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• Evaporative fraction increased by 16–49 % on cloudy days compared with clear days. • Clearness index controlled evaporative fraction through direct and indirect paths. • High NDVI reduced evaporative fraction response to cloudiness. Cloudiness influences gross primary productivity and evapotranspiration in terrestrial ecosystems by increasing diffuse radiation and improving the photosynthesis of shaded leaves. However, the effect of cloudiness on energy partitioning in terrestrial ecosystems is not well understood. Using latent heat (LE) and sensible heat (H) datasets observed by the eddy covariance systems and meteorological variables, the regulation of biophysical factors on energy partitioning under different sky conditions was investigated at 6 plantation ecosystems (2 deciduous broadleaf and 4 coniferous plantations) in eastern China from 2019 to 2021. The averages of evaporative fraction (EF) during the growing season were 0.34±0.06–0.50±0.04. The EF values increased by 16−49% on cloudy days, which was attributed to the lower decrease in LE compared to H. On the half-hourly scale, EF was mainly driven by air temperature (T a), relative extractable soil water (REW) and clearness index (CI) (i.e., a metric to quantify cloudiness). For plantation ecosystems, CI had an indirect effect on EF mainly through net radiation (R n). Moreover, the positive effect of cloudy sky conditions on EF was higher under low T a , vapor pressure deficit (VPD) and REW conditions. The increment of EF on cloudy days was weaker at high normalized difference vegetation index, which may be explained by higher T a and VPD limiting canopy transpiration during the mid-growing season. Cloudiness can also influence energy partitioning by regulating stomatal behavior in plantation ecosystems. These results contribute to understanding the drivers of energy partitioning in response to cloudiness in plantation ecosystems. [ABSTRACT FROM AUTHOR]