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As zoonotic diseases coevolved with early agriculture, social distancing within dense human settlements could have conferred a selective advantage in terms of infection risk. Here, we consider the case of Trypillia mega-settlements after 4000 BC, as virulent diseases began affecting humans in the Black Sea region. Through epidemiological susceptible-infected-recovered-susceptible (SIRS) models situated on clustered networks and on a site plan of a Trypillia mega-settlement, we show the adaptive benefits of decreasing either occupation density or the frequency of interactions with other communities across the settlement. We explore critical thresholds in these parameters that may shed light on the fluctuations of population densities at Trypillia mega-settlements before and after approximately 3600 BCE. Our findings suggest that disease was probably a significant driver of human settlement patterns by late Neolithic times.

A variable proportion of finds from the Neolithic and Chalcolithic of ‘Old Europe’ has come from places outside settlements, cemeteries, production sites, ritual sites, or caves. Such finds tend to be described as ‘chance/isolated/single/stray’ finds or, when in groups, as ‘hoards’. The frequent, modernist cause invoked for these finds is that they were either ‘hidden’ in times of mortal danger, represented a ‘gift to the gods’, or simply ‘lost’. One reason for these explanatory shortcomings is the over-attention to the types of objects deposited in the landscape and the frequent lack of attention to the often-distinctive place of deposition. We believe that we have misnamed, overlooked, or not accurately characterised an entire class of sites, which we term ‘landscape deposition sites’, whose defining feature was the transformation of a place by the deposition of a significant object or group of objects to create a qualitatively different place. The creation of such landscape deposit sites varied in time and space throughout Old Europe, but all sites were affected by this new dimension of the extended cultural domain.In this article, we consider the interpretations of metal deposition in North-west Europe and the light they shed on an earlier and geographically different region. The primary aim of this paper is an exploration of the variable relationships between landscape deposit sites and the coeval finds made in special deposits in settlements and cemeteries in the 5th and 4th millennia bc, which will lead to proposed new interpretations of landscape deposition sites.

AbstractThe Varna I cemetery, on the Bulgarian Black Sea coast, is one of the most remarkable sites in European prehistory, with the world’s earliest large-scale assemblage of gold artifacts. Modeling of the first series of 14 accelerator mass spectrometry (AMS) dates yielded a duration of occupation at the site of ca. 150 years, ~4600–4450 cal BC. However, there were insufficient paired human–animal dates for a full consideration of the question of the marine reservoir effect. Here, a fuller set of 71 dates from 53 graves is presented. We identify a small reservoir effect in a number of individuals based on 14C, as well as carbon and nitrogen stable isotopes. We test the effect of this by building a series of different Bayesian models. Our favored model, including a correction for some of the human determinations, shows activity at the cemetery starting at 4596–4516 cal BC and ending 4427–4341 cal BC (at 95.4% probability). The overall span of activity covers ~120–260 years (93.6% prob.). The modeling shows that Varna I falls toward the beginning of the Bulgarian Late Copper Age.

Farming was first introduced to Europe in the mid-seventh millennium bc, and was associated with migrants from Anatolia who settled in the southeast before spreading throughout Europe. Here, to understand the dynamics of this process, we analysed genome-wide ancient DNA data from 225 individuals who lived in southeastern Europe and surrounding regions between 12000 and 500 bc. We document a west–east cline of ancestry in indigenous hunter-gatherers and, in eastern Europe, the early stages in the formation of Bronze Age steppe ancestry. We show that the first farmers of northern and western Europe dispersed through southeastern Europe with limited hunter-gatherer admixture, but that some early groups in the southeast mixed extensively with hunter-gatherers without the sex-biased admixture that prevailed later in the north and west. We also show that southeastern Europe continued to be a nexus between east and west after the arrival of farmers, with intermittent genetic contact with steppe populations occurring up to 2,000 years earlier than the migrations from the steppe that ultimately replaced much of the population of northern Europe.