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Muon spin resonance and 7Li NMR relaxation measurements in frustrated two-dimensional S = 1/2 Heisenberg antiferromagnets on a square lattice are presented. It is found that, in both Li2VOSiO4 and Li2VOGeO4, spin dynamics at frequencies orders of magnitude below the Heisenberg exchange frequency are present. These dynamics are associated with the motions of walls separating coexisting collinear domains with a magnetic wave vector rotated by 90 degrees.