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The interaction between the natural ambient winds found in the atmospheric boundary layer (ABL) and buoyant flames is crucial in broad applications in the scientific and engineering fields. Unlike the buoyancy-driven pool fires in still air that have been studied extensively, the complexity of physics changes significantly in wind presence. This study aims at analysing the characteristics of boundary-layer turbulence in the presence of large fires. The eddy dissipation concept, finite volume discrete ordinate method, and one k-equation model are used for combustion, thermal radiation, and sub-grid scale closure using the Large-Eddy Simulation (LES), respectively. A numerical model on simple cases is validated first to assess its capability to reproduce available experimental observations for a purely buoyant fire in still air. A forced-flow boundary layer combustion in a small chamber with a smooth inflow is further considered. In general, good agreement between the simulation results and available experimental data was achieved for temperature and velocity profiles. The unsteady inflow condition used to consider incoming atmospheric turbulence is generated through a precursor simulation. The wind interaction with the line fire changes the atmospheric boundary layer profile affecting the heat transfer ahead of the flame, thereby creating counter-rotating structures downstream. It is shown that the buoyancy-dominated flow due to the flame reaction induced local pressure variation and perturbed shear flow near the ground, thereby altering the wind speed through which the plume rises. Richardson number was also used as a dominant non-dimensional group to analyse the variation of enhanced flow vertical velocity with distance from the fire source. Thus, it is understood that the pronounced longitudinal shear spreading at the surface affects the behaviour of short term or puff releases, suggesting the shedding of small eddies during the combustion process.