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Central European weather and climate are closely related to atmospheric mass advection triggered by the North Atlantic Oscillation (NAO), which is a relevant index for quantifying internal climate variability on multi-annual timescales. It remains unclear, however, how large-scale circulation variability affects local climate characteristics when downscaled using a regional climate model. In this study, 50 members of a single-model initial-condition large ensemble (LE) of a nested regional climate model are analyzed for a NAO–climate relationship. The overall goal of the study is to assess whether the range of NAO internal variability is represented consistently between the driving global climate model (GCM; the Canadian Earth System Model version 2 – CanESM2) and the nested regional climate model (RCM; the Canadian Regional Climate Model version 5 – CRCM5). Responses of mean surface air temperature and total precipitation to changes in the NAO index value are examined in a central European domain in both CanESM2-LE and CRCM5-LE via Pearson correlation coefficients and the change per unit index change for historical (1981–2010) and future (2070–2099) winters. Results show that statistically robust NAO patterns are found in the CanESM2-LE under current forcing conditions. NAO flow pattern reproductions in the CanESM2-LE trigger responses in the high-resolution CRCM5-LE that are comparable to reanalysis data. NAO–response relationships weaken in the future period, but their inter-member spread shows no significant change. The results stress the value of single-model ensembles for the evaluation of internal variability by pointing out the large differences of NAO–response relationships among individual members. They also strengthen the validity of the nested ensemble for further impact modeling using RCM data only, since important large-scale teleconnections present in the driving data propagate properly to the fine-scale dynamics in the RCM. [ABSTRACT FROM AUTHOR]