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The invasive ricaniid species Pochazia shantungensis (Chou & Lu, 1977), an Asiatic pest in China and Korea, is reported for the first time in Italy. This is the fourth confirmed record from Europe after Turkey, France, and Germany. The list of the host plants recorded in Europe is given. Illustrations of male and female genital structures (external and internal) are also presented.

The paper provides new distributional data for 5 planthopper and 61 leafhopper species, recorded from the Bieszczady Mountains (south-eastern Poland), including 35 taxa listed as new for the region.

A new monospecific genus of flatid planthoppers (Hemiptera: Fulgoromorpha: Flatidae), Makaya gen. nov., is described for Makaya volontany sp. nov. (type species) from the island of Madagascar. Habitus, male and female external and internal genital structures of the new species are illustrated and compared with related taxa. Makaya volontany is endemic to Madagascar where it is currently known from different types of dry forests in the western part of the island.

Sap-feeding hemipteran insects live in associations with diverse heritable symbiotic microorganisms (bacteria and fungi) that provide essential nutrients deficient in their hosts’ diets. These symbionts typically reside in highly specialized organs called bacteriomes (with bacterial symbionts) or mycetomes (with fungal symbionts). The organization of these organs varies between insect clades that are ancestrally associated with different microbes. As these symbioses evolve and additional microorganisms complement or replace the ancient associates, the organization of the symbiont-containing tissue becomes even more variable. Planthoppers (Hemiptera: Fulgoromorpha) are ancestrally associated with bacterial symbionts Sulcia and Vidania, but in many of the planthopper lineages, these symbionts are now accompanied or have been replaced by other heritable bacteria (e.g., Sodalis, Arsenophonus, Purcelliella) or fungi. We know the identity of many of these microbes, but the symbiont distribution within the host tissues and the bacteriome organization have not been systematically studied using modern microscopy techniques. Here, we combine light, fluorescence, and transmission electron microscopy with phylogenomic data to compare symbiont tissue distributions and the bacteriome organization across planthoppers representing 15 families. We identify and describe seven primary types of symbiont localization and seven types of the organization of the bacteriome. We show that Sulcia and Vidania, when present, usually occupy distinct bacteriomes distributed within the body cavity. The more recently acquired gammaproteobacterial and fungal symbionts generally occupy separate groups of cells organized into distinct bacteriomes or mycetomes, distinct from those with Sulcia and Vidania. They can also be localized in the cytoplasm of fat body cells. Alphaproteobacterial symbionts colonize a wider range of host body habitats: Asaia-like symbionts often colonize the host gut lumen, whereas Wolbachia and Rickettsia are usually scattered across insect tissues and cell types, including cells containing other symbionts, bacteriome sheath, fat body cells, gut epithelium, as well as hemolymph. However, there are exceptions, including Gammaproteobacteria that share bacteriome with Vidania, or Alphaproteobacteria that colonize Sulcia cells. We discuss how planthopper symbiont localization correlates with their acquisition and replacement patterns and the symbionts’ likely functions. We also discuss the evolutionary consequences, constraints, and significance of these findings.

We study the classification problem of simple transitive 2-representations of the 2-category of finite-dimensional bimodules over a radical square zero Nakayama algebra. This results in a complete classification of simple transitive 2-representations whose apex is a finitary two-sided cell. We define a notion of localization of 2-representations. We construct previously unknown simple transitive 2-representations as localizations of cell 2-representations. Using the universal property of our construction we prove that any simple transitive 2-representation with finitary apex is equivalent to a localization of a cell 2-representation., Comment: 24 pages

SummaryThe evolutionary success of sap-feeding hemipteran insects in the suborder Auchenorrhyncha was enabled by nutritional contributions from their heritable endosymbiotic bacteria. However, the symbiont diversity, functions, and evolutionary origins in this large insect group have not been broadly characterized using genomic tools. In particular, the origins and relationships among ancient betaproteobacterial symbiontsVidania(in Fulgoromorpha) andNasuia/Zinderia(in Cicadomorpha) are uncertain. Here, we characterized the genomes ofVidaniaandSulciafrom threePyropsplanthoppers (family Fulgoridae) to understand their metabolic functions and evolutionary histories. Like in previously characterized planthoppers, these symbionts share nutritional responsibilities, withVidaniaproviding seven out of ten essential amino acids.Sulcialineages across the Auchenorrhyncha have a highly conserved genome but with multiple independent rearrangements occurring in an early ancestor of Cicadomorpha or Fulgoromorpha and in a few succeeding lineages. Genomic synteny was also observed within each of the betaproteobacterial symbiont generaNasuia,Zinderia, andVidania, but not across them, which challenges the expectation of a shared ancestry for these symbionts. The further comparison of other biological traits strongly suggests an independent origin ofVidaniaearly in the planthopper evolution and possibly ofNasuiaandZinderiain their respective host lineages.Originality-Significance StatementWe sequenced and characterized the genomes of two ancient nutritional symbionts,SulciaandVidania, in three species from the genusPyropsin the species- and symbiont-rich but understudied insect clade, Fulgoromorpha (planthoppers). We describe—for the first time—several independent genome rearrangements inSulcia, which is often cited as a premier example of extreme genome stability spanning hundreds of millions of years. We also show a global lack of synteny across the genomes of the Auchenorrhynchan betaproteobacterial symbionts (Vidania,Nasuia, andZinderia). This result is unexpected given previous hypotheses of a common origin for these symbionts >250 million years ago alongsideSulcia. Taken together, we suggest an independent origin ofVidaniaand possibly ofNasuiaandZinderiasymbiont lineages as well. This hypothesis further links the potential acquisition of novel nutritional endosymbiont lineages with the emergence of auchenorrhyncham superfamilies.

A new monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae), Tarehylava gen. nov., is described for T. avymaina sp. nov. (type species), based on four females collected in the spiny forest ecoregion of south-western Madagascar. Habitus, as well as external and internal female genital structures of the new genus and species are described and illustrated.

Stroiński, Adam, Balderi, Matteo, Marraccini, Daniele, Mazza, Giuseppe (2022): First records of Pochazia shantungensis (Chou & Lu, 1977) (Hemiptera: Fulgoromorpha: Ricaniidae) in Italy. Zootaxa 5188 (3): 275-282, DOI: https://doi.org/10.11646/zootaxa.5188.3.4

A new monospecific genus of flatid planthoppers (Hemiptera: Fulgoromorpha: Flatidae), Avaratra gen. nov., is described for Avaratra spinifemora sp. nov. (type species) from the island of Madagascar. Habitus, male and female external and internal genital structures of the new species are illustrated. Avaratra spinifemora gen. et sp. nov. is endemic to Madagascar and probably represent a species specialized to dry costal forest, the vegetation widely distributed in Madagascar, gradually extending with the climate change. Taking into account a few data and poor collecting in these areas, particularly for planthoppers, it is possible that the species occurs in other similar places along the west northern coast.

Laberia palliata Stål, 1866 Figs 3–16 Perinetia reticulata Lallemand & Synave, 1954: 82; syn. nov. Aylaella reticulata – Demir & Özdikmen 2009: 271. Type material Holotype of Laberia palliata Stål, 1866 (Fig. 3) LOCALITY UNKNOWN • ♀; “ Mauri- / tius.”, “ Stevens.”, “Typus”, “140 / 64”, “ Laberia pallida Stål / Holotype (Flatidae)”, “ NHRS-GULI / 000006158 ”; NHRS. Holotype of Perinetia reticulata Lallemand & Synave, 1954 (Fig. 4) MADAGASCAR • ♂; “MADAGASCAR / Rég. Périnet / G. Olsoutiett col. 193”, “TYPE”, “ V. Lallemand et / H. Synave det., 195? / Perinetia g. n. / reticulata sp. n. ”, “Museum Paris / MNHN (EH) / 19341”; MNHN. Additional material examined MADAGASCAR – Alaotra-Mangoro Region, Moramanga District • 1 ♂; “ Analamazotra / Perinet, Madag / OЛСуФЪЕва XI.930”; ZIN • 1 ♀; “Périnet / Madagascar / Seyrig 12.II. 931”; ZIN • 1 ♀; “ Madagascar / province de Toamasina / Andasibe, 1049 m / S18°53,410, E48°22,881 ”, “Museum Paris / frt humid, brd piste nicke / 04-XI-2011 / Th. Bourgoin Rèc. ”, “Museum Paris / MNHN (EH) / 19347”; MNHN • 1 ♂, 1 ♀; “ Madagascar –CE / border of Andasibe N.P. / “Parc de Orchidés”/ at light; 3.-6. xi.2010 / P. Bănař & local coll. lgt.”, “Collectio / Moravské museum / Brno”; MMBC. – Analamanga Region, Manjakandriana District • 1 ♀; “ MADAGASCAR CENTRE / EST LAC MANTASOA / AMBOHIBOATAVO ”, “ III-1973 / A. Peyrieras ”, “Museum Paris / MNHN (EH) / 16576”; MNHN • 1 ♀; “ MADAGASCAR CENTRE / EST LAC MANTASOA / AMBOHIBOATAVO ”, “ III-1973 / A. Peyrieras ”, “Museum Paris / MNHN (EH) / 16577”; MNHN. – Vatovavy Fitovinany Region, Ifanadiana District • 1 ♂; “ Madagascar –CE / Ranomafana N.P.; 11.– / 18.xi. 2010, 958m / S21°15′22.6″ E47°25′17.8″ / at light; P. Bănař lgt.”, “Collectio / Moravské museum / Brno”; MMBC. – Vatovavy Fitovinany Region: Ifanadiana District, Ranomafana commune • 1 ♀; “COLL. MUS. CONGO / Madagascar: Nandihizina / XII – 1937, J. Vadon ”, “ Laberia paliata Stål, det / RGFennah”; MRAC. Redescription MEASUREMENT. Total length 15–18 mm. HEAD. Head with compound eyes narrower than pronotum (Fig. 6A–B). Disc of vertex, pronotum and mesonotum in one plane (Fig. 7A). Vertex (Fig. 6A–D): ratio A/B = 1.0–1.1, with margins slightly elevated, carinate; anterior margin of vertex triangularly produced, apex of vertex obtusely angled; posterolateral margins of vertex slightly diverging posteriad; posterior margin distinctly triangularly incised, to the level of anterior margins of compound eyes; disc of vertex flat, without median carina, with incomplete furrow spreading from posterior margin to anterior ¼ of vertex length in mid line. Frons (Fig. 7C–D): ratio C/E = 0.41–0.49; D/E = 1.58–1.67; frons longer than wide, widest below level of antennal bases; anterior margin prolonged, lateral margins carinately elevate; disc of frons with three carinae, median carina reaching frontoclypeal suture, lateral carinae not exceeding level of lower margins of compound eyes, median and lateral carinae connected at base. Postclypeus with median and lateral carinae; anteclypeus medially carinate; anterior margin of preocular field covered with row of short setae (Fig. 7C–D). Rostrum reaching between metacoxae; apical segment shorter than subapical one. Compound eyes ellipsoidal, posteriorly with narrow callus. Ocelli distinct. Antennal fovea emarginate, posteriorly elevated, shifted to posterior margin of gena. Scape short, cylindrical. Pedicel bulbous, slightly longer than wide, arista about 3 times as long as pedicel; antennal plate organs evenly distributed on pedicel, organized in rows, reaching base of pedicel. Plate organs of crenelated plate type in the merged-tip type group, with ear-like lobes paired organized on the external circle of them, isolated in the internal area (Fig. 8). THORAX. Pronotum (Figs 6, 7E–F): ratio F/B = 0.44–0.54; disc of pronotum arcuate, with median carina and lateral impressions, lateral carinae not reaching posterior margin, anterior angle incised between compound eyes, exceeding anteriorly half of compound eyes length; posterior margin of pronotum widely triangularly incised, incision reaching level of posterior margins of compound eyes; lateral lobes with curved anteriad; distinctly elevated postocular carinae, not reaching posterior margin of pronotum; lateral carina and horizontal carina of side of pronotum absent. Mesonotum (Figs 6A–B, E–F, 7E–F): ratio G/F = 5.3–6.13, G/B+F = 1.80–2.08, G/H = 0.99–1.03; in mid line about as long as wide, diamond-shaped, with disc flat and lateral portions declivous, with five parallel carinae; median carina reaching almost to scutellum, lateral carinae reaching posterior margins, anterolateral carinae pronounced at base than obsolescent posteriad; mesoscutellar groove arcuate posteriad. Tegula with two parallel carinae (Fig. 9A). TEGMINA (Figs 9B–F, 10A–D). Ratio I/J = 1.86–2.02; membranous, with dense network of veins and veinlets. Costal margin curved at base, arcuate to anteroapical angle; anteroapical angle widely arcuate, posterior margin arcuate, posteroapical angle widely obtuse; tornus straight; apex of clavus reaching ⅔ of tegmen length. Costal area present, wide, with apex reaching almost to ⅔ of tegmen length, merely basad of apex of clavus; costal area with prominent, reticulate network of veinlets. Basal cell rounded, slightly longer than wide. Stems ScP+R, MP and CuA leaving basal cells separately, in ScP+R→MP→CuA forking sequence on corium. Stem ScP+R with short stalk, shorter than basal cell, both branches ScP+RA and RP with few furcations, together with veinlets resulting in reticulate pattern, ultimate terminal of RP reaching anteroapical angle. Common stalk of MP about as long as basal cell, both branches MP1+2 and MP 3+4 with number of furcations, together with veinlets resulting in reticulate pattern, the earliest terminal of MP1+2 reaching anteroapical angle, the ultimate terminal of MP3+4 exceeding posteroapical angle, reaching tornus. Common stem CuA distinctly longer than basal cell; branch CuA 1 weaker, with a few furcations, reaching tornus with three terminals, branch CuA 2 stronger, parallel to claval suture and vein CuP, reaching margin with single terminal. Claval suture and vein CuP distinct. Claval veins Pcu and A1 fused in basal half of clavus length, joined vein Pcu+A1 reaching apex of clavus; irregular veinlets between CuP and Pcu present. Costal cell narrow, with a few veinlets between stems Pc+CP and ScP+RA. Nodal line not materialized. Chitinized pterostigmal area absent. Veinlet icu present, connecting CuA 2 and apex of clavus. Veins with short, scarce setae on ventral surface. HIND WING. Membranous, costal margin slightly curved at base, then almost straight, slightly concave at level of wing coupling apparatus, anteroapical angle widely rounded, apical margin distinctly curved, posteroapical angle widely rounded, anal lobe widely rounded. Basal cell slightly longer than wide, stems ScP+R and M leaving basal cell with a short common stalk. Stem ScP+R forked at level of wing coupling apparatus, ScP+RA 1 reaching margin distinctly basad of anteroapical angle, with 3 terminals; single rarp veinlet present. RP forked apicad of transverse veinlet rp-mp, about at the level of ending RA vein; RP with 4–6 terminals reaching margin basad of and at anteroapical angle. Transverse veinlets rp-mp 3–4. Stem MP forked usually about level of first fork of RP, with 4–5 terminals, single imp vein present between MP 1+2 and MP 3+4. Stem CuA forked distinctly basad of stem ScP+R forking, before half of wing, multifurcation with 11–16 terminals and with 10–12 icu veins. Stem CuP single, stem Pcu subparallel to CuP in basal portion, apically curved towards CuP. Veins with short scarce setae on ventral surface. LEGS (Figs 10E–F, 11). Profemur slightly flattened, with rows of short setae along margins, subquadrate in cross section, with rows of short setae along margin; basiprotarsomere as long as midprotarsomere, apical protarsomere slightly longer than cumulative length of basi-and midprotarsomeres; tarsal claws distinct, arolium bilobate, wide. Mesofemur (Fig. 10E) slightly longer than profemur, subquadrate in cross section, with rows of short setae along margins, basimesotarsomere (Figs 10E, 11A) as long as midmesotarsomere, apical mesotarsomere slightly longer than cumulative length of basi-and midmesotarsomeres; tarsal claws distinct, arolium bilobate, wide. Metafemur (Fig. 11B) shorter than metatibia, metatibia with 5–7 lateral spines, subquadrate in cross sections, with margins covered with rows of short setae, with asymmetrical 8 apical teeth (Fig. 11B–D). Basimetatarsomere (Fig. 11C–D) long, about as long as cumulative length of mid- and apical metatarsomeres, apical margin incised, with 12 apical teeth, with subapical setae, except the external ones; midmetatarsomere long, lateral teeth distinct, ventromedian margin arcuately convex, with bunch of setae; apical metatarsomere with rows of longitudinal setae; tarsal claws distinct, arolium bilobate. MALE TERMINALIA (Figs 12–13). Anal tube (Figs 12A–C, 13A) tubular, slightly longer than wide with posterior margin concave in dorsal view, produced ventrally in lateral view. Pygofer (Figs 12A–B, 13C) with upper part distinctly wider than ventral, posterior margin strongly sinuated, without any processes. Corpus connectivi (Fig. 13D) short and robust; alae connectivi large, bearing well developed crista. Periandrium (Fig. 13D–F) covering half of aedeagus, slightly asymmetrical, with two pairs of broad spine-like processes; dorsal processes bigger than ventral ones; ventral basal spine-like process on right side and a short membranous bulb, covered with minute teeth. Aedeagus (Fig. 13G–H) distinctly longer than periandrium, long and narrow, apically, with a pair of dorsal sclerotized strong spines, surrounding a pair of hook-like longer processes. FEMALE TERMINALIA. Pregenital sternite (Figs 14A–D, 16A–B) well developed with posterior margin with huge submedian protruding lobes with margin between them weakly arcuate. Anal tube (Figs 14A, 15A–D, 16C–E) round, short: shorter than wide in dorsal view. Paraproct small, apical margin widely rounded, reaching posterior margin of anal tube. Epiproct slightly shorter than paraproctal lobe. Gonoplac (Figs 14A–B, 15E–F, 16F) quadrangular, slightly sclerotized. Gonospiculum bridge short and robust (Fig. 16I–K). Gonocoxa VIII (Fig. 16G–H) trapezoidal, with a deep concave incision on its anterior margin. Endogocoxal lobe (Fig. 16G–H) bearing a membranous endogonocoxal process with minute apical teeth, slightly shorter than the anterior connective lamina. Anterior connective lamina with some strong 5 subterminal and 3 apical well sclerotized teeth (Fig. 16G–H). Fibula anterior slightly curved. Gonapophysis IX elongated, membranous with median lobes united medially, straight, densely covered dorsally with minute, scale-like denticulations, almost as long as lateral lobes, which bear the posterior fibulae; lateral margin with a long, developed finger-like lobe, covered dorsally with minute, scale-like denticulations. COLORATION (Figs 3A–C, 4A–F, 5). General coloration green. In dry and older specimens, general coloration reddish-brown or brown. Hind wing membranous, transparent, with one dark, round spot on anal area. Abdomen in ʻfreshʼ specimens with orange tergites and sternites yellowish, yellowish-brown to brown in oldest specimens. Distribution (Fig 17A) Madagascar (East): Alaotra-Mangoro Region: Moramanga District: Andasibe commune; Analamanga Region: Manjakandriana District, Mantasoa commune; Haute Matsiatra Region: Ambohimahasoa District, Morafeno commune; Vatovavy Fitovinany Region: Ifanadiana District, Ranomafana commune. Note The genus Laberia was described by Stål (1866), based on single female specimen labelled as originating from Mauritius (Fig. 3D), as a new genus belonging to Ricaniidae. This seems to be a case of mislabelling, as since the original description it was never reported on Mauritius, and all specimens available came from Madagascar. Melichar (1898: 296) moved this genus to Nogodinidae, and Fennah (1978: 118) placed it in the tribe Bladinini Kirkaldy, 1907, subtribe Gaetulina Fennah, 1978. The subsequent changes in placement of Laberia in Tropiduchidae resulted from taxonomic views and opinions concerning Gaetuliini (see above). However, the type material of Laberia has never been re-examined since its description and its characters and taxonomic status needed to be reconciled with the taxonomic changes that took place around the families Nogodinidae-Tropiduchidae and their respective delimitation. A few additional specimens of the species were found dispersed in various collections enabling this revisionary study that led to our separation of the genus into a new tropiduchid Elicinae tribe, Laberiini trib. nov. Additionally, during this study, we also discovered that another ‘mythical’ taxon from Madagascar, Perinetia reticulata Lallemand & Synave, 1954, originally described as a species of a monotypic genus of the family Acanaloniidae Amyot & Serville, 1834, was conspecific with Laberia palliata Stål, 1866. The generic name Perinetia was proposed by various authors several times for various species of animals. Currently, all these usages have been replaced by new names. Hence, the generic name Perinetia becomes again available for the purposes of taxonomy. However, the oldest usage of this name for a genus is now believed to be a junior objective synonym. All subsequent proposals with Perinetia as generic name have already been replaced. The usages of the generic name Perinetia are presented below: 1936 Perinetia Collenette: 165 [Insecta: Lepidoptera: Lymantriidae]; type species: Perinetia leucocloea Collenette, 1936: 166, pl. 12, fig. 8; by original designation. Remark. Perinetia Collenette, 1936 has priority, while the other names should be treated as junior homonyms (Article 60 of the International Code of Zoological Nomenclature – ICZN 1999). Perinetia Collenette, 1936 is a junior subjective synonym of Stenaroa Hampson, 1910: 444 [Insecta: Lepidoptera: Lymantriidae]. 1952 Perinetia Seyrig: 193. [Insecta: Hymenoptera: Ichneumonidae]; type species: Perinetia nigrifacies Seyrig, 1952: 194; by original designation. Remark. This name was proposed to be replaced by Madagascesa Koçak & Kemal, 2008: 6. 1954 Perinetia Lallemand & Synave: 81 [Insecta: Hemiptera: Acanaloniidae]; type species: Perinetia reticulata Lallemand & Synave, 1954: 82. Remark. This name was proposed to be replaced by Aylaella Demir & Özdikmen, 2009: 271. 1959 Perinetia Barnard: 81 [Crustacea: Malacostraca: Philosciidae]; type species: Philoscia (Perinetia) reducta Barnard, 1958: 81. Remark. This name was proposed to be replaced by Barnardetia Xing & Chen, 2013: 399. 1964 Perinetia Descamps: 203, 206 [Insecta: Orthoptera: Eumastacidae]; type species: Perinetia annulipes Descamps, 1964: 208, figs 349–351; by original designation and monotypy. Remark. This name was proposed to be replaced by Perinetella Descamps & Wintrebert, 1965: 96. However, this name was preoccupied by Perinetella Synave, 1956: 211; type species: Perinetella nigroflava Synave, 1956: 2012, figs 9–10; by original designation and monotypy (Insecta: Hemiptera: Flatidae), and a new replacement name was proposed – Descampsiella Özdikmen, 2008: 67. 1988 Perinetia Domergue: 135 [Reptilia: Serpentes: Colubridae]; type species: Perinetia coulangesi Domergue, 1988; by original designation and monotypy. Remark. This name was proposed to be replaced by Brygophis Domergue & Bour, 1989: 805., Published as part of Stroiński, Adam, Bourgoin, Thierry & Szwedo, Jacek, 2022, Laberiini, a new tribe of Tropiduchidae planthoppers from Madagascar (Hemiptera: Fulgoroidea), pp. 23-54 in European Journal of Taxonomy 836 (1) on pages 31-48, DOI: 10.5852/ejt.2022.836.1913, http://zenodo.org/record/7051851, {"references":["Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276.","Lallemand V. & Synave H. 1954. Homopteres nouveaux de Madagascar. 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Annals and Magazine of Natural History (8) 5: 430 - 464, 465 - 496. https: // doi. org / 10.1080 / 00222931008692804","Seyrig A. 1952. Les Ichneumonides de Madagascar. IV Ichneumonidae Cryptinae. Memoires de l'Academie malgache 19 (39): 1 - 213.","Kocak A. O. & Kemal M. 2008. Nomenclatural notes on the genus group names in the families Braconidae and Ichneumonidae (Hymenoptera). Centre for Entomological Studies Miscellaneous Papers 144: 6.","Barnard K. H. 1958. Terrestrial isopods and amphipods from Madagascar. Memoires de l'Institut scientifique de Madagascar (serie A) 12: 67 - 111.","Descamps M. 1964. Revision preliminaire des Euschmidtiinae (Orthoptera-Eumastacidae). Memoires du Museum national d'Histoire naturelle. Ser. A, Zoologie 30: 1 - 321.","Descamps M. & Wintrebert D. 1965. Contribution a l'etude des eumastacides malgaches (OrthopteraEumastacidae). Memoires du Museum national d'Histoire naturelle, Ser. A, Zoologie 34: 1 - 188.","Synave H. 1956. Les Flatidae de Madagascar (Hemiptera-Homoptera). Memoires de l'Institut des Sciences de Madagascar (Ser. E) 7: 197 - 217.","Ozdikmen H. 2008. Nomenclatural changes for some Orthoptera (Ensifera and Caelifera). Zootaxa 1763: 67 - 68. https: // doi. org / 10.11646 / zootaxa. 1763.1.6","Domergue C. A. 1988. Notes sur les serpents de la region malgache. 8. Colubridae nouveaux. Bulletin du Museum national d'Histoire naturelle Sect. A Zoologie Biologie et Ecologie animales 10 (1): 135 - 146.","Domergue C. A. & Bour R. 1989. Brygophis nom nouveau pour Perinetia Domergue, 1988, preemploye (Reptilia, Colubridae). Bulletin du Museum national d'Histoire naturelle Sect. A Zoologie Biologie et Ecologie animales 10 (ser. 4): 805 - 806."]}

Tribe Laberiini trib. nov. urn:lsid:zoobank.org:act: F8F8C80E-EA21-4759-AF47-C4BB195F1082 Type genus Laberia Stål, 1866; here designated. Diagnosis Head capsule narrower than thorax, projecting in front of eyes. Pedicel with plate organs on the whole surface reaching to its base. Rostrum long, ending between metacoxae. Tegmina broad, extending far beyond abdomen, held flattening tectiform, with their ventral faces not facing each other; costal area present and wide, with reticulate venation; nodal line not materialized. Hind tibia with 5–7 lateral spines. Metatarsal apical teeth asymmetrical. Male gonostyli symmetrical, not fused. Female: gonoplacs flat quadrangular, wide, without apical marginal teeth. Gonapophyses IX with a long, lateral, digitated process. Anal tube short, ring-shaped. Content Monotaxic tribe currently with one genus – Laberia Stål, 1866. A new simplified key to higher taxa of Elicinae to accommodate the new tribe Laberiini trib. nov. is proposed here: 1. Tegmina usually with distinct nodal line and venation distinctly reticulate distally; gonoplacs elongated, usually with marginal teeth; gonapophyses IX in shape of isosceles triangle.................................................................................................................................... Tropiduchinae Stål, 1866 – Nodal line of tegmina not distinct; gonoplacs rounded without marginal teeth; gonapophyses IX not triangular............................................................................................... Elicinae Melichar, 1915... 2 2. Tegmina short, reticulate, with transverse costal veins; ScP+R and MP with common stem................................................................................................................ Patollini Szwedo & Stroinski, 2013 – Tegmina short or surpassing abdomen; ScP+R and MP leaving basal cell separately..................... 3 3. Tegmina steeply tectiform, surpassing abdomen; in dorsal view lateral margins of gonapophyses IX straight.............................................................................................................................................. 4 – Tegmina flatly tectiform; in dorsal view lateral margins of gonaphysis distinctly protruded with tooth or long digitate process..................................................................................................................... 5 4. Tegmina not reticulated; metatibia with one spine; metatarsal apical teeth asymmetrical...................................................................................................... Bucini Gnezdilov, Bartlett & Bourgoin, 2016 – Tegmina reticulated; metatibia without spines; metatarsal apical teeth regular............................................................................................................................................. Parathisciini Gnezdilov, 2013 5. In dorsal view lateral margin of gonapophysis protruded with a short tooth-like process.......................................................................................................................................... Elicini Melichar, 1915 – In dorsal view lateral margin of gonapophysis bearing a long digitate process............................................................................................................................................................... Laberiini trib. nov., Published as part of Stroiński, Adam, Bourgoin, Thierry & Szwedo, Jacek, 2022, Laberiini, a new tribe of Tropiduchidae planthoppers from Madagascar (Hemiptera: Fulgoroidea), pp. 23-54 in European Journal of Taxonomy 836 (1) on pages 29-30, DOI: 10.5852/ejt.2022.836.1913, http://zenodo.org/record/7051851, {"references":["Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276.","Melichar L. 1915. Monographie der Lophopinen. Annales Historico-Naturales Musei Nationalis Hungarici 13: 337 - 385.","Szwedo J. & Stroinski A. 2013. An extraordinary tribe of Tropiduchidae from the Eocene Baltic amber (Hemiptera: Fulgoromorpha: Fulgoroidea). Zootaxa 3647 (2): 371 - 381. https: // doi. org / 10.11646 / zootaxa. 3647.2.8","Gnezdilov V. M., Bartlett C. R. & Bourgoin T. 2016. A new tribe of Tropiduchidae (Hemiptera: Fulgoroidea) with revision of the genus Buca and description of asymmetric hind leg spinulation. Florida Entomologist 99 (3): 406 - 416. https: // doi. org / 10.1653 / 024.099.0311","Gnezdilov V. M. 2013. Contribution to the taxonomy of the family Tropiduchidae Stal (Hemiptera, Fulgoroidea) with description of two new tribes from Afrotropical Region. Deutsche entomologische Zeitschrift 60 (2): 179 - 191."]}

Tribe Laberiini trib. nov. urn:lsid:zoobank.org:act: F8F8C80E-EA21-4759-AF47-C4BB195F1082 Type genus Laberia Stål, 1866; here designated. Diagnosis Head capsule narrower than thorax, projecting in front of eyes. Pedicel with plate organs on the whole surface reaching to its base. Rostrum long, ending between metacoxae. Tegmina broad, extending far beyond abdomen, held flattening tectiform, with their ventral faces not facing each other; costal area present and wide, with reticulate venation; nodal line not materialized. Hind tibia with 5–7 lateral spines. Metatarsal apical teeth asymmetrical. Male gonostyli symmetrical, not fused. Female: gonoplacs flat quadrangular, wide, without apical marginal teeth. Gonapophyses IX with a long, lateral, digitated process. Anal tube short, ring-shaped. Content Monotaxic tribe currently with one genus – Laberia Stål, 1866. A new simplified key to higher taxa of Elicinae to accommodate the new tribe Laberiini trib. nov. is proposed here: 1. Tegmina usually with distinct nodal line and venation distinctly reticulate distally; gonoplacs elongated, usually with marginal teeth; gonapophyses IX in shape of isosceles triangle.................................................................................................................................... Tropiduchinae Stål, 1866 – Nodal line of tegmina not distinct; gonoplacs rounded without marginal teeth; gonapophyses IX not triangular............................................................................................... Elicinae Melichar, 1915... 2 2. Tegmina short, reticulate, with transverse costal veins; ScP+R and MP with common stem................................................................................................................ Patollini Szwedo & Stroinski, 2013 – Tegmina short or surpassing abdomen; ScP+R and MP leaving basal cell separately..................... 3 3. Tegmina steeply tectiform, surpassing abdomen; in dorsal view lateral margins of gonapophyses IX straight.............................................................................................................................................. 4 – Tegmina flatly tectiform; in dorsal view lateral margins of gonaphysis distinctly protruded with tooth or long digitate process..................................................................................................................... 5 4. Tegmina not reticulated; metatibia with one spine; metatarsal apical teeth asymmetrical...................................................................................................... Bucini Gnezdilov, Bartlett & Bourgoin, 2016 – Tegmina reticulated; metatibia without spines; metatarsal apical teeth regular............................................................................................................................................. Parathisciini Gnezdilov, 2013 5. In dorsal view lateral margin of gonapophysis protruded with a short tooth-like process.......................................................................................................................................... Elicini Melichar, 1915 – In dorsal view lateral margin of gonapophysis bearing a long digitate process............................................................................................................................................................... Laberiini trib. nov.

The tropiduchid monotypic genus Laberia Stål, 1866, endemic to Madagascar, is placed in a new tribe of its own Laberiini trib. nov. The tribe is placed in the subfamily Elicinae, and can be distinguished from other representatives of the subfamily by the unique combination of morphological characters of the head, tegmina, legs and genital structures. The genus Laberia and its only species Laberia palliata Stål, 1866 are redescribed, chresonymy is presented and nomenclatorial questions are clarified. Distributional data from Madagascar and doubtful presence of the taxon in Mauritius are discussed. Taxonomic content of Elicinae is briefly discussed.

Genus Laberia Stål, 1866 Laberia Stål, 1866: 234. Type species: Laberia palliata Stål, 1866: 234; by monotypy. Perinetia Lallemand & Synave, 1954: 81; syn. nov. Type species: Perinetia reticulata Lallemand & Synave, 1954: 82; by monotypy and original designation. Aylaella Demir & Özdikmen, 2009: 271; syn. nov. Type species: Perinetia reticulata Lallemand & Synave, 1954: 82. Description BODY. Head capsule with vertex longer than wide, strongly projecting in front of eyes. Anterior margin triangular, lateral margins slightly diverging posteriorly, posterior margin anteriorly rounded concave; no median carina. Frons with lateral margins slightly diverging down to below level of the antenna, then converging to continue with lateral margin of postclypeus; median carina weak and thin, continuing on the clypeus; a pair of short dorsal latero-median carinae vanishing at the level of the compound eyes. No median ocellus. Rostrum long, with tip between metacoxae. Pronotum with anterior margin strongly produced, rounded, postocular carina interrupted behind compounds eyes, laterally continuing in an anteriorly curved ridge. Mesonotum with a weak median carina and a pair of lateral ones, anteriorly bifid. Tegulae large, bicarinate. Tegmina broad, extending far beyond the end of abdomen, with venation distinctly reticulate; costal area present and wide, with reticulate venation; ScP+R, MP and CuA branched separately on the basal cell. LEGS. Hind tibia with 5–7 lateral spines. Metatarsus: basimetatarsomere with asymmetrical row of apical teeth, midmetatarsomere with symmetrical row of apical teeth. Metatibiotarsal formula: 8/12/2. MALE TERMINALIA. Male gonostyli (Figs 12, 13B) symmetrical, not fused medioventrally, more or less trapezoidal, with a small basal triangular expansion on its dorsal margin; capitulum not developed. FEMALE TERMINALIA. Gonoplacs wide, flat, quadrangular, with no apical marginal teeth. Gonapophyses IX with a long lateral digitated process. Anal tube short, ring-shaped. Content Laberia palliata Stål, 1866, Published as part of Stroiński, Adam, Bourgoin, Thierry & Szwedo, Jacek, 2022, Laberiini, a new tribe of Tropiduchidae planthoppers from Madagascar (Hemiptera: Fulgoroidea), pp. 23-54 in European Journal of Taxonomy 836 (1) on page 30, DOI: 10.5852/ejt.2022.836.1913, http://zenodo.org/record/7051851, {"references":["Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276.","Lallemand V. & Synave H. 1954. Homopteres nouveaux de Madagascar. Le Naturaliste malgache 6: 79 - 82.","Demir E. & Ozdikmen H. 2009. Two new replacement names for genera in Dictyopharidae and Acanaloniidae (Hemiptera: Auchenorrhyncha). Proceedings of the Entomological Society of Washington 111 (1): 271. https: // doi. org / 10.4289 / 0013 - 8797 - 111.1.271"]}

Tribe Elicini Melichar, 1915 Elicini Melichar, 1915: 379. Type genus Elica Walker, 1857: 86. Content and distribution Currently, Elicinae includes 46 genera and 158 species and Elicini remains the most diverse tribe within the subfamily with 37 genera (1.5% of the Fulgoromorpha) and 143 species (1% of the Fulgoromorpha) (Bourgoin 2022; Fig. 2). Due to its complex history chronologically listed here, that we complete, we provide a graphical historical synopsis of the generic composition of the tribe (Fig. 2A–B). 1915 Elicaini [sic!] Melichar (Melichar: 379; definition, list of genera, key) 1978 Gaetuliina (Fennah: 118; definition, list of genera comprised) 1978 Elicina (Fennah: 118; new status, definition, list of genera comprised) 1984 Gaetuliina Fennah (Fennah: 82; update of definition, transfer of genera from Issidae) 1985 Gaetuliina (O’Brien & Wilson: 89; difficulties of recognition of Nogodinidae after inclusion of Gaetuliinae) 1999 Gaetulina [sic!] (Szwedo & Stroiński: 203; taxonomy) 1999 Gaetulini [sic!] (Szwedo & Stroiński: 297; taxonomy, discussion of characters) 2007 Gaetuliini Fennah (Gnezdilov: 295; new status within Tropiduchidae) 2008 Gaetuliini (Fletcher: 119; transfer of the genus Busas Jacobi, 1909 to Gaetuliini) 2009 Gaetuliina (Stroiński & Gnezdilov: 459; taxonomy) 2009 Gaetuliini (Stroiński & Gnezdilov: 459, 460; redescription of Busas Jacobi, 1909, list of genera) 2010 Gaetuliina (Szwedo & Stroiński: 132; taxonomy) 2010 Gaetulina Fennah, 1978 [sic!] (Szwedo & Stroiński: 137; taxonomy) 2010 Gaetulini Fennah, 1978 [sic!] (Szwedo & Stroiński: 136, 137; fossil record, taxonomy) 2012b Gaetuliini Fennah, 1978 (Gnezdilov: 637, 638; taxonomy, distribution, description of new genus) 2013 Gaetuliina Fennah, 1978 (Szwedo & Stroiński: 371; taxonomy) 2013 Gaetuliini Fennah, 1978 (Szwedo & Stroiński: 380; characters, fossil record) 2013 Elicini Melichar, 1915 (Gnezdilov: 184; taxonomy) 2013 Elicaini Melichar, 1915 (Gnezdilov: 184; taxonomy, synonymy) 2013 Gaetuliina Fennah, 1978 (Gnezdilov: 184; taxonomy, synonymy) 2015 Elicini Melichar, 1915 (Wang M. et al. 2015: 563, 564; taxon concept history, new genus Connelecita Wang & Bourgoin, 2015) 2015 Elicini Melichar, 1915 (Gnezdilov & Bourgoin: 601; two new genera from Madagascar: Bolitropis Gnezdilov & Bourgoin, 2015 and Bambomada Gnezdilov & Bourgoin, 2015) 2019 Elicini Melichar, 1915 (Szwedo 2019 in Szwedo et al. 2019, fossil genera: Senogaetulia Szwedo, 2019 and Dakrutulia Szwedo, 2019) The fossil taxa of Elicini are known from the Eocene Baltic amber (Tritophania Jacobi, 1938) and terminal Eocene deposits of Bembridge Marls, Isle of Wight, United Kingdom (Senogaetulia Szwedo, 2019 and Dakrutulia Szwedo, 2019) (Jacobi 1938; Szwedo & Stroiński 1999; Szwedo et al. 2019). The tribe Elicini is widely distributed in the New World, in South Africa and Madagascar, in south east continental Asia and Indonesia, and in Australia. It exhibits a bimodal latitudinal distribution with peaks in north and south subtropical regions, while it is less represented in the equatorial zone., Published as part of Stroiński, Adam, Bourgoin, Thierry & Szwedo, Jacek, 2022, Laberiini, a new tribe of Tropiduchidae planthoppers from Madagascar (Hemiptera: Fulgoroidea), pp. 23-54 in European Journal of Taxonomy 836 (1) on pages 28-29, DOI: 10.5852/ejt.2022.836.1913, http://zenodo.org/record/7051851, {"references":["Melichar L. 1915. Monographie der Lophopinen. Annales Historico-Naturales Musei Nationalis Hungarici 13: 337 - 385.","Walker F. 1857. Catalogue of the Homopterous insects collected at Singapore and Malacca by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society 1: 82 - 100. https: // doi. org / 10.1111 / j. 1096 - 3642.1856. tb 00958. x","Bourgoin T. 2022. FLOW (Fulgoromorpha Lists on The Web): a world knowledge base dedicated to Fulgoromorpha (Insecta: Hemiptera: Fulgoromorpha). Paris. Version 8, updated 2022 - 02 - 03. Available from http: // hemiptera-databases. org / flow / [accessed 13 Jul. 2022].","Fennah R. G. 1978. The higher classification of the Nogodinidae (Homoptera: Fulgoroidea) with a description of a new genus and species. Entomologist's Monthly Magazine 113: 113 - 120.","Bourgoin T., Wang R. R., Asche M., Hoch H., Soulier-Perkins A., Stroinski A., Yap S. & Szwedo J. 2015. From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha). Zoomorphology 134: 63 - 77. https: // doi. org / 10.1007 / s 00435 - 014 - 0243 - 6","Gnezdilov V. M. & Bourgoin T. 2015. New genera and new species of the tribe Elicini (Hemiptera: Fulgoroidea: Tropiduchidae) with key to Tropiduchid genera known from Madagascar. Annales Zoologici 65 (4): 599 - 618. https: // doi. org / 10.3161 / 00034541 ANZ 2015.65.4.007","Szwedo J., Drohojowska J., Popov Y., Simon E. & Wegierek P. 2019. Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 110 (3 - 4): 331 - 396. https: // doi. org / 10.1017 / S 175569101900001 X","Jacobi A. 1938. Eine neue Bernsteinzikade (Rhynchota: Homoptera). Sitzungberichte der Gesellschaft Naturforschender Freunde zu Berlin, 15, 188 - 189.","Szwedo J. & Stroinski A. 1999. Redescription of Tritophania patruelis Jacobi, 1938 from Eocene Baltic amber (Hemiptera: Nogodinidae). Annales Zoologici 49 (3): 203 - 207."]}

A new monotypic genus of nogodinid planthoppers (Hemiptera: Fulgoromorpha: Nogodinidae) is described from Dominican amber. The adult female of Jatoba losbrachi gen. et sp. nov. can be distinguished...

Nikomiklukha Gnezdilov, 2010 Type species: Issus praecedens Walker, 1857; by original designation by Gnezdilov 2010: 48. Sarimissus Wang, Zhang & Bourgoin, 2019: Gnezdilov, 2019: 1302 [sic!; error]2010 Gnezdilov, p. 48 [new genus] 2019 Gnezdilov, p. 1302 Composition: Nikomiklukha praecedens (Walker, 1857), Nikomiklukha sumatrana Gnezdilov, 2010, Nikomiklukha maclayi Gnezdilov, 2010., Published as part of Qin, Dao-Zheng & Stroi��ski, Adam, 2022, Sarillanus nom. nov., to replace Sarimissus Meng, Qin et Wang, 2020 nec Sarimissus Wang, Zhang et Bourgoin, 2019 (Hemiptera: Issidae), pp. 247-250 in Zootaxa 5093 (2) on page 248, DOI: 10.11646/zootaxa.5093.2.8, http://zenodo.org/record/5905274, {"references":["Gnezdilov, V. M. (2010) Three new genera and three new species of the family Issidae (Hemiptera: Fulgoromorpha) from Borneo and Sumatra. Tijdschrift voor Entomologie, 153, 41 - 52.","Walker, F. (1857) Catalogue of the Homopterous insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society. London, 1 (4), 141 - 175.","Wang, M. L., Zhang, Y. L. & Bourgoin, T. (2019) On the tribe Sarimini with two new genera from south of China (Hemiptera, Fulgoromorpha, Issidae). Zootaxa, 4706 (2), 375 - 383. https: // doi. org / 10.11646 / zootaxa. 4706.2.10","Gnezdilov, V. M. (2019) On the synonymy and distribution of the planthopper genera Euroxenus Gnezdilov, 2009 and Nikomiklukha Gnezdilov, 2010 (Hemiptera, Auchenorrhyncha, Fulgoroidea: Issidae). Entomological Review, 99 (9), 1299 - 1303. https: // doi. org / 10.1134 / S 0013873819090070"]}

Qin, Dao-Zheng, Stroiński, Adam (2022): Sarillanus nom. nov., to replace Sarimissus Meng, Qin et Wang, 2020 nec Sarimissus Wang, Zhang et Bourgoin, 2019 (Hemiptera: Issidae). Zootaxa 5093 (2): 247-250, DOI: https://doi.org/10.11646/zootaxa.5093.2.8

Sarimissus Wang, Zhang et Bourgoin, 2019 Type species. Sarimissus maculifrons Wang, Zhang et Bourgoin, 2019: 377; by original designation and monotypy. 2019 Wang, Zhang & Bourgoin, p. 376 [new genus, description] 2019 Gnezdilov, p. 1302 [synonymy under Nikomiklukha Gnezdilov, 2010] 2020 Nikomiklukha: Chang et al. 2020, p. 33 Composition. Monotypic genus��� Sarimissus maculifrons Wang, Zhang & Bourgoin, 2019., Published as part of Qin, Dao-Zheng & Stroi��ski, Adam, 2022, Sarillanus nom. nov., to replace Sarimissus Meng, Qin et Wang, 2020 nec Sarimissus Wang, Zhang et Bourgoin, 2019 (Hemiptera: Issidae), pp. 247-250 in Zootaxa 5093 (2) on page 248, DOI: 10.11646/zootaxa.5093.2.8, http://zenodo.org/record/5905274, {"references":["Wang, M. L., Zhang, Y. L. & Bourgoin, T. (2019) On the tribe Sarimini with two new genera from south of China (Hemiptera, Fulgoromorpha, Issidae). Zootaxa, 4706 (2), 375 - 383. https: // doi. org / 10.11646 / zootaxa. 4706.2.10","Gnezdilov, V. M. (2010) Three new genera and three new species of the family Issidae (Hemiptera: Fulgoromorpha) from Borneo and Sumatra. Tijdschrift voor Entomologie, 153, 41 - 52.","Chang, Z. - M., Yang, L. & Chen X. - S. (2020) Two new genera with species of the tribe Sarimini (Hemiptera, Fulgoromorpha, Issidae) from China. ZooKeys, 956, 31 - 47. https: // doi. org / 10.3897 / zookeys. 956.47784"]}

Simple Summary Although previous studies have recently explored the phylogenetic relationships among the planthopper families, the taxonomic relationships between Ricaniidae and other families of Fulgoroidea need to be further explored. Meanwhile, the morphological definitions of the two largest genera, Pochazia Amyot & Serville, 1843 and Ricania Germar, 1818 (the type genus of Ricaniidae) remain controversial, and their monophyly status has never been established. This study aims to clarify the relationship of Ricaniidae with other families of Fulgoroidea and to provide evidence to clarify the differences between these two related genera for species attribution. Our results support the monophyly of Ricaniidae and the sister group status of the two families Flatidae and Ricaniidae but fail to support the monophyly of Pochazia and Ricania. Diagnoses between these two genera cannot be resolved until more evidence is acquired. This study provides new evidence toward the phylogenetic analysis and revision of the distinguishing characteristics of related genera in this family. Abstract Ricaniidae is a relatively small planthopper family with about 69 genera and 442 species worldwide. Members of this family occur throughout the warm temperate and tropical regions. Some species cause devastating damage to major agricultural and economic plants. However, the relationship between Ricaniidae and other families of Fulgoroidea needs to be further explored. The morphological definitions of the two biggest genera, Pochazia Amyot & Serville, 1843 and Ricania Germar, 1818 (the type genus of Ricaniidae) remain controversial. In this study, mitogenomes of five representatives in these two genera were decoded using the next-generation sequence method and genome assembly. Results showed that their complete mitogenomes are circular DNA molecules with 15,457 to 16,411 bp. All protein-coding genes (PCGs) begin with the start codon ATN, GTG or TTG and end with TAA, TAG, an incomplete stop codon single T or an incomplete stop codon single A. A lost DHU arm was discovered in the trnS gene of the five mitogenomes and the trnV gene within Pochazia confusa, Pochazia guttifera and Ricania simulans. The remnant tRNAs folded into clover-leaf structures. The sliding window, genetic distance, and Ka/Ks analyses indicated that the cox1 gene is the slowest evolving and is relatively conserved. The phylogenetic tree topologies support (Delphacidae + (((Issidae + (Lophopidae + Caliscelidae)) + (Flatidae + Ricaniidae)) + (Achilidae + (Dictyopharidae + Fulgoridae)))) as the best topology, as recognized by both PhyloBayes, RAxML and MrBayes based on four data sets (PCG, PCGRNA, PCG12, PCG12RNA). The monophyly of Ricaniidae and the sister group status of two families Flatidae and Ricaniidae are supported, but all analyses failed to support the monophyly of Pochazia and Ricania. The diagnoses between these two genera cannot be resolved until more evidence is acquired.

We use the theory of Tambara modules to extend and generalize the reconstruction theorem for module categories over a rigid monoidal category to the non-rigid case. We show a biequivalence between the $2$-category of cyclic module categories over a monoidal category $\mathscr{C}$ and the bicategory of algebra and bimodule objects in the category of Tambara modules on $\mathscr{C}$. Using it, we prove that a cyclic module category can be reconstructed as the category of certain free module objects in the category of Tambara modules on $\mathscr{C}$, and give a sufficient condition for its reconstructability as module objects in $\mathscr{C}$. To that end, we extend the definition of the Cayley functor to the non-closed case, and show that Tambara modules give a proarrow equipment for $\mathscr{C}$-module categories, in which $\mathscr{C}$-module functors are characterized as $1$-morphisms admitting a right adjoint. Finally, we show that the $2$-category of all $\mathscr{C}$-module categories embeds into the $2$-category of categories enriched in Tambara modules on $\mathscr{C}$, giving an ''action via enrichment'' result., Comment: 62 pages

Bitara gen. nov. Type species. Bitara augusta sp. nov.; by present designation and monotypy. Diagnosis. Similar to Oechalinella Wang, 2017 in Wang et al. 2017, in general appearance. Differ in shape of the head, at lateral margin as long as wide, with anterior margin arcuate (nearly three times as long as wide, with anterior margin acutely angulate in Oechalinella); pronotum with median carina not reaching anterior margin (median carina reaching anterior margin in Oechalinella); male pygofer, medially fused gonostyli and aedeagus asymmetrical, male pygofer with posterodorsal elongate process on right side, left side without process (in Oechalinella male pygofer with posterior margin sinuate, with short lobate semicircular expansion on posterodorsal margin in right side, lobate subquadrate expansion in posterodorsal margin in left side); dorsal margin not excavate at level of anal tube base (dorsal margin distinctly excavate to accommodate base of anal tube in Oechalinella); aedeagal complex straight caudad (aedeagal complex curved in Oechalinella); endosome bulbous, spiniferous, with three distinct long spines (endosome small and smooth in Oechalinella); genital styles tips of dorsal projections teeth-like, directed mediad (margin of dorsal projections smooth and not curved median in Oechalinella); female pregnital sternite narrow, posterior margin medially deeply concave, with median sclerotised process; anal tube distinctly shorter than gonoplac, tubular; gonoplac unilobate, with row of distinct teeth on posterior margin; bases of gonapophyses VIII asymmetrical—left gonapophysis base with lobate, with hook-like process; right base of gonapophysis lobate, smooth. Description. Head including compound eyes narrower than pronotum. Vertex chevron-like in dorsal view, with lateral margins as long as wide, anterior margin arcuate. Frons twice as long as broad in middle, roughly quadrate except abruptly narrowing in ventral forth; lateral margins of frons thickened, carinate, median carina simple. Frontoclypeal suture widely angulate. Clypeus with median longitudinal eminence. Pronotum with anterior margin produced anteriad between compound eyes, with median carina not reaching anterior margin, disc of pronotum not elevated. Mesonotum tricarinate, longer than broad, at same plane as vertex and pronotum, flat, with carinae connected at base, lateral carinae reaching posterior margin. Tegmen hyperpterous long and narrow, membranous, without granulation, with three defined lines formed by veins and veinlets: nodal, first and second postnodal lines. Male pygofer, medially fused gonostyli and aedeagus asymmetrical. Female pregenital sternite narrow, posterior margin medially deeply concave, with median sclerotised process.Anal tube distinctly shorter than gonoplac, tubular. Gonoplac unilobate, with row of distinct teeth on posterior margin. Bases of gonapophyses VIII asymmetrical. Etymology. Generic name is derived from the name of the Sepik language Bitara (Berinomo), spoken in East Sepik Province, Papua-New Guinea, in the area where the specimens were collected. Gender: feminine., Published as part of Stroiński, Adam & Szwedo, Jacek, 2021, Bitara gen. nov. of Tropiduchidae (Hemiptera: Fulgoromorpha) east of Wallace line, pp. 127-139 in Zootaxa 5057 (1) on pages 128-129, DOI: 10.11646/zootaxa.5057.1.8, http://zenodo.org/record/5585596, {"references":["Wang, R. R., Li, X. L., Szwedo, J., Stroinski, A., Liang, A. P. & Bourgoin, T. (2017) Testing Tropiduchini Stal 1866 (Hemiptera: Tropiduchidae) monophyly, a young inter-tropical taxon of mainly insular species: taxonomy, distribution patterns and phylogeny, with the description of a new genus from Papua New Guinea. Systematic Entomology, 42 (2), 359 - 378. https: // doi. org / 10.1111 / syen. 12219"]}

Bitara augusta sp. nov. (Figs 1–46) Diagnosis. Tegmen with cells C1 and C3 of similar length, terminal MP 2 single. Metatibia with 3 lateral spines (including genual one) and row of 6 apical teeth (4+2) forming irregular line without diastema. Process of right lobe of pygofer, arcuately curved, tapering caudad. Spines of endosoma of different sizes; one oriented caudad, ventral one oriented cephalad; dorsal one the shortest, oriented dorsocephalad. Description. Measurements: Total length 1.03–1.16 cm. Head. Vertex: proportion A/B = 1.64–1.81. Frons: proportion C/E = 0.42–0.45; proportion D/E = 0.53–0.58. Thorax. Pronotum: proportion F/B = 0.64–0.68. Mesonotum: proportion G/F+B = 2.22–2.40, proportion G/F = 5.45–5.90, proportion G/H = 1.09–1.18. Tegmina: proportion I/J = 2.41–2.64. Coloration (Figs 1–5). General colour ochraceous, speckled with lighter and darker irregular markings. Anterior and lateral margins of vertex mahogany-red; disc of vertex lighter straw-coloured. Frons ochraceous, with lighter speckles, median carina coloured as background, lateral carinations mahogany-brown. Postclypeus and anteclypeus ochraceous.Anterolateral and posterior margin of pronotum mahogany-reddish, disc of pronotum strawcolored. Mesonotum with lateral carinae mahogany-reddish, disc light straw-coloured, lateral margins ochraceous, speckled. Tegulae ochraceous. Tegmen transparent, membranous, venation ochraceous. Legs uniformly coloured, ochraceous. Head (Figs 1–13). Head with compound eyes narrower than pronotum. Vertex anteriorly projected, much wider than long, chevron-like in dorsal view, anterior margin angulately rounded, posterior margin deeply arcuately incised, lateral margins slightly diverging posteriad, all margins elevated, median carina almost reaching anterior margin; disc of vertex flat. Frons twice as long as broad in middle, roughly quadrate except abruptly narrowing in ventral fourth; lateral margins of frons thickened, carinate, median carina simple. Frontoclypeal suture broadly angulate. Clypeus with median longitudinal eminence. Rostrum reaching hind coxae bases; apical segment longer than broad, shorter than subapical segment. Callus small, posteriad of compound eye (Fig. 5). Ocelli small, below midpoint on compound eye. Antenna with scape very short, pedicel elongated, cylindrical (about as long as wide), with plate organs present on leading surface, tip of pedicel and ventral surface, trichoid sensilla type 1 and type 2 present: plate organs of crenelated type surrounded by a ring of elevated spines, higher than sensorial crests of the plate organ of pedicel. Thorax (Figs 1–7, 11, 14–17). Pronotum short, approximately crescent-shaped in dorsal view; anterior margin roundly produced anteriad between compound eyes; tricarinate, disc not elevated, delimited by lateral carinae, lateral carinae distinctly elevated, median one not so, not reaching anterior margin; posterior margin carinate; sideways T-shaped lateral carina between eye and tegula (base toward tegula). Mesonotum longer than broad, on same plane as vertex and pronotum, flat; conspicuously tricarinate, lateral carinae anteriorly curved to reach median carina, posteriorly parallel, reaching posterior margin; median carina approximately reaching mesoscutellar suture. Mesoscutellar suture straight; posterior margin of mesoscutellum acute. Tegula not carinate. Procoxa shorter than profemur, subquadrate, with margins slightly carinate, smooth. Profemur slightly shorter than protibia, subquadrate in cross-section with rows of setae along. Protibia subquadrate in cross-section; short decumbent setae along margins. Protarsus shorter than half of protibia length, basiprotarsomere scaphoid, apical protarsomere longer than combined length of basiprotarsomere and midprotarsomere. Mesocoxa slightly shorter than mesofemur, subquadrate, with margins smooth. Mesofemur subquadrate in crosssection, shorter than mesotibia, short, decumbent setae along margins. Mesotibia subquadrate in cross-section, with rows of setae along, slightly longer than protibia; Mesotarsus shorter than half of mesotibia length, basimesotarsomere scaphoid, apical mesotarsomere longer than combined length of basiprotarsomere and midprotarsomere. Metacoxa with coxal process widely triangular, very wide at base, spinose at apex. Metatibia distinctly longer than metafemur, not widened apically, with 3 lateral spines and row of 6 apical teeth (4+2) forming irregular line, without diastema: both lateral teeth same size; 4 internal spines different - two lateral internal teeth same size and same as lateral teeth; pair of middle teeth same size and distinctly shorter than others. Basimetatarsomere about as long as cumulative length of mid- and apical metatarsomeres, with apical row of 6 teeth; all teeth almost same size (see also Wang et al. 2017 Fig. 7F). Wings as in Oechalinella (for veins and cells nomenclature see Wang et al. 2017, Figs. 3E, 4D). Tegmen hyperpterous, long and narrow, membranous, without granulation, with three defined transverse lines formed by veins and veinlets (proximal to distal): nodal, first and second postnodal lines. Costal area present, narrower than costal cell, nearly reaching the level of tip of clavus, with sparse cross veins. Costal cell extending beyond midlength the length of tegmen and without transverse veinlets. Stems ScP+R and MP leaving basal cell with a short common stalk; stem ScP+R forked basad of tegmen midlength; branch ScP+RA separated from RP before nodal line. ScP+RA 1 forked before at nodal line, slightly basad of apex of costal area; first fork of RP distinctly apicad of nodal line, at level of first postnodal line. Stem MP forking first at level of nodal line; stalk MP 1+2 markedly longer than stalk MP 3+4; MP 1+2 forked at level of first postnodal line; stalk MP 3+4 forked at level of nodal line. Stem CuA forked basad of ScP+R fork, proximad of tegmen midlength, distad of claval veins junction. Postnodal and subapical row of cells of similar length. Clavus closed (i.e., with CuP reaching margin and claval veins reaching margin), with apical tip acute, basad of end of costal area; claval veins Pcu and A 1 fused basad of half of CuP. Posterocubital cell, postcubital cell and anal cell without transverse veinlets. Hind wings hyaline, elongate, slightly shorter than tegmen, with costal cell widened at base, with anal lobe wide. Stems ScP+R, MP and CuA, fused at base. Stem ScP+R forked at level of wing coupling apparatus (see Wang et al. 2017, Figs 4E & 5D), at level of CuA 1 branching; branch ScP+RA with single or two terminals, reaching margin well basad of apex of wing; branch RP with three terminals reaching margin at apical angle of wing. Stem MP not forked before rp-mp and mp-cua veinlets, three terminals MP 1, MP 2 and MP 3+4 reaching margin forked well apicad. Stem CuA forked slightly basad of stem ScP+R forking, branch CuA 1 forked again basad of mpcua veinlet; terminals CuA 1a, CuA 1b and CuA 2 forked apically, reaching margin with five-six terminals. Stem CuP single. Stem Pcu distinctly curved before apex, fused for a distance with branching of A 1, A 2 single. Stems CuA and CuP connected more distad. Veinlets rp-mp slightly apicad of mp-cua at about same level, apicad of CuA 2 forking, cua-cup more basal, apicad of CuA forking. Male terminalia (Figs 18–29). Pygofer asymmetrical, roughly triangular in lateral view, upper margin declivous, lower margin straight; with posterodorsal elongate process on right side, left side without process; dorsal margin not excavate at level of anal tube base; ventral margin deeply, arcuately excavate. Gonostyli asymmetrical, medially fused into a plate, convex medially in ventral plane, in lateral view left gonostylus elongate, with subapical triangular process, dorsal edge with an oblique hook-like process at about half of gonostylus length, directed cephalad; in lateral view right gonostylus subtriangular, caudodorsal margin arcuate, dorsal margin without hook. Phallic complex straight caudad; periandrium tubular, short, membranous, endosoma bulbous, spiniferous, with three asymmetric distinct spines. Anal tube elongate, tubular, reaching almost to apex of gonostyli; with ventrocaudal, triangular expansion, epiproct and paraproct of similar size. Female terminalia (Figs 30–46). Pregenital sternite narrow, posterior margin medially deeply concave, with median sclerotised process; anal tube distinctly shorter than gonoplac, tubular; gonoplac unilobate, with row of distinct teeth on posterior margin; bases of gonapophyses VIII asymmetrical—left gonapophysis base lobate, with hook-like process; right base of gonapophysis lobate, smooth; gonapophysis VIII with teeth on apical portion of dorsal margin, ventral margin arcuate, with three apical teeth; endogonocoxal process sabre-like, as long as gonapophysis VIII. Type material. Holotype, male: [D.N. Guinea 150. / Standlager a. Aprilfluss / 183, 12.-14.IX.1912 / Kais. Augustafl. Exp. / Bürgers S.G.] (MNHU). Paratypes, 2 males, 1 female: [D.N. Guinea 150. / Standlager a. Aprilfluss / 183, 12.-14.IX.1912 / Kais. Augustafl. Exp. / Bürgers S.G.] —male (MNHU); [D.N. Guinea / Regenberg 550 m / 8.-15. V.13 / Kais. Augustafl. Exp. / Bürgers S.G.], [331 / 9.X.13.] —male (MIZ); [D. N. Guinea / Standlager b. Malu / 12.–13.III.1912 / Dr. Bürgers S.G.] —female (MNHU). Distribution. New Guinea, Papua New Guinea, Madang Province.

A new genus and species of the tropiduchid planthopper of the tribe Tropiduchini is described as Bitara augusta gen. et sp. nov. from Madang Province, Papua New Guinea. It is the 18th species of Tropiduchini and the 9th genus known east of the Wallace line. It is another taxon with characteristic asymmetric genitalia within this tribe.

Ricanula peronata sp. nov. (Figs 7–9, 11) Etymology. The name is derived from the Latin word ‘ peronatus ’, referring to the periandrium with boot-shaped ventral processes in ventral view. Diagnosis. The species is similar to Ricanula curva sp. nov., but differs from the latter by having periandrium with boot-shaped ventral processes in ventral view (Fig. 9F); ventral processes of aedeagus oriented ventrally in lateral view (Fig. 9G) (apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view; ventral processes of aedeagus S-curved in lateral view in Ricanula curva sp. nov.). Description. Measurements. Length (inc. tegmen): male 6.5–7.4 mm, female 7.3–8.8 mm. Head. Vertex (Figs 7A, C) without median carina. Frons: median and lateral carinae of frontal disc surpassing half of disc, ending about the level of antennae; apical parts of median carina weakly visible. Thorax. Pronotum (Figs 7A, C) with small round depressions submedially on each lateral side. Mesonotum: lateral carinae (Figs 7A, C) not reaching posterior margin; antero-lateral carinae not connected with anterior margin. Tegmen: postero-apical part of tegmen with two eye-spot black cells, posterior margin arcuate. Longitudinal veins ScP+RA and RP, MP arising as short common stem from basal cell. Claval veins Pcu and A 1 fused on midlength of CuP vein. Hind wing (Fig. 7F) without transverse veinlets. Hing legs: Basitarsomere of metatarsus with 8 apical teeth. Metatibiotarsal formula 2/6/8. Male terminalia. Anal tube (in dorsal view, Fig. 8B) nearly square, posterior margin strongly concave, basal margin slightly convex, lateral margins straight; anus placed before midlength, paraproct surpassing the posterior margin. Pygofer (in lateral view, Fig. 8A) with dorsal posterior angle without process. Genital styles (in lateral view, Fig. 8A) broadly triangular; ventral margin weakly sinuate; dorsal margin weakly convex, with small concavity before spine-like process. Phallic complex (Figs 9A–C): Periandrium (Figs 9D–F) with boot-shaped ventral processes in ventral view, apical part of ventral processes straight; dorsal periandrium with U-shaped structure with membranous apical part sclerotized base in dorsal view; lateral margin of periandrium with small rod-shaped processes in lateral view, rodshaped processes hidden in the periandrium (in ventral view); ventral periandrium distinctly convex. Aedeagus (Figs 9G–I) apically with two pairs of processes. Median split asymmetrical: ventral split present only in 1/5; dorsal split very deep, reaching almost basal part. All processes single armed: lateral processes longer than ventral processes, about two thirds of aedeagus; ventral processes oriented ventrally in lateral view. Female terminalia (Figs 8C–I). Pregenital sternite (Fig. 8I): posterior margin medially with two prominent processes, margin between processes with wide and shallow incision. Anal tube (in dorsal view, Fig. 8C) ovoid, with widest part medially, basal margin weakly convex, posterior margin widely concave, lateral margins arcuate; anus placed after midlength, paraproct surpassing the posterior margin. Gonoplac (Fig. 8H): posterior margin with two rows of small teeth. Coloration. General color brown (Figs 7A–B). Median part of frons (Fig. 7D) black brown. Eyes brown (Figs 7B–C), ornamented with irregular brown patches. Gena (Fig. 7B) black brown with two yellowish spots. Tegmen (Figs 7B, E) brown, costal margin with about 14 transverse black brown stripes from base to a little beyond middle, between the transverse brown stripes filled with light yellow stripes, sub-medially of tegmen with a large flavescent spot marked by 2 central transverse back lines. Wings brown, each side of A 2 with a longitudinal grayish narrowed band (Fig. 7F). Abdomen and terminalia brown. Type material. Holotype, male, China: 20 Apr. 1964, Yunnan, Xishuangbanna, Menglun, 650 m, coll. Baolin Zhang. Paratypes (8 males, 17 females, China): 1 male: 13 Jul. 1958, 650– 700 m, coll. Chunpei Hong; 2 males, 4 females: 21/ 30 Apr. 1974, coll. Yao Chou, Feng Yuan et Yinyue Hu; 1 female: 22 May 1982, 1 female: 24 May 1982, coll. Qinmei Wang et Jingruo Zhou; 1 male, 1 female: 18 May 1991, 1 female: 20 May 1991, 1 male, 1 female: 22 May 1991, 2 females: 26 May 1991, 3 males, 6 females: 31 May 1991, coll. Yinglun Wang et Wanzhi Cai; Yunnan, Xishuangbanna, Menglun. Distribution. China (Province Yunnan)., Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on pages 363-366, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953

Ricanula pulverosa (Stål, 1865) (Figs 10–11) Ricania pulverosa Stål, 1865: 162; Stål, 1870: 767; Atkinson, 1886: 61; Noualhier, 1896: 256; Schmidt, 1905: 176; Distant, 1906: 380; Schumacher, 1915: 130; Melichar, 1898: 240; Esaki, 1932: 1805; Kato, 1933: 7; Zia, 1935: 537; Lallemand, 1942: 70; Jacobi, 1944: 22; Chou et al., 1985: 81. Ricanula pulverosa, Melichar, 1923: 130; Metcalf, 1955: 97; Yang, 1989: 193. Specimens examined. 1 male: 1 Jun. 1974, Yunnan, Xishuangbanna, Mengla, coll. Yao Chou et Feng Yuan; 1 male: 18 Apr. 1982, Yunnan, Xishuangbanna, Mengla, coll. Sumei Wang et Jingruo Zhou. Measurements. Length (inc. tegmen): male 7.8 mm. Remarks. First time after original description, Yang (1989) described and illustrated the male and female genitalia of the species based on Taiwan specimens. The morphological characters examined here are consistent with Yang’s (1989) description, but the male genitalia in this study show somewhat intraspecific variations as follows: 1) anal tube nearly rectangular in dorsal view (Fig. 10G) (irregularly hexagonal in Yang, 1989, Fig. 12G); 2) lateral processes of aedeagus reaching only 1/4 of the dorsal processes (Fig. 10I) (lateral processes reaching 1/2 of the dorsal processes in Yang, 1989, Figs 12H, I). Distribution. China (Provinces Shaanxi, Zhejiang, Yunnan, Fujian, Hainan, Taiwan), Cambodia, Myanmar, Thailand, Vietnam, Japan, India, Indonesia., Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on page 366, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953, {"references":["Stal, C. (1865) Homoptera nova vel minus congita. Ofversigt af Kongliga Svenska Vetenskaps-Akadamiens Forhandlingar, 22, 145 - 165.","Stal, C. (1870) Hemiptera insularum Philippinarum. Bidrag till Philippinska oarnes Hemipter-fauna. Ofversigt af Kongliga Svenska Vetenskaps-Akademiens Forhandlingar, 27, 607 - 776. https: // doi. org / 10.5962 / bhl. title. 61898","Atkinson, E. T. (1886) Notes on Indian Rhynchota, No. 6. Journal and Proceedings of the Asiatic Society of Bengal, 55, 143 - 223.","Noualhier, J. M. (1896) Note sur les Hemipteres recoltes en Indo-Chine et offerts au Museum par M. Pavie. Bulletin du Museum national d'histoire naturelle, 10, 251 - 259.","Schmidt, E. (1905) Die Ricaniiden des Stettiner Museums. Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin, 66, 168 - 198.","Distant, W. L. (1906) The fauna of British India, including Ceylon and Burma. Rhynchota. Vol. III (Heteroptera-Homoptera). Taylor and Francis, London, 503 pp.","Schumacher, F. (1915) Homoptera in H. Sauter's Formosa-Ausbeute. Supplementa Entomologica, 4, 108 - 142.","Melichar, L. (1898) Monographie der Ricaniiden (Homoptera). Annalen des K. K. Naturhistorischen Hofmuseums, 8 (2 - 3), 197 - 359.","Kato, M. (1933) Three colour illustrated insects of Japan. Fascicle 4. Homoptera: Fulgoridae and Others. Koseikaku, Tokyo, 127 pp.","Zia, Y. (1935) Note sur les Flatinae et les Ricaniinae de la chine du sud et du Tonkin (Homoptera Fulgoridae). Sinensia, 6 (5), 525 - 540.","Lallemand, V. (1942) Notes sur quelques especes recueillies par le R. Piel (Musee Heude, Shanghai) et le R. P. de Cooman (Hoa Binh, Tonkin). Notes d'Entomologie Chinoise. Musee Heude, 9 (4), 69 - 77.","Jacobi, A. (1944) Die Zikadenfauna der Provinz Fukien in Sudchina und ihre tiergeographischen Beziehungen. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 5 - 66.","Chou, I., Lu, J. S., Huang, J. & Wang, S. Z. (1985) Homoptera, Fulgoroidea. Economic Insect Fauna of China. Fasc. 36. Science Press, Beijing, 152 pp.","Melichar, L. (1923) Homoptera, fam. Acanaloniidae, Flatidae et Ricaniidae. Genera Insectorum. Part 182. L. Desmet-Verteneuil, Bruxelles, 185 pp.","Metcalf, Z. P. (1955) General catalogue of the Hemiptera. Fascicle IV. Fulgoroidea. Part 16. Ricaniidae. Smith College, Northampton, Massachusetts, 199 pp.","Yang, C. T. (1989) Ricaniidae of Taiwan (Homoptera: Fulgoroidea). In: Collected Papers on Fulgoroidea of Taiwan. Taiwan Museum Special Publication Series, 8, pp. 171 - 204."]}

Ricanula Melichar, 1898 Ricania (Ricanula) Melichar, 1898: 218. Ricanula Schmidt, 1912: 75. Type species. Ricania noualhieri Melichar, 1898, designated by Schmidt (1912). Remarks. Ricanula can be distinguished from other genera in Ricaniidae by the combination of following characters: costal area of tegmen with sparse and curved transverse veinlets; postero-apical part of tegmen with eyes-spot black cells; tegmen with one line of transverse veinlets; median and posterior parts of tegmen with numerous irregular transverse veinlets. Diagnostic characters. Head. Head including eyes narrower than mesonotum/thorax. Vertex transverse, distinctly wider than long at midline, with all margins well carinate; disc of vertex with or without median carina. Frons with all margins well carinated, at upper margin longer than high at midline, widest at the level of lower margin of compound eyes; lateral margins covering base of pedicel, not incised near the level of ocelli. Frontal disc tricarinate, carinae distinctly separated basally, median carina straight; lateral carinae arcuate, almost parallel to lateral margins. Fronto-clypeal suture arched/arcuate. Clypeus distinctly narrower than frons, without median carina. Compound eyes, with small callus in lower part of posterior margin. Ocelli present. Rostrum-apical segment shorter than subapical one. Antenna pedicel elongate, cylindrical, with functional area at the top. Thorax. Pronotum distinctly longer than vertex at midline; disc of pronotum with median carina and two lateral impressions. Mesonotum elongate, diamond shape, longer in midlength than wide at lateral angles and longer in midlength longer than combined length of vertex and pronotum at midlength; lateral angles placed before midlength; median carina, lateral and antero-lateral carinae present; median carina and lateral carinae connected basally; median carina reaching scutellum, lateral carinae reaching posterior margin, anterolateral carinae not connected with lateral and not surpassing level of lateral angles of mesonotum. Tegmina membranous, elongately-triangular, flattened, with distinct venation and transverse veinlets. Costal margin weakly arcuate, apical angle broadly rounded, placed distad to claval angle, posterior margin weakly sinuate, postclaval margin (tornus) absent. Costal area with transverse veinlets and oblique shallowly incisions; postero-apical part with eyes-spot black cells. Longitudinal veins ScP+RA, MP and CuA leaving basal cell separated. CuA vein first fork placed before the connection of claval veins Pcu+A 1. Tegmen with single apical line of transverse veinlets, median and posterior part of tegmen with numerous irregular transverse veinlets. Hind wing with precostal cell present; ScRA and MP forking distinctly after midlength of wing, CuA forking distinctly before half of wing. Hing legs. Metatibia with 2 lateral spines; apically with 6 well developed spines, external lateral spines bigger than internal lateral spines; basitarsomere of metatarsus a little longer than cumulative length of second and apical tarsomeres. Male terminalia. Anal tube with ventral margin strongly concaved in lateral view. Pygofer higher than wide; dorsal part narrower than ventral one. Genital styles broadly triangular (dorsal and ventral margins nearly parallel in R. fujianensis and R. cacaonis), with sharp spine-like process at the end of dorsal margin. Periandrium with or without processes, with long lateral split surpassing the half of its length; dorsal periandrium a bit shorter than ventral one. Aedeagus long and narrow, apically with 1–2 pairs of symmetrical, well sclerotized, spinose processes. Female terminalia. Pregenital sternite with well-developed lateral lobes, median portion of the posterior margin with or without processes. Anal tube not surpassing half of upper margin of the gonoplac. Gonoplac well developed, laterally flattened; posterior margin of the gonoplac with 2–3 rows of small teeth; membranous parts of gonoplac well developed, placed medially on ventral margin. Gonapophysis VIII sabre-like, v-shape in cross section, with teeth at dorsal margin; endogonocoxal process with spiniferous microsculptures, well sclerotized medially; lateral parts membranous, reaching apex of gonapophysis VIII. Gonaphophyses IX and gonospiculum bridge well developed. Bursa copulatrix with two isometric pouches: first pouch with well visible cell and sclerotized ornamentation; second pouch with numerous pores., Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on pages 354-355, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953, {"references":["Melichar, L. (1898) Monographie der Ricaniiden (Homoptera). Annalen des K. K. Naturhistorischen Hofmuseums, 8 (2 - 3), 197 - 359.","Schmidt, E. (1912) Diagnosen neuer Fulgoriden-Gattungen und Arten nebst einigen Bemerkungen. Stettiner Entomologische Zeitung, 73, 67 - 102."]}

Ricanula peronata sp. nov. (Figs 7–9, 11) Etymology. The name is derived from the Latin word ‘ peronatus ’, referring to the periandrium with boot-shaped ventral processes in ventral view. Diagnosis. The species is similar to Ricanula curva sp. nov., but differs from the latter by having periandrium with boot-shaped ventral processes in ventral view (Fig. 9F); ventral processes of aedeagus oriented ventrally in lateral view (Fig. 9G) (apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view; ventral processes of aedeagus S-curved in lateral view in Ricanula curva sp. nov.). Description. Measurements. Length (inc. tegmen): male 6.5–7.4 mm, female 7.3–8.8 mm. Head. Vertex (Figs 7A, C) without median carina. Frons: median and lateral carinae of frontal disc surpassing half of disc, ending about the level of antennae; apical parts of median carina weakly visible. Thorax. Pronotum (Figs 7A, C) with small round depressions submedially on each lateral side. Mesonotum: lateral carinae (Figs 7A, C) not reaching posterior margin; antero-lateral carinae not connected with anterior margin. Tegmen: postero-apical part of tegmen with two eye-spot black cells, posterior margin arcuate. Longitudinal veins ScP+RA and RP, MP arising as short common stem from basal cell. Claval veins Pcu and A 1 fused on midlength of CuP vein. Hind wing (Fig. 7F) without transverse veinlets. Hing legs: Basitarsomere of metatarsus with 8 apical teeth. Metatibiotarsal formula 2/6/8. Male terminalia. Anal tube (in dorsal view, Fig. 8B) nearly square, posterior margin strongly concave, basal margin slightly convex, lateral margins straight; anus placed before midlength, paraproct surpassing the posterior margin. Pygofer (in lateral view, Fig. 8A) with dorsal posterior angle without process. Genital styles (in lateral view, Fig. 8A) broadly triangular; ventral margin weakly sinuate; dorsal margin weakly convex, with small concavity before spine-like process. Phallic complex (Figs 9A–C): Periandrium (Figs 9D–F) with boot-shaped ventral processes in ventral view, apical part of ventral processes straight; dorsal periandrium with U-shaped structure with membranous apical part sclerotized base in dorsal view; lateral margin of periandrium with small rod-shaped processes in lateral view, rodshaped processes hidden in the periandrium (in ventral view); ventral periandrium distinctly convex. Aedeagus (Figs 9G–I) apically with two pairs of processes. Median split asymmetrical: ventral split present only in 1/5; dorsal split very deep, reaching almost basal part. All processes single armed: lateral processes longer than ventral processes, about two thirds of aedeagus; ventral processes oriented ventrally in lateral view. Female terminalia (Figs 8C–I). Pregenital sternite (Fig. 8I): posterior margin medially with two prominent processes, margin between processes with wide and shallow incision. Anal tube (in dorsal view, Fig. 8C) ovoid, with widest part medially, basal margin weakly convex, posterior margin widely concave, lateral margins arcuate; anus placed after midlength, paraproct surpassing the posterior margin. Gonoplac (Fig. 8H): posterior margin with two rows of small teeth. Coloration. General color brown (Figs 7A–B). Median part of frons (Fig. 7D) black brown. Eyes brown (Figs 7B–C), ornamented with irregular brown patches. Gena (Fig. 7B) black brown with two yellowish spots. Tegmen (Figs 7B, E) brown, costal margin with about 14 transverse black brown stripes from base to a little beyond middle, between the transverse brown stripes filled with light yellow stripes, sub-medially of tegmen with a large flavescent spot marked by 2 central transverse back lines. Wings brown, each side of A 2 with a longitudinal grayish narrowed band (Fig. 7F). Abdomen and terminalia brown. Type material. Holotype, male, China: 20 Apr. 1964, Yunnan, Xishuangbanna, Menglun, 650 m, coll. Baolin Zhang. Paratypes (8 males, 17 females, China): 1 male: 13 Jul. 1958, 650– 700 m, coll. Chunpei Hong; 2 males, 4 females: 21/ 30 Apr. 1974, coll. Yao Chou, Feng Yuan et Yinyue Hu; 1 female: 22 May 1982, 1 female: 24 May 1982, coll. Qinmei Wang et Jingruo Zhou; 1 male, 1 female: 18 May 1991, 1 female: 20 May 1991, 1 male, 1 female: 22 May 1991, 2 females: 26 May 1991, 3 males, 6 females: 31 May 1991, coll. Yinglun Wang et Wanzhi Cai; Yunnan, Xishuangbanna, Menglun. Distribution. China (Province Yunnan).

Ricanula curva sp. nov. (Figs 4–6, 11) Etymology. The name is derived from the Latin word ‘ curvus ’, referring to ventral processes of aedeagus being Scurved in lateral view; apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view. Diagnosis. The species is similar to R. peronata sp. nov., but differs from the latter by having apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view (R. peronata —ventral processes boot-shaped in ventral view); ventral processes of aedeagus S-curved in lateral view (R. peronata —ventral processes of aedeagus oriented ventrally in lateral view). Description. Measurements. Length (inclu. teg.): male 6.7–8.4 mm, female 7.2–9.6 mm. Head. Vertex (Figs 4A, C) with or without median carina. Frons: lateral carinae of frontal disc finishing basally at the same level as median carina, ending about the level of antennae. Thorax. Pronotum (Figs 4A, C) with small round depressions submedially on each lateral side (in some specimens weakly visible). Mesonotum: lateral carinae of mesonotum (Figs 4A, C) reaching posterior margin; anterolateral carinae connected with anterior margin. Tegmen: postero-apical part of tegmen with two eye-spot black cells, posterior margin almost straight. Longitudinal veins ScRA and RP arising as short common stem from basal cell, MP veins forked on basal cell. Claval veins Pcu and A 1 fused before midlength of CuP vein. Hind wing with r-m transverse veinlets present in distal part of wing (Fig. 4F). Hing legs: Basitarsomere of metatarsus with 8 apical teeth. Metatibiotarsal formula 2/6/8. Male terminalia (Figs 5A–B, 6A–I). Anal tube (in dorsal view, Fig. 5B) nearly as rectangle; posterior margin strongly concave, basal margin slightly convex, lateral margins straight; anus placed before midlength, paraproct slightly surpassing the posterior margin. Pygofer (in lateral view, Fig. 5A): dorso-posterior angle with process. Genital styles (Fig. 5A) broadly triangular in lateral view; ventral margin convex; dorsal margin weakly convex, with small concavity before spine-like process. Phallic complex (Figs 6A–C): Periandrium (Figs 6D–F) with ventral processes, apical part of ventral processes of periandrium curved laterally (in dorsal and ventral view), ventral periandrium distinctly convex in middle; dorsal periandrium with U-shaped structure with membranous apical part sclerotized base in dorsal view; lateral margin of periandrium with rod-shaped processes in lateral view. Aedeagus (Figs 6G–I) with two pairs of processes. Median split asymmetrical: ventral split present only in 1/5; dorsal split very deep, reaching almost basal part. All processes single armed: lateral processes longer than ventral processes, about 2/3 of aedeagus, curved dorsally at 2/3 of its length; ventral processes S-curved ventrally in lateral view. Female terminalia (Figs 5C–I). Pregenital sternite: posterior margin medially with two processes, margin between processes with strong and deep incision (Fig. 5I). Anal tube (in dorsal view, Fig. 5C) elongate, with posterior part wider than basal one; basal margin almost straight, posterior margin widely concave medially, lateral margins arcuate; anus placed a bit before midlength, paraproct surpassing the posterior margin. Gonoplac posterior margin with two rows of teeth. Coloration. General color brown to dark brown (Figs 4A–B). Lateral margins of frons yellow, area alongside frontoclypeal suture yellow, clypeus brown with yellow patch medially under frontoclypeal suture, rostrum yellowish with brown apex (Fig. 4D). Eyes (Fig. 4B) sordid brown, ornamented with irregular black brown patches. Gena (Fig. 4B) brown with two yellow spots. Tegmen (Figs 4A–B, E) brown to dark brown; costal margin with about 13–16 transverse brown stripes from base to a little beyond middle, between the transverse brown stripes filled with light yellow stripes, tegmen sub-medially with a large flavescent spot marked by 2 central transverse brown lines. Wings (Fig. 4F) brown, each side of A 2 with a grayish narrowed band longitudinally. Abdomen and terminalia brown. Type material. Holotype, male, China: 28 May 1982, Guangxi, Tianlin, Langping, coll. Jikun Yang. Paratypes (13 males, 21 females, China): 4 males, 2 females: 11 Apr. 1978, Guangxi, Baise, Yangwei, coll. Xianyu Qian; 1 male: 12 May 1980, Guangxi, Longzhou, Longhu, coll. Zhuyin Wang; 1 male: 17 May 1982, Guangxi, Longzhou, Nonggang, 240m, coll. Fasheng Li; 2 males: 29 May 1982, 1 female: 30 May 1982, Guangxi, Tianlin, Langping, coll. Jikun Yang; 1 female: 25 Jun. 1982, Guangxi, Longsheng, coll. Jikun Yang; 3 males, 14 females: 3 Jun. 1984, 1 female: 9 Jun. 1984, Guangxi, Lingtian commune, coll. Zhengliang Wu et Xiaolin Lu; 1 male, 1 female: 6 May 1993, 1 male, 1 female: 8 May 1993, Guangxi, Longzhou, Daqing Mountain, coll. Sikong Liu. Distribution. China (Province Guangxi)., Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on pages 359-363, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953

Two new species of the genus Ricanula Melichar, 1898 within the family Ricaniidae (Hemiptera: Fulgoromorpha) from China are described and illustrated: R. curva sp. nov. and R. peronata sp. nov. Ricania cacaonis Chou et Lu, 1977 is redescribed and transferred to genus Ricanula. Ricanula pulverosa (Stål, 1865) is re-illustrated.

Ricanula curva sp. nov. (Figs 4–6, 11) Etymology. The name is derived from the Latin word ‘ curvus ’, referring to ventral processes of aedeagus being Scurved in lateral view; apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view. Diagnosis. The species is similar to R. peronata sp. nov., but differs from the latter by having apical part of ventral processes of periandrium strongly curved laterally in dorsal and ventral view (R. peronata —ventral processes boot-shaped in ventral view); ventral processes of aedeagus S-curved in lateral view (R. peronata —ventral processes of aedeagus oriented ventrally in lateral view). Description. Measurements. Length (inclu. teg.): male 6.7–8.4 mm, female 7.2–9.6 mm. Head. Vertex (Figs 4A, C) with or without median carina. Frons: lateral carinae of frontal disc finishing basally at the same level as median carina, ending about the level of antennae. Thorax. Pronotum (Figs 4A, C) with small round depressions submedially on each lateral side (in some specimens weakly visible). Mesonotum: lateral carinae of mesonotum (Figs 4A, C) reaching posterior margin; anterolateral carinae connected with anterior margin. Tegmen: postero-apical part of tegmen with two eye-spot black cells, posterior margin almost straight. Longitudinal veins ScRA and RP arising as short common stem from basal cell, MP veins forked on basal cell. Claval veins Pcu and A 1 fused before midlength of CuP vein. Hind wing with r-m transverse veinlets present in distal part of wing (Fig. 4F). Hing legs: Basitarsomere of metatarsus with 8 apical teeth. Metatibiotarsal formula 2/6/8. Male terminalia (Figs 5A–B, 6A–I). Anal tube (in dorsal view, Fig. 5B) nearly as rectangle; posterior margin strongly concave, basal margin slightly convex, lateral margins straight; anus placed before midlength, paraproct slightly surpassing the posterior margin. Pygofer (in lateral view, Fig. 5A): dorso-posterior angle with process. Genital styles (Fig. 5A) broadly triangular in lateral view; ventral margin convex; dorsal margin weakly convex, with small concavity before spine-like process. Phallic complex (Figs 6A–C): Periandrium (Figs 6D–F) with ventral processes, apical part of ventral processes of periandrium curved laterally (in dorsal and ventral view), ventral periandrium distinctly convex in middle; dorsal periandrium with U-shaped structure with membranous apical part sclerotized base in dorsal view; lateral margin of periandrium with rod-shaped processes in lateral view. Aedeagus (Figs 6G–I) with two pairs of processes. Median split asymmetrical: ventral split present only in 1/5; dorsal split very deep, reaching almost basal part. All processes single armed: lateral processes longer than ventral processes, about 2/3 of aedeagus, curved dorsally at 2/3 of its length; ventral processes S-curved ventrally in lateral view. Female terminalia (Figs 5C–I). Pregenital sternite: posterior margin medially with two processes, margin between processes with strong and deep incision (Fig. 5I). Anal tube (in dorsal view, Fig. 5C) elongate, with posterior part wider than basal one; basal margin almost straight, posterior margin widely concave medially, lateral margins arcuate; anus placed a bit before midlength, paraproct surpassing the posterior margin. Gonoplac posterior margin with two rows of teeth. Coloration. General color brown to dark brown (Figs 4A–B). Lateral margins of frons yellow, area alongside frontoclypeal suture yellow, clypeus brown with yellow patch medially under frontoclypeal suture, rostrum yellowish with brown apex (Fig. 4D). Eyes (Fig. 4B) sordid brown, ornamented with irregular black brown patches. Gena (Fig. 4B) brown with two yellow spots. Tegmen (Figs 4A–B, E) brown to dark brown; costal margin with about 13–16 transverse brown stripes from base to a little beyond middle, between the transverse brown stripes filled with light yellow stripes, tegmen sub-medially with a large flavescent spot marked by 2 central transverse brown lines. Wings (Fig. 4F) brown, each side of A 2 with a grayish narrowed band longitudinally. Abdomen and terminalia brown. Type material. Holotype, male, China: 28 May 1982, Guangxi, Tianlin, Langping, coll. Jikun Yang. Paratypes (13 males, 21 females, China): 4 males, 2 females: 11 Apr. 1978, Guangxi, Baise, Yangwei, coll. Xianyu Qian; 1 male: 12 May 1980, Guangxi, Longzhou, Longhu, coll. Zhuyin Wang; 1 male: 17 May 1982, Guangxi, Longzhou, Nonggang, 240m, coll. Fasheng Li; 2 males: 29 May 1982, 1 female: 30 May 1982, Guangxi, Tianlin, Langping, coll. Jikun Yang; 1 female: 25 Jun. 1982, Guangxi, Longsheng, coll. Jikun Yang; 3 males, 14 females: 3 Jun. 1984, 1 female: 9 Jun. 1984, Guangxi, Lingtian commune, coll. Zhengliang Wu et Xiaolin Lu; 1 male, 1 female: 6 May 1993, 1 male, 1 female: 8 May 1993, Guangxi, Longzhou, Daqing Mountain, coll. Sikong Liu. Distribution. China (Province Guangxi).

Sap-sucking hemipterans host specialized, heritable microorganisms that supplement their unbalanced diet with essential nutrients. These microbes show unusual features that provide a unique perspective on the evolution of life but have not been systematically studied. Here, we combine microscopy with high-throughput sequencing to revisit 80-year-old reports on the diversity of symbiont transmission modes in a broadly distributed planthopper family Dictyopharidae. We show that in all species examined, the ancestral nutritional endosymbionts Sulcia and Vidania are complemented by co-primary symbionts, either Arsenophonus or Sodalis, acquired several times independently by different host lineages. Like in other obligate sap-feeders, the ancestral symbionts produce essential amino acids, whereas co-primary symbionts contribute to the biosynthesis of B vitamins. These symbionts reside within separate bacteriomes within the abdominal cavity, although in females, Vidania also occupies bacteriocytes in the rectal organ. Notably, the symbionts are transmitted from mothers to offspring in two alternative ways. In most examined species, all nutritional symbionts simultaneously infect the posterior end of the full-grown (vitellogenic) oocytes and next gather in their perivitelline space. In contrast, in other species, Sodalis colonizes the cytoplasm of the anterior pole of young (previtellogenic) oocytes forming a cluster separate from the “symbiont ball” formed by late-invading Sulcia and Vidania. Our data add to evidence on frequent replacements of gammaproteobacterial symbionts combined with the relative functional stability of the nutritional functions during the evolution of sap-feeding insects, and show how newly-arriving microbes may utilize different strategies to establish long-term heritable symbiosis.Significance statementSup-sucking hemipterans host ancient heritable microorganisms that supplement their unbalanced diet with essential nutrients, and which have repeatedly been complemented or replaced by other microorganisms. They need to be reliably transmitted to subsequent generations through the reproductive system, and often they end up using the same route as the ancient symbionts. We show for the first time that in a single family of planthoppers, the complementing symbionts that have established infections independently utilize different transmission strategies, one of them novel, with the transmission of different microbes separated spatially and temporarily. These data show how newly-arriving microbes may utilize different strategies to establish long-term heritable symbiosis.

In communicating resource systems (CRS), a set of independent, hierarchically composed resources communicate to realize a business process. Examples of CRSs include RESTful Web services, cloud computing platforms and hierarchical distributed systems. Unfortunately, their complexity, distributed nature, and usually long lifespan can lead to problems like livelocks, deadlock occurrences and invocation loops. This makes CRSs hard to maintain and manage; it also leads to a loss of QoS or system failures. Moreover, it is important to take into account, in system management, the exact way how the system gets to some desired or undesired state. Therefore, it is crucial to analyze processes, executed in a system to find and fix the above problems. Unfortunately, CRS process models are hardly ever available. Therefore, we present a new distributed algorithm, dRMA, discovering process models from CRS event logs. In order to reduce the representational bias between the discovered process model and the real process, we have defined a Communication Net, which can express features of CRSs: directed communication channels, hierarchy, and a resource perspective. Finally, we have evaluated dRMA on a cluster, and have showed that distributing jobs can significantly reduce the time of process model discovery.

Two new species of ricaniid planthopper Ricanoides (Hemiptera: Fulgoromorpha: Ricaniidae) are described and illustrated: Ricanoides phiaoacensissp. nov. from Vietnam and Ricanoides srenoiensissp. nov. from Cambodia. Male and female external and internal genital structures of the new species are illustrated and discussed. Key for the identification of all species belonging to the genus based on male genital structures is given.

A new monotypic genus of flatid planthoppers (Hemiptera: Fulgoromorpha: Flatidae), Selizitapia gen. nov., is described for Selizitapia pennyi gen. et sp. nov. (type species) from the island of Madagascar. Habitus, male and female external and internal genital structures of the new species are illustrated and compared with similar taxa. Selizitapia pennyi gen. et sp. nov. is endemic to Madagascar where it is known to date only from one locality in the Central Plateau and is associated with tapia woodland formation.

Selizitapia pennyi gen. et sp. nov. urn:lsid:zoobank.org:act: FE9BEDC2-9BD1-4E72-8E70-5E91C943E49E Figs 1–7 Diagnosis The only species in the genus. Etymology The species is dedicated to Dr Norm Penny (1946–2016) – outstanding researcher of Neuropterida, and collection manager at the Department of Entomology at the California Academy of Sciences, San Francisco, California. He kindly provided us with invaluable material of Madagascan Flatidae making the previous and current studies possible. Material examined Holotype MADAGASCAR: ♂; “ MADAGASCAR, Province / Fianarantsoa, Italaviana, / 35 km SSE of Antsirabe / 20º10.40’S, 47º05.16’E / 8-24 April 2004 ”; “ California Acad of Sciences / coll: M. Irwin, R. Harin- Hala / malaise trap – in Uapaca / forest elev 1360 m / MA-24-45”; “CASLOT 044412”; “ HOLOTYPE ”; “ Selizitapia pennyi sp. nov. / Świerczewski & Stroiński det.”; CAS (dry-mounted, abdomen detached, dissected and stored in glycerol in a glass microvial under the specimen). Paratypes All specimens with the locality collection data the same as the holotype apart from the collection codes and the dates of collecting. All dry-mounted, abdomens of some specimens detached, dissected and stored in glycerol in a glass microvial under the specimen. MADAGASCAR – Fianarantsoa Province, Italaviana • 1 ♀; 9–19 Dec. 2002; MA-24-02; CASLOT 044446; CAS • 1 ♂; 30 Apr.–11 May 2003; MA-24-15; CASLOT 044453; CAS • 1 ♂; 28 Jan.–13 Feb. 2005; MA-24-57; CASLOT 044465; CAS • 1 ♀; 28 Jan.–13 Feb. 2005; MA-24-67; CASLOT 044465; CAS • 1 ♀; 30 Apr.–11 May 2003; MA-24-15, CASLOT 044463; MIZ • 1 ♀; 2–14 Oct. 2004; MA-24- 57; CASLOT 044485; CAS • 1 ♂; 30 Mar.–9 Apr. 2003; MA-24-12, CASLOT 044507; MIZ • 1 ♀; 30 Mar.–9 Apr. 2003; MA-24-12; CASLOT 044507; CAS • 1 ♀; 10–20 Mar. 2003; MA-24-10; CASLOT 044521; CAS • 1 ♀; 17–27 Sep. 2003; MA-24-29; CASLOT 044527; CAS • 1 ♀; 16–26 Oct. 2003; MA-24-32; CASLOT 044596; CAS. Description SIZE. Total length 6.8–7.0 mm. COLORATION. Mostly uniformly stramineous with costal area, apical cells and postclaval margin fuscous; dark brown markings between basal part MP and CuA; teeth of gonoplac dark brown (Fig. 1A–C). HEAD. Vertex: A/B = 16.67. Frons: C/E = 1.00; D/E = 1.20. THORAX. Pronotum: F/B = 8.33; mesonotum: G/F = 3.20; G/B+F = 2.86; G/H = 0.89; tegmen: I/J = 3.67; I/K = 2.39. MALE TERMINALIA. Anal tube, in lateral view, with ventral margin as obtuse angle and dorsal margin arcuate (Fig. 5A). Genital style with posterior margin weakly convex, ventral and dorsal margins almost straight, postero-ventral angle bluntly rounded, not extending to the posterior margin (Fig. 5A). Basal lobes of ventral periandrium small with sinuate margin (Fig. 5D). FEMALE TERMINALIA. Pregenital sternite with anterior and posterior margins in median portion almost straight (Fig. 7A). Anal tube with apical margin, in dorsal view, strongly rounded (Fig. 7B); in lateral view, with basal part wider than apical part, anus placed almost at midlength (Fig. 7C). Gonoplacs with membranous part at ventral margin (Fig. 7D). Gonapophysis VIII with subapical 5 massive teeth at dorsal margin (Fig. 7E). Ductus receptaculi and diverticulum ductus widened distally (Fig. 7G). Distribution and habitat Madagascar; so far only known from one locality in the central part of the island (35 km SSE of Antsirabe). The type series was collected in a tapia woodland formation, in the season of 2003 from March to May and from September to October., Published as part of Świerczewski, Dariusz & Stroiński, Adam, 2021, Selizitapia gen. nov. (Hemiptera: Fulgoromorpha: Flatidae) from tapia woodlands of Madagascar, pp. 124-139 in European Journal of Taxonomy 750 (750) on pages 128-136, DOI: 10.5852/ejt.2021.750.1367, http://zenodo.org/record/5451685

Selizitapia gen. nov. urn:lsid:zoobank.org:act: 7CB34782-C76D-43AE-808C-4911F4CB0A2E Figs 1–7 Type species Selizitapia pennyi gen. et sp. nov., here designated. Diagnosis The new genus differs from other taxa of Selizini in Madagascar by the following characters: 1) wings subrectangular (wings strongly constricted apically in Stenocyarda Fennah, 1965); 2) frons with median carina not diverged (frons with Y-shaped median carina in Urana); 3) mesonotum without gibbosities (mesonotum with four gibbosities in Urana); 4) dorsal part of periandrium unilobate (dorsal part of periandrium bilobate in Lembakaria and trilobate in Kelyflata Świerczewski & Stroiński, 2019); 5) lateral split of periandrium exceeding ⅓ of its length (lateral split of periandrium reaching ⅓ of its length in Peyrierasus Stroiński & Świerczewski, 2013). Etymology The generic name is an arbitrary combination of the name of the tribe ‘Selizini’, which the new genus belongs to, and the name of the forest formation – ‘tapia’, which the insect is associated with. Gender feminine. Description HEAD. Head with compound eyes, in dorsal view, slightly narrower than thorax. Vertex transverse, constricted in middle, medially slightly overlapped by pronotum: posterior margin carinate and strongly elevated, anterior margin carinate medially, covered by posterior margin, lateral parts obsolete; lateral margins carinate and subparallel (Figs 1A, 2A, C–E). Frons convex, widest at its lower third in frontal view; lateral margins carinate, arcuate and elevated, without incisions; upper margin almost straight; disc of frons with single, well-visible, median carina, laterally with obsolete ridges; frontoclypeal suture arcuate (Figs 1B, 2A–B). Clypeus smooth, weakly convex, without carinae (Figs 1B, 2B). Rostrum with apical segment shorter than subapical one, apex reaching hind coxae level. Compound eyes oval, with narrow callus at posterior margin. Lateral ocelli present. Antennae placed very close to medioventral margin of eyes; scapus small, ring-like, with single setae; pedicel shorter than diameter of eye but distinctly longer than scapus, bulbous, functional area at the top and on dorsal surface with trichoid sensilla type 1, antennal plate organs present on apical concavity and basally delimiting lateral margins of dorsal functional surface (Figs 2F, 3A). THORAX. Pronotum, in dorsal view, shorter than mesonotum at midline: anterior margin arcuate with median portion almost straight, reaching anterior margin of compound eyes, posterior margin concave; disc of pronotum wrinkled, without carinae, with lateral impressions and central groove; postocular eminences conical (Figs 1A, 2A, C–E). Mesonotum with scutellum widely deltoid, wider than long at midline, scutellum with elevated apex; disc of mesonotum medially depressed with shallow groove; lateral carinae as ridges, only visible in posterior part and connected with posterior margin (Figs 1A, 2E). Tegmina longer than wide, subrectangular, with distinct venation and numerous transverse veinlets in apical part, without nodal line and with single apical line; costal margin sinuate, costal and sutural angle rounded, apical margin slightly rounded, postclaval sutural margin straight. Costal area short, with dense transverse veinlets, ending at the level of fusion of claval veins (Figs 1C, 3E–F). Costal cell about the same width as costal area, tapering apicad. Basal cell longer than wide. Tegmen with longitudinal veins ScP+RA and RP arising as short common stem from basal cell before bulla. Vein ScP+RA with fork distinctly after RP fork, ending on costal margin with 4 terminals; vein RP with fork before MP fork, ending on costal margin with 8–9 terminals; vein MP with fork distinctly apicad to CuA fork, ending on apical and postclaval margins; CuA with the first fork distinctly before RP fork. Apical cells subrectangular. Veins of apical half of tegmen wrinkled. Sensory and wax gland-plates concentrated on bulla and costal area, with a few scattered on the whole tegmen (Figs 1C, 3E–F). Clavus ending a bit before the end of costal area; Pcu and A1 joined slightly anterior to clavus apex; A1 slightly elevated; sensory and wax gland-plates concentrated on the area between Pcu and A 1 and basal part of the area after A 1 vein; single transverse veinlet after Pcu-A 1 connection (Figs 1C, 3E). LEGS. Pro- and mesofemora slightly shorter than tibiae, subrectangular in cross section. Pro- and mesotibiae with shallow groove on external side; apical tarsomere of anterior and median legs longer than cumulative length of second and basal tarsomeres. Metatibiae longer than metafemora, triangular in cross section with two lateral spines and apical row of spines – first lateral spine placed subapically, second lateral spine placed a bit after midlength, apical spines in formula 2 longer (external) + 5 shorter (internal); basitarsomere of metatarsus a bit longer than cumulative length of second and apical tarsomeres, with apical spines lined as semicircle – 2 external spines a bit longer than 7 shorter internal spines; each internal spine bearing single, distinct seta; second segment of tarsomere with two lateral spines and median pad with setae. Metatibiotarsal formula: 2-(2+5)/(2+7)/2 (Figs 3B–D). MALE TERMINALIA. Anal tube, in lateral view, elongate, with breaking point before anal opening, tapering apicad; anal opening placed a bit after midlength; basal part wider than apical part (Figs 4A–B, 5A); in dorsal view, rhomboid, with rounded apex (Figs 4C–D, 5B). Pygofer, in lateral view, with dorsal and ventral margin almost the same length, subparallel; anterior margin weakly concave, posterior margin convex. Genital style triangular, bearing short, hook-like capitulum with apex oriented anteriad (Figs 4A–B, E, 5A). PHALLIC COMPLEX. Periandrium without any additional processes; in lateral view, about as long as aedeagus; lateral split reaching ⅔ of periandrium (Fig. 5C). Dorsal part of periandrium, in dorsal view, a bit shorter than ventral part, unilobate, smooth, spearhead-shape. Ventral part of periandrium elliptic, tapering apicad, basally with lateral lobes (Fig. 5D). Aedeagus, in lateral view, long and narrow, oriented ventrad, medially with acute process oriented apicad (Fig. 5E); in dorsal view bipartite, symmetrical, with deep median split, reaching ¾ of its length (Fig. 5F). FEMALE TERMINALIA. Pregenital sternite with lateral lobes distinctly separated (Figs 6A–B, 7A). Anal tube, in lateral view, covering gonoplac and reaching its posterior margin (Figs 6B, 7C); in dorsal view, elliptic (Figs 6C, 7B). Gonoplac unilobate, rounded posteriorly, oriented ventrad, covering gonapophysis VIII (Figs 6B, 7D); posterior margin with one row of stout teeth, positioned at some distance one from another; teeth of both gonoplacs fitting together in a zip-like manner (Figs 6E–F, 7D). Gonapophysis VIII widely triangular, flattened, slightly oblique in respect to longitudinal body axis (Fig. 7E); endogonocoxal process as long as gonapophysis, wide, tapering apicad, with spiniferous microsculpture. Gonospiculum as in Fig. 7H–I. Bursa copulatrix with single pouch, rounded, cells with weakly sclerotized central areas (Fig. 7F). Spermatheca well developed; ductus receptaculi longer than diverticulum ductus, both parts smooth (Fig. 7G). Tergites of abdomen membranous in median portion (Fig. 6C–D). Diversity and distribution The genus is monotypic and contains a single species from Madagascar., Published as part of Świerczewski, Dariusz & Stroiński, Adam, 2021, Selizitapia gen. nov. (Hemiptera: Fulgoromorpha: Flatidae) from tapia woodlands of Madagascar, pp. 124-139 in European Journal of Taxonomy 750 (750) on pages 126-128, DOI: 10.5852/ejt.2021.750.1367, http://zenodo.org/record/5451685, {"references":["Swierczewski D. & Stroinski A. 2019 a. Lembakaria gen. nov. - a new genus of Selizini from Madagascar spiny forest ecoregion (Hemiptera: Fulgoromoprha: Flatidae). Annales Zoologici 69 (3): 575 - 588. https: // doi. org / 10.3161 / 00034541 ANZ 2019.69.3.007"]}

A new monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae) from New Georgia Island (Solomon Islands), Kazukurugen. nov., is described for K. zingiberissp. nov. (type species). Habitus, female, external and internal genital structures of the new species are described and illustrated.

A new monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae) from Namibia, Sensoricagen. nov., is described for S. namibiensissp. nov. (type species). Habitus, female, external and internal genital structures of the new species are described and illustrated.

A new genus of ricaniid planthopper (Hemiptera: Fulgoromorpha: Ricaniidae), Jeromicanusgen. nov., is described to comprise three species – two species from Vietnam Jeromicanus orientalissp. nov. (type species) and J. vietnamensissp. nov. and one species from Laos – L. laosensissp. nov. Habitus, male and female external and internal genital structures of the new species are illustrated.

The monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae), LambertoniaLallemand, 1950 (type species – L. insignisLallemand, 1950) is redescribed and illustrated for the first time. Habitus, female external and internal genital structures are described and illustrated.

Hagneia gen. nov. (Figs 1–68) Type species. Hagneia kallea sp. nov., here designated. Locus typicus (Figs 6–7). Vietnam, Province Cao B ằng, Nguyên Bình District, Nui Pia Oac Sud (22.594395, 105.885330). Etymology. Generic name is dedicated to my dearest best friend Agnieszka Gruszczyńska. Her given name derives from the Classic Greek ἁγνείη—(h)agnein, meaning pure, holy. Gender feminine. Diagnosis. Hagneia gen. nov. can be distinguished by the combination of following characters: frontal disc with 3 carinae separated basally; clypeus with median carina; mesonotum with 5 carinae; costal area wider than postcostal cell; longitudinal veins ScP+RA, MP and CuA leaving basal cell separated; posterior part of tegmen with 2 lines of transverse veinlets; basitarsomere of metatarsus shorter than cumulative length of second and apical tarsomeres, with not fully developed row of apical teeth: ventral apical part of basitarsomere with pad of strong setae; aedeagus with 3 symmetrical pairs of well sclerotized apical spinose processes; female posterior margin of the gonoplac smooth (without any teeth). Description. HEAD. Head with compound eyes (in dorsal view) narrower than mesonotum. Vertex (Figs1–5, 11–12, 14–15) transverse, with or without median carina, distinctly wider than long at midline, all margins well carinated. Frons (Figs 4, 15–16) with all margins well carinated; at upper margin longer than high at midline, widest at the level mid of compound eyes; lateral margins covering base of pedicel, incised near the level of antenna. Frontal disc with 3 carinae, distinctly separated basally; median carina distinctly surpassing half of disc, lateral carinae distinctly shorter, ending about the lower part of compound eyes (in some specimens weakly visible). Compound eyes with very small callus at postero-ventral margin and posterior margin. Ocelli present. Antenna (Figs 17–20): pedicel cylindrical, with functional area (trichoid sensilla type 1 and antennal plate organs) at the top and on the tip of frontal side surface. Clypeus (Figs15–16) distinctly narrower than frons, with median carina. Rostrum with apical segment a little shorter than subapical one. THORAX. Pronotum (Figs 1–5, 11–14) distinctly longer than vertex at midline; disc of pronotum with median carina and two lateral impressions. Mesonotum (Figs 1–5, 11–14) elongated, diamond shape, distinctly longer at midline than combined length of vertex and pronotum; median carina, lateral and antero-lateral carinae present; median carina and lateral carinae connected basally; median carina reaching scutellum, lateral carinae not reaching posterior margin; anterolateral carinae connected with lateral at the level of lateral angles, lateral angles placed before midlength. Tegmina (Figs 1–5, 21–26) membranous, elongately-triangular, flattened, with distinct venation and transverse veinlets. Costal margin weakly arcuate, apical angle broadly rounded, placed distad to claval angle, posterior margin arcuate; tornus absent. Costal area with dense transverse veinlets ending before tip of clavus; wider than postcostal cell, wider in basal half, tapering apicad and with wave shape apex. Costal cell narrower than costal area, with transverse veinlets, in some specimens weakly incomplete and weakly visible. Basal cell elongately oval, distinctly longer than wide (about 2 times). Longitudinal veins ScP+RA, MP and CuA leaving basal cell separated; all first forks of longitudinal veins placed distinctly before half of tegmen; veins ScRA and RP arising as short common stem from basal cell (vein forked just after leaving basal cell); first fork RP before first fork ScRA and after first fork of MP 1+2; common stem of MP 1+2 and MP 3+4 longer than ScRA but shorter than CuA; first fork of MP 3+4 closer than MP 1+2, in few specimens forks at about the same levels. CuA with protruded model of forking. Tegmen with 2 lines of transverse veinlets, apical and subapical cells distinctly longer than wide, apical one shorter than subapical; nodal line absent; median portion of tegmen with numerous irregular transverse veinlets. Cubital cell with transverse veinlets at basal part. Clavus closed; CuP ending at margin, in some specimens fused with the last branch of CuA, which is very variable (single or bifurcate, straight or curved); claval veins fused after midlength of CuP vein; posterocubital cell (basal and posterior part) and postcubital cell with well visible transverse veinlets. Hind wing with precostal cell present; ScRA and MP forking distinctly after midlength of wing, CuA forking distinctly before half of wing; rp-m, m-cua and icu transverse veinlets present in distal part of wing. Hing legs (Figs 27–32): metatibia distinctly longer than metafemur, partly flattened and widened at distal part; metatibia with 2 lateral spines placed distally to each other in distal part; apical row of teeth of metatibia with 7 well developed spines different in size; external lateral spines bigger than internal lateral spines; 5 internal spines different in size with asymmetrical diastema in formula 1+4 (with external spines formula 2 +5); basitarsomere of metatarsus shorter than cumulative length of second and apical tarsomeres, with not fully developed row of apical teeth: 2 lateral teeth equal in size and big and 3 internal small placed externally/asymmetry 1(2)1; ventral apical part of basitarsomere with pad of strong setae; mesotarsomere with pad of strong setae on ventral side. MALE TERMINALIA (Figs 33–48). Anal tube (in lateral view, Figs 33–34, 42) elongate, massive, distinctly surpassing posterior margin of pygofer; posterior part narrower than basal one; anus placed before midlength, ventral margin straight. Anal tube (in dorsal view, Figs 35–36, 39) elongate, subrectangular, widest about midlength; anus place before midlength, postero-ventral angle widely rounded; posterior margin in dorsal view with median concavity, lateral margins slightly arcuate. Pygofer (in lateral view, Figs 35–36, 39) higher than wide; dorsal part narrower than ventral one, dorso-posterior angle with well-developed process. Genital styles (Figs 35–38, 40–41), in lateral view, longer than wide with sharp spine-like process at the end of dorsal margin; apical part wider than basal; ventral margin almost straight (in lateral view), weakly sinuate in ventro-lateral view; dorsal margin weakly convex, with small concavity before spine-like process; caudo-dorsal angle widely rounded and surpassing the base of process, posterior margin weakly convex. Phallix complex (Figs 43–48): periandrium (Figs 43–45) without any processes, elongate, with long lateral split surpassing the half of its length; dorsal part of periandrium a bit shorter than ventral one, median part partly membranous, apically with median split; ventral part with widened apex and additional small lateral lobes. Aedeagus (Figs 46–48) long and narrow, apically with 3 pairs of symmetrical,, well sclerotized, spinose processes and short median split. Median split asymmetrical: ventral split present only in 1/4 ending with elongate triangular lobe; dorsal split very deep, reaching almost basal part. All processes single armed and oriented basad: dorsal processes biggest than other, distinctly curved in 1/3 of its length, posterior part with membranous ventral margin; median processes shorter, massive and fully sclerotized; ventral processes with small membranous part at basal part on dorsal margin, with little curved tip. FEMALE TERMINALIA (Figs 49–68). Pregenital sternite with well-developed and distinctly separated lateral lobes; posterior margin medially with processes (Figs 49–51, 61). Anal tube (in lateral view, Figs 55–57, 63) not reaching the posterior margin of gonoplac; anal tube (in dorsal view, Figs 53–54, 62) ovoid, wider before anus; anus placed a bit after midlength; anal style (paraproct) and anal segment (epiproct) short. Gonoplac (Figs 49–50, 55–60, 64) well developed, unilobate, laterally flattened; posterior margin of the gonoplac smooth (without any teeth); membranous part of gonoplac placed basally on ventral margin. Gonapophysis VIII (Fig. 65) sabre-like, “v” shape in cross section, with teeth at dorsal margin; endogonocoxal process tapering apicad, shorter than gonapophysis VIII, with median sclerotized core surrounded by membranous part. Gonaphophyses IX and gonospiculum bridge well developed, as in Figs 66–67. Bursa copulatrix (Fig. 68) of two pouches connected by narrow part; first pouch elongate, with cells and sclerotised ornamentation (except dorsal side); second pouch smaller, more oval than the first one, without cells but with sclerotized plates. Spermatheca (Fig. 68) well developed; ductus receptaculi elongate and narrow, ribbed; diverticulum ductus about as long as ductus receptaculi, with long narrow smooth ductus, apically with ovoid and smooth bulla. Distribution. Vietnam, Province Cao Bằng, Nguyên Bình District.

A new monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae), Hagneia gen. nov., is described for Hagneia kallea sp. nov. (type species). Habitus, male and female external and internal genital structures of the new species are illustrated.

Hagneia kallea sp. nov. (Figs 1–68) Etymology. Specific epithet derives from Classical Greek κάλλος, ~εος kallos, ~eos meaning good, beauty, noble. Diagnosis. Only one species in the genus. Male: dorso-posterior angle of pygofer with process oriented dorsad; female: pregenital sternite medially with 2 separated processes at posterior margin. Description. Total length 0.8–1.3 cm. HEAD. Vertex: proportion A/B = 9.6–11.25; all margins slightly elevated; in dorsal view, lateral margins almost straight and parallel; anterior margin widely arcuate, posterior margin with major curvature than anterior one. Frons: proportion C/E = 1.25–1.32; proportion D/E = 1.43–1.5; upper margin straight, lateral margins weakly arcuate, distinctly curved to frontoclypeal suture in lower part. Frontal disc delicately rugose vertically, near the frontoclypeal suture concave; lateral carinae of disc parallel to lateral margins (in some specimens weakly visible). Frontoclypeal suture widely arcuate. Clypeus with short median carina present in median part, surface of clypeus in median part weakly convex. Rostrum reaching mid coxae. THORAX. Pronotum: proportion F/B = 2.3–2.75; disc of pronotum delicately rugose; anterior margin widely arcuate; posterior margin weakly concave. Mesonotum: proportion G/F+B = 6.5–6.85, proportion G/F = 9.04–9.5, proportion G/H = 1.12.–1.19. Tegmina: proportion I/J = 1.41–1.47. Hind wing ScRA with 3 terminals, MP with 2–3 terminals, CuA wit 7–11 terminals. MALE TERMINALIA. Dorso-posterior angle of pygofer with process oriented dorsad; posterior margin in 3/4 straight, in 1/4 convex (lower part). FEMALE TERMINALIA. Pregenital sternite with posterior margin medially with 2 separated processes. COLORATION (Figs 1–5). Frons black with dark yellow lateral margins and narrow band alongside lateral margins. Clypeus and postclypeus medially dirty brown, lateral areas black; lateral part of head black with yellow narrow band alongside anterior margin and dark yellow patch around base of antenna. Compound eyes with irregular black and brown patches. Vertex black with lateral margins and narrow band alongside, lateral margins brown, pronotum (lateral lobes black with brown margin), mesonotum and tegulae black. Tegmina black with white tearshaped patch in middle and white-yellowish triangular patch near of end of costal area and huge transverse band alongside posterior margin. Hind wing with brown transverse band subapically posterior margin. Pro- and mesofemur on external and internal lateral side black, dorsally brown, pro-, mesotibia and tarsomeres brown. Metafemur, metatibia and metatarsus brown. Abdomen (male and females) black with brown narrow (dorsally) anterior margin (in females band bigger) of the tergites, ventral side of pygofer brown; terminalia black. Type material. Holotype, male: [Vietnam, Cao B ằng Province, / Nguyên Bình District, / Phia Oac National Parc, / 22°35’39.8”N 105°53’07.2”E], [1282–1290 m a.s.l., 8– 12.07.2019, / day and light trapping / leg. T. Bourgoin, J. Gunczy, G. / Kunz, A. Soulier and A. Stroiński]—deposited in MIZ. Praratypes, 6 ♂♂, 7 ♀♀: [Vietnam, Cao B ằng Province, / Nguyên Bình District, / Phia Oac National Parc, / 22°35’39.8”N 105°53’07.2”E], [1282–1290 m a.s.l., 8– 12.07.2019, / day and light trapping / leg. T. Bourgoin, J. Gunczy, G. / Kunz, A. Soulier and A. Stroiński]— 5 ♂♂, 6 ♀♀ deposited in MIZ, 1 ♂, 1♀ deposited in MNHN. 4 ♀♀ preserved in MIZ were used for endosymbiotic studies and they lack abdomens. Distribution. Vietnam: Cao Bằng Province, Nguyên Bình District, Phia Oac National Parc— 22°35’39.8”N 105°53’07.2”E (22.594395, 105.885330).

Given a finite dimensional algebra $A$, we consider certain sets of idempotents of $A$, called self-injective cores, to which we associate 2-subcategories of the 2-category of projective bimodules over $A$. We classify the simple transitive 2-representations of such 2-subcategories, vastly generalizing the main result of arXiv:1805.05724., 21 pages