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We report on a 60-year-old red pine (Pinus resinosa Aiton) spacing trial experiment located in Ontario, Canada, that included the combinations between six initial spacings (from 1.2 to 3.0 m) and the presence or absence of a commercial thinning (CT) regime, as well as their impacts on quadratic mean diameter (QMD) and stand volume yield. The CT regime, initiated at age 30, targeted a residual basal area (BA) of 38 m2·ha−1after each of four entries. Without thinning, as initial spacing increased, QMD increased; gross and net volume production peaked in the 2.1–2.4 m spacings. With thinning, similar trends with spacing were evident for QMD, although piece sizes were larger and differences between spacings were lower. The immediate increase of mean tree size caused by tree selection explained most of the differences in QMD between thinned and unthinned plots. Thinning to a common target BA resulted in similar standing volume across spacings. Cumulative gross yield was similar between spacings of

Knowledge of the long-term effects of silviculture treatments is crucial to forest management. The long-term effects of thinning, a common and widely used silviculture treatment, is little documented for upland black spruce ( Picea mariana (Mill.) Britton, Sterns & Poggenb.) stands. We revisited a partial cutting experiment installed in 1961 in a 65-year-old unmanaged upland black spruce stand. The aim was to document the long-term effects of thinning on tree and stand growth and to complete previously published results of the first 15 years of response to thinning by determining its influence in terms of merchantable volume. Free thinning was applied following three intensities: 0%, 25%, and 50% of total basal area removal. The retrospective analysis of growth rings showed that the response over the first 15 years was less significant when determined in net merchantable volume instead of net total volume. Heavily thinned plots, nonetheless, showed a net stand merchantable volume increment 33% greater than that of the unthinned plots. In the longer term, a spruce budworm ( Choristoneura fumiferana Clemens) outbreak affected the site; nevertheless, the heavily thinned plots maintained a superior tree growth rate and did not show senescence mortality like the other plots, allowing stand volume to catch up to that of the unthinned plots after 33 years. Results suggested that thinning upland black spruce stands may be useful in mitigating reductions in volume production associated with growing stands to longer rotations as called for by certain ecosystem-based management approaches.

Regeneration of northern white-cedar ( Thuja occidentalis L.) is often deficient after harvesting in mixedwood stands growing on mesic sites even where browsing pressure is low. We compared the effectiveness of silviculture treatments on early regeneration of white-cedar after single-tree selection cutting (25% of basal area removed), shelterwood seed cut (50% of basal area removed), and group selection cutting (gaps of 625 m2) in three yellow birch ( Betula alleghaniensis Britt.) – softwood uneven-aged stands in Quebec, Canada. Three years after harvesting, the combination of factors that maximized abundance of white-cedar seedlings was single-tree selection cutting with artificial seeding on exposed mineral seedbeds (68.8% of plots with the presence of white-cedar). Early growth of planted white-cedar seedlings (40 cm tall) and biomass production were proportional to light availability, i.e., best under group selection cutting (mean height increment = 14.8 cm/year, mean root collar diameter increment = 3.0 mm/year). Browsing pressure has regional impacts depending on herbivore occupancy of the area. Our study was conducted near the northern limit of white-tailed deer ( Odocoileus virginianus (Zimmerman, 1780)), and deer were locally absent, while snowshoe hare ( Lepus americanus Erxleben, 1777) did not have a consistent effect on seedling abundance and early survival, limiting height growth only during the first year following planting.

We present the 6 year effects of different cutting patterns (patch-selection cutting with 20, 30, and 40 m diameter gaps, 1 ha patch clear-cut, and uncut control) and spot scarification, on seedbed coverage and regeneration dynamics in yellow birch ( Betula alleghaniensis Britton) – conifer stands in eastern Quebec, Canada. After 3 years, yellow birch had established better in cutting patterns with gaps than in the patch clear-cut and in the control, while its density was 7 times higher in scarified than in nonscarified subplots. After 6 years, scarified openings and the borders of openings had 3–5 times more seedlings >30 cm in height than nonscarified openings and the understory between the gaps. The loss of advance growth in openings was the main result for conifer species, although recruitment of new balsam fir ( Abies balsamea (L.) Mill.) seedlings was accelerated by scarification. Despite the abundance of red spruce ( Picea rubens Sarg.) seed-trees on the site, our treatment combinations failed to promote its natural regeneration. Varying gap size did not change the total density of competing vegetation but modified the composition of this shrub layer. Our 6 year results suggest that maintaining conifer species, and the mixed composition of the stands, is uncertain over the long term.

This paper examined the impact of commercial thinning on selected wood properties of jack pine ( Pinus banksiana Lamb.). Wood properties evaluated include wood density (ring density, earlywood ring density, and latewood ring density), percentage of latewood in the ring, and wood bending properties. Nonlinear, mixed-effect models have been developed using data from three commercially thinned sites in eastern Canada. Ring density followed the same pattern as percentage of latewood, in which cambial age, relative height, and ring width were found to have important effects. Earlywood and latewood ring densities changed within the juvenile wood zone until a plateau was reached. Ring width affected earlywood and latewood ring densities mainly in narrow rings. Wood bending stiffness (measured by modulus of elasticity) and strength (measured by modulus of rupture) increased with cambial age and wood density; whereas, wood strength was also affected by ring width. Commercial thinning did not influence the developed models, but it had an indirect effect through increased ring width.

The mortality pattern of 14 black spruce (Picea mariana (Mill.) BSP) stands from the Saguenay region (Quebec), originating from fire or clear-cutting in the early 20th century, was reconstructed based on the dendrochronological dating of dead trees. Most of the spruces died during the 1970s and 1980s, which suggests the possible impact of the most recent spruce budworm (Choristoneura fumiferana Clem.) outbreak. This hypothesis is supported by the observation of major growth reductions synchronous with known outbreak periods and by the existence of a significant partial correlation between the mortality rates and the occurrence of outbreaks. A significant partial correlation between the mortality rates and the relative density of the studied stands suggests that competition amongst individuals might act as a predisposing factor to the death of trees, while the outbreaks act as an inciting factor. The observed mortality pattern is similar to self-thinning, since most of the dead trees were from the smallest size classes.

The widespread use of careful logging in the province of Quebec raises many questions about the effects of this practice on the structure and productivity of boreal black spruce (Picea mariana (Mill.) BSP) stands compared with fire-origin stands that are currently being harvested. The objective of this study is to describe and compare the evolution pattern of the diameter structure of stands originating from fire or logging dating back to the first half of the 20th century. The dendrochronological approach allowed the reconstitution of the dynamics of 40- to 96-year-old stands. Second-growth stands differ from fire-origin stands by their highly hierarchized initial structure. After the recruitment period, the degree of hierarchy of the structure of second-growth and fire-origin stands is comparable. After logging, the diameter structure of black spruce stands keeps a higher skewness index than that of stands originating from fire. For both stand types the skewness decreases or remains constant with time, which invalidates the model proposed by Mohler et al. (C.L. Mohler, P.L. Marks, and D.G. Sprugel. 1978. J. Ecol. 66: 599–614). This observation questions the existence of a strong asymmetric competition among trees in black spruce stands. Results indicate that the yield of black spruce stands originating from clearcuts from the early 20th century is higher than or equal to fire-origin stands mainly because (i) basal area was maintained to levels higher than or equal to fire-origin stands and (ii) most advanced growth was taller that 1 m at the time of logging.

Analysis was performed on 112 stems of black spruce (Piceamariana (Mill.) BSP) from the Réserve faunique des Laurentides to compare the growth in height, DBH, and volume of layers released by clear-cutting to the growth of individuals of comparable age originating from seeds after fire. The sampled stands originated from fires and clear-cuttings that occurred between 1894 and 1941. Stem analysis also permitted the calculation of specific volume increment, which corresponds to the annual volume increment divided by the surface of the cambium. Results show that height, DBH, and volume measured 60 years after clear-cutting were better correlated to the height of the advanced growth at the moment of release (Ho) than to the number of years of suppression. The mean annual increments in height and DBH of released layers were positively related to height at the year of logging when the height was less than 2 m. When layers were taller, mean annual increments in height and DBH were negatively related to initial height. These relationships were however variable, since for two-thirds of the stand's life, the periodic annual increments in height and diameter did not differ significantly (α = 0.05) between small (Ho o ≤ 2 m), and large second-growth spruces (Ho > 2 m). The superiority of taller layers is consequently due to greater heights before release. Volume growth rate of layers was positively related to initial height until approximately 60 years after clear-cutting. However, no differences in specific volume increment could be associated with initial height. Therefore, the relationship between initial height and volume increment can be attributed to the difference of cambial area between small, medium, and large second-growth spruces. Black spruce originating from seeds had greater height, DBH, and specific volume increments than second-growth trees, until about 50 years after stand origin. Thereafter, growth rates are comparable. Consequently, after 40 years, stands originating from seeds are comparable in height, volume, and DBH to layers that reached between 1 and 2 m in height after clear-cutting. Neither drainage class nor point density had a significant effect on the sampled black spruces. Stem analysis also revealed an important growth reduction that can be associated with the last spruce budworm (Choristoneurafumiferana Clem.) outbreak in the Réserve faunique des Laurentides. We can deduce from the results of this study that the performance of second-growth stands compared with fire-origin stands will mainly depend on the density of the advanced growth and its height structure.