly depended on the growth form, from highest values in herbs to lowest in woody perennials. The main component associated with interspecific variation in RGR was the specific leaf area (SLA), and a quantitative grouping of the different growth forms appeared along the regression line between both parameters. The slow-growing species, i.e. woody perennial shrubs, had the lowest SLA and the fast-growing perennial herbs, the highest, while woody semi-deciduous shrubs appeared intermediate. Decreases in RGR due to water stress were analysed in terms of the relative contribution of the leaf mass ratio (LMR), SLA and the net assimilation rate (NAR). Pooling all species, the decrease in RGR caused by water deficit was mainly explained by decreases in SLA. However, this general pattern was strongly dependent of growth form. Thus, in the woody perennial plants, the decrease in RGR was accompanied by a three-fold decrease in NAR which, however, increased in perennial herbs. SLA increased with decreasing water supply in woody perennial plants, and decreased in woody semi-deciduous shrubs and perennial herbs. Finally, decreases in LMR partly explained decreases in RGR in perennial herbs and woody perennial shrubs. This different response of the different growth forms may reflect differences in seedling adaptation and surviving strategies to drought periods. [ABSTRACT FROM AUTHOR]

This study investigated the belowground development and strategy of late-successional European beech ( Fagus sylvatica L.) in ageing natural Scots pine (Pinus sylvestris L.) and Silver birch (Betula pendula Roth.) woodlands in a French volcanic mid-elevation area. For this purpose root biomass, root profile and fine-root architecture of competitor trees were examined in 53 mixed pine–beech and 42 birch–beech woodlands along a stand maturation gradient, using the root auger technique (0–75-cm). The total beech fine-root biomass highly correlated with aerial dimensions such as stem height and girth, whereas it moderately correlated with its age, thus indicating the effects of competition. Basic stand biometric data such as stand density and basal area had no significant effect on beech root biomass. Conversely, competition indices taking into account the vertical dimensions of competitor trees were efficient, probably due to redundancy with beech height. At similar age and height, beeches under birch had a greater belowground development than beeches under pine. Each species exhibited specific rooting pattern and plasticity of fine-root architecture along the gradients of stand maturation and competition. Beech had a heart-shaped rooting habit in both mixings, which strongly increased along stand maturation. Its fine-root system adopted a foraging strategy to respond to increasing stand competition. The Scots pine fine-root system was plate-like and showed a low morphological plasticity, thus presumably a conservative strategy. Silver birch exhibited a high biomass and a foraging capacity in the topsoil but a loose root system in the subsoil. The coexistence of pine and beech roots in the upper soil presumably leads to a high belowground competition. Beech root system becomes predominant throughout the soil profile and it adopts an efficient foraging strategy, but at the expense of its belowground development. Conversely, the niche partitioning strategy between beech and birch may explain why beech develops strongly belowground in spite of the fact that birch has a dense rooting and a competitive fine-root architecture. As a consequence, beech mid-term regeneration and development may be facilitated under birch as compared with pine. [ABSTRACT FROM AUTHOR]

In a hypersaline mangrove scrub forest in northern Florida, coexisting trees of Laguncularia racemosa and Avicennia germinans were either fertilized with nitrogen or phosphorus, or not fertilized (controls). We aimed to test whether nutrient additions differentially altered photosynthetic performance and resource utilization in these two species. In control trees, photosynthetic rates were higher in L. racemosa than A. germinans. However, leaf nitrogen concentrations were higher in A. germinans than L. racemosa. Avicennia germinans responded to fertilization with nitrogen by increasing leaf nitrogen concentrations and rates of photosynthesis such that they were equivalent to photosynthesis in L. racemosa. Laguncularia racemosa did not show a response to nitrogen additions. Neither species showed strong responses to phosphorus fertilization. Avicennia germinans had high photosynthetic water-use efficiency (photosynthesis/transpiration), but low photosynthetic nitrogen-use efficiency (photosynthesis/leaf nitrogen). In contrast, L. racemosa had comparatively low photosynthetic water use efficiency and high photosynthetic nitrogen use efficiency. Leaf level characteristics lead us to hypothesize that coexistence of A. germinans and L. racemosa should occur where nitrogen levels are low and salinity is moderate, or at least moderate for some period of the year. [ABSTRACT FROM AUTHOR]

The split cuts of Cook, Kannan and Schrijver are general-purpose valid inequalities for integer programming which include a variety of other well-known cuts as special cases. To detect violated split cuts, one has to solve the associated separation problem. The complexity of split cut separation was recently cited as an open problem by Cornuéjols & Li CL01. In this paper we settle this question by proving strong &{scr;&{scr;𝒩𝒫&}scr;&}scr;-completeness of separation for split cuts. As a by-product we also show &{scr;&{scr;𝒩𝒫&}scr;&}scr;-completeness of separation for several other classes of inequalities, including the MIR-inequalities of Nemhauser and Wolsey and some new inequalities which we call balanced split cuts and binary split cuts. We also strengthen &{scr;&{scr;𝒩𝒫&}scr;&}scr;-completeness results of Caprara & Fischetti CF96 (for [ABSTRACT FROM AUTHOR]

Provides a summary and synthesis of the results from the Gap Model Comparison Workshop that was held in July 1999 at Pingree Park, Colorado. How to scale individual plant responses to the population level; Results of a number of model comparison exercises; Information on regeneration and mortality mechanisms.

Paeonia ostii is a traditional ornamental and medicinal species that has attracted considerable interest for its high oil value. To facilitate the effective and rational cultivation and application of P. ostii in China, it is necessary to determine its potential spatial habitat distribution and environmental requirements. Using high-resolution environmental data for current and future climate scenarios, the potential suitable area and climatic requirements of P. ostii were modelled. Among the 11 environmental variables investigated, growing degree days, precipitation of the wettest month, mean temperature of the coldest quarter, global UV-B radiation, annual precipitation, and soil pH played major roles in determining the suitability of a habitat for the cultivation of P. ostii. Under the current environmental conditions in China, a total area of 20.31 × 105 km2 is suitable for growing P. ostii, accounting for 21.16% of the country's total land area. Under the two future climate scenario/year combinations (i.e., representative concentration pathways [RCPs], RCP2.6 and RCP8.5 in 2050), this species would increase its suitable area at high latitudes while decrease at low latitudes. These results present valuable information and a theoretical reference point for identifying the suitable cultivation areas of P. ostii. [ABSTRACT FROM AUTHOR]

Two trials ('Wonowon' and 'Iron Creek') in the Prince George Forest Region of interior British Columbia were begun in the mid 1980s to evaluate site preparation treatments for establishing white spruce (Picea glauca [Moench] Voss) in the Boreal White and Black Spruce biogeoclimatic zone. The 14 treatments (9 or 10 per trial) were: [B.C.] Ministry, Sinkkila, and Bracke mounds; Bracke mounds manually supplemented with 20-, 14-, or 6-cm cappings of mineral soil; fertilized Sinkkila mounds; Bracke patches; fertilized Bracke patches; bladed strips; plowed ground; herbicide; and untreated controls, separately with bothstandard and nominally superior 'alternate' planting stock. With minor aberrations, each trial consists of 5 randomized complete blocks each with one 80-tree plot per treatment; planting was in spring, 1984at Wonowon, 1987 at Iron Creek. All trees in mounding treatments andthe inner 48 trees in other plots were monitored for performance through 1998 at Wonowon, 1996 at Iron Creek. The herbicide and plowing treatments, and mounds capped thickly enough with mineral soil to inhibit weed regrowth, were clearly superior to others. In the mounding treatments at Wonowon, survival rate increased with capping thickness,but, while the 14 and 20 cm cappings were best, significant differences among them were few after 15 growing seasons. In both trials, patch scarification gave poorer results than did planting without site preparation. [ABSTRACT FROM AUTHOR]

This study examined how the availability of inorganic nitrogen (N) modified the anatomical characteristics of white spruce (Picea glauca (Moench) Voss) roots related to their hydraulic properties. Seedlingswere grown for one growing season in 4 L capacity pots filled with sand under one of three N levels: low (10 ppm), medium (50 ppm) and high (125 ppm). First order lateral roots with intact tips were sampledfrom dormant seedlings in October. Root segments were collected from4, 10, and 14 cm distances above the root tip for fixation and sectioning and for maceration. Additional specimens were collected from the 4 and 14 cm distances for maceration and scanning electron microscopy of xylem pits. Root diameter and surface area occupied by the xylem in root cross sections increased basipetally in all treatments but exceptions were found. Higher N-levels significantly increased root diameter and surface area occupied by the xylem. In the two higher N treatments secondary root development was more advanced near the root tip than in the low N treatment. There was a strong positive correlation between root diameter and cross-sectional root area occupied by the xylem (30-50% of the root cross section) but not in portions with little secondary development. Non-conducting space within the xylem occupied 10-13% of its cross-sectional surface. Tracheids of the primary xylem were larger, had larger lumens but thinner cell walls than those of the secondary xylem. Low N treatment seedling tracheids had smaller total cross-sectional area, less lumen, and less cell wall surface area than the two other N treatments. Tracheid diameter means were between 19-20 Mum in the high and medium N treatments, and 15.2 ??m in the low N treatment. The range was 4.5-51.3 Mum. Tracheid lengthwas not significantly affected by N. The average tracheid was about 1000 Mum long, and the range was 110-3530 Mum. Pit-border diameters ranged between 4.1-20.6 Mum (average 10-11 Mum) and were not affected by t [ABSTRACT FROM AUTHOR]