Search

Siobla sturmii (Klug, 1817) (Figs 4–6) Tenthredo (Allantus) sturmii Klug, 1817: 116. Macrophya pacifica F. Smith, 1874: 378. Macrophya castanea Jakowlew, 1888: 373. Encarsioneura frigida Mocsáry, 1909: 15–16. Diagnosis. This species is similar to S. jucunda Mocsáry, but differs from the latter by the following: the microsculptures on the 2nd and 3rd abdominal tergites fine, matt; hairs on dorsum of head and of thorax black; malar space of male as long as the radius of a lateral ocellus, the 4th and 5th abdominal tergites blackish brown, the lobe of pennis narrow; malar space of female as long as the diameter of the middle ocellus, the 9th abdominal tergite yellowish brown, vein C almost entirely blackish brown. Distribution. China (Heilongjiang (Wuying), Jilin (Mt. Changbai), Liaoning (Xinbin), Shanxi (Mt. Wutai), Hebei (Mt. Xiaowutai)), Japan (Hokkaido, southern Kuriles, Honshu, Kyushu), South Korea, Russian Far East (Primorskiy Kray; Sakhalin), Siberia, Europe. Primary type examined. 1♀, lectotype (HNHM, here designated): “Sibirica or., Raddefka”; “ Encarsioneura frigida, det. Mocsáry” [red label]; “ Lectotype, Encarsioneura frigida Mocsáry, 1909, Des. Niu G, 2010”; “DEI-GISHym 1005”; “ Siobla sturmii (Klug, 1817), Det. Niu G, 2010”. 1♀, paralectotype (HNHM, here designated): “Sibirica or., Raddefka”; “ Encarsioneura frigida, det. Mocsáry” [red label]; “ Paralectotype, Encarsioneura frigida Mocsáry, 1909, Des. Niu G, 2010”; “DEI-GISHym 1000”; “ Siobla sturmii (Klug, 1817), Det. Niu G, 2010”. 1♂, paralectotype (HNHM, here designated): “Sibirica or., Raddefka”; “ Encarsioneura frigida, det. Mocsáry” [red label]; “ Paralectotype, Encarsioneura frigida Mocsáry, 1909, Des. Niu G, 2010”; “DEI-GISHym 1004”; “ Siobla sturmii (Klug, 1817), Det. Niu G, 2010”. Remarks. No distinct difference was found between the types of Siobla frigida and S. sturmii. The former is treated as a junior synonym of the latter following Malaise (1931)., Published as part of Niu, Gengyun & Wei, Meicai, 2020, Review of some Siobla (Hymenoptera: Tenthredinidae) species, pp. 208-224 in Zoological Systematics 46 (3) on pages 210-212, DOI: 10.11865/zs.2020303, http://zenodo.org/record/7175799, {"references":["Smith, F. 1874. Descriptions of new species of Tenthredinidae, Ichneumonidae, Chrysididae, Formicidae, etc. of Japan. Transactions of the Entomological Society of London, 1874: 373 - 409.","Jakowlew, A. 1888. Quelques nouvelles especes des mouches a scie de l'Empire Russe. Trudy Russkogo Entomologiceskogo Obscestva v S. Peterburge, 22: 368 - 375.","Mocsary, A. 1909. Chalastogastra nova in collectione Musei nationalis Hungarici. Annales Historico-Naturales Musei Nationalis Hungarici, 7: 1 - 39.","Gimmerthal, B. A. 1834. Einige in Livland aufgefundene und benannte Sagewespen (Tenthredinae). Bulletin de la Societe Imperiale des Naturalistes de Moscou, 7: 122 - 128.","Malaise, R. 1931. Blattwespen aus Wladiwostock und anderen Teilen Ostasiens. Entomologisk Tidskrift, 52: 97 - 159."]}

Siobla plesia Malaise, 1945 stat. nov. (Figs 11–13) Siobla sturmii plesia Malaise, 1945: 127. Type locality: Sichuan, China. Diagnosis. This species is similar to S. yunanensis, but differs from the latter by the following: mesoscutellum flat. Female: basal three antennomeres with black spots, abdominal sternites yellowish brown; Male: dorsum of flagellum without black stripe, posterior margin of pronotum reddish brown, outer of basal fumer of the hind leg with reddish spot, abdominal sternites reddish brown. Distribution. China (Sichuan (Ganzi)). Primary type examined. 1♀, Lectotype (USNM, here designated): “ 9 mi. SW of Tatsienlu, Jun 25–7.23, 8500–13000ft ”; “Szechuen CHINA DCGraham”; “Type No. 27440, USNM ”; “ S. sturmii (Kl), var. plesia n.var. Malaise det. 1936”; “ Siobla plesia, TYPE. Roh.”. 1♂, paralectotype (USNM, here designated): “ 9 mi. SW of Tatsienlu, Jun 25–7.23, 8500–13000ft ”; “Szechuan CHINA DC Graham”; “Type No. 27440, USNM ”; “ S. sturmii (Kl), var. plesia n. var. Malaise det. 1936”; “ Siobla plesia, allotype. Roh.”. Other material examined. 1♀ 1♂, Maiba, Kangding, Sichuan, E101º34.856′, N30º03.631′, 3525 m, 2009-VII-01, Wei Meicai; 1♀ 1♂, China, Kangding, 2800 m, Sichuan Prov. 18.VI.1990, F. S. Naito leg., F.S.Huang leg.; Exchange KUK; 1♀ 1♂, China, Mt. Zheduoshan 3100 m, Sichuan Prov. 21.VI.1990. M. T. Naito leg. M. Takeda leg.; Exchange KUK. Remarks. Malaise (1945) reported plesia as a subspecies of S. sturmii (Klug, 1817). Here we treat plesia as an independent species as it is distinct from S. sturmii. In S. sturmii, the female has following characters: abdominal tergites 2–4 and tergite 9 yellowish brown; pale margin of pronutum narrow; hairs on head and thorax brown; hind tarsi somewhat dark; the male abdominal tergites 2–3 yellowish brown; sternites 5–7 and basal plate black; the female lancet more slender with sharper serrulae, the area below pore line (aso of Ross, 1945) quite narrow and bent, dorsal margin round (Figs 13f, h); the inner margin of male harpe weakly incised (Fig. 13j); the dorsal margin of the valviceps convex above middle (Fig. 13l). In S. plesia, the female has follwing characters different from S. Sturmii: abdominal tergites 2–5, and tergites 7–9 yellow (Figs 11a–c); pale margin of pronutum broad (Fig. 13c); hairs on head (Figs 12a, c, e) and thorax (Figs 13a, c) yellowish brown; hind tarsi yellow (Figs 11a–c); the male abdominal tergites 2–4 yellowish brown; sternites 5–7 and basal plate yellow with dark macula (Figs 11e–g); the female lancet less slender with obtuse serrulae, the area below pore line broad and not bent, dorsal margin almost truncate (Figs 13e, g); the inner margin of male harpe weakly convex (Fig. 13i); the dorsal margin of the valviceps convex below middle (Fig. 13k)., Published as part of Niu, Gengyun & Wei, Meicai, 2020, Review of some Siobla (Hymenoptera: Tenthredinidae) species, pp. 208-224 in Zoological Systematics 46 (3) on page 218, DOI: 10.11865/zs.2020303, http://zenodo.org/record/7175799, {"references":["Malaise, R. 1945. Tenthredinoidea of South-Eastern Asia with a general zoogeographical review. Opuscula Entomologica, Suppl. 4: 1 - 288.","Ross, H. H. 1945. Sawfly genitalia: terminology and study techniques. Entomological News, 56: 261 - 268."]}

Siobla iridipennis Malaise, 1934 (Fig. 14) Siobla iridipennis Malaise, 1934: 23; Malaise, 1945: 117–118. Description. Holotype ♀. Body length 9.0 mm (Figs 14a–c). Yellow brown, following parts black: head and antenna entirely, narrow lateral margin of pronotum, bottom of parapsis, metapostnotum, propleuron, ventral half of mesepisternum, middle of mesepimeron, metapleuron except for dorsal margin and small posterior corner, narrow anterior margin of tergite 2, narrow base of basal ovipositor sheath and most of apical ovipositor sheath; legs black, apical half of fore femur on anterior side, ventral side of fore tibia and of tarsus, ventral side of middle tarsus pale brown; most of middle femur and hind femur except for extreme apex yellow brown. Wing evenly infuscate, with feeble purple reflection, paler toward apex, pterostigma and veins black brown. Hairs on head black brown, hairs on dorsal of thorax pale brown and on mesopleuron brown, setae on sheath pale brown. Punctures on clypeus large and sparse, interspaces broad and smooth (Fig. 14f); punctures on dorsum and lateral of head denser, with narrow but distinct smooth interspaces (Figs 14e, g); frons, lateral fovea and middle fovea densely punctured, with very narrow interspaces; interspaces between punctures on middle of inner orbit about as broad as diameter of a puncture, smooth and shiny; punctures dense on anterior of mesoscutal middle lobe, large and sparse on middle and posterior area, interspaces smooth and broader than diameter of a puncture; punctures on mesoscutal lateral lobe smaller and denser (Fig. 14h), interspaces smaller than diameter of a puncture, top area with feeble microsculptures; punctures on anterior slope of mesoscutellum shallow and sparse, interspaces smooth and as broad as a puncture, strongly shiny; posterior third of mesoscutellum coarsely punctured, mesoscutellar appendage smooth on middle and microsculptured laterally; punctures on metascutellum large and sparse, shiny; metapostnotum densely microsculptured, feebly shiny; punctures on propleuron minute and sparse (Fig. 14i); upper half of mesepisternum coarsely and densely punctured, anterior margin sparsely punctured, punctures on ventral half of mesepisternum minute and very sparse, interspaces broad and shiny, a large ventral anterior corner of mesepisternum smooth and shiny; middle of mesepimeron with large and dense punctures, bottom of anepimeron and most of katepimeron densely microsculptured, posterior margin of katepimeron smooth; dorsum of metepisternum evenly punctured, without microsculpture, ventral of metepisternum smooth and shiny; depressed part of metepimeron with shallow punctures and fine microsculptures, dorsal margin with dense punctures, posterior corner smooth; abdominal tergites 1 and 2 quite smooth (Fig. 14j), small middle area with several punctures; other tergites with dense and large punctures, interspaces smaller than punctures, surface smooth, strongly shiny. Hairs on dorsum of head 2 times as long as diameter of lateral ocellus, hairs on mesopleuron about 2.2 times diameter of lateral ocellus. Clypeus about 2 times as broad as long, middle length distinctly longer than lateral length, apical margin roundly protruding (Fig. 14f); distance between eyes at level of toruli 1.2 times longest axis of eye; malar space 0.6 times diameter of lateral ocellus; middle fovea broad and deep, lateral fovea small and narrow, much deeper than middle fovea; supraantennal tubercle distinctly elevated, broader than high and about as high as frontal wall; frontal wall low and obtuse; interocellar furrow deep and narrow, postocellar furrow shallow, lateral furrows fine, weakly curved and slightly divergent backwards; postocellar area flat, clearly lower than top of ocelli, without middle ridge, broader than long (1.5: 1); in dorsal view temple almost as long as eye (0.95: 1), lateral sides weakly bent and slightly convergent backwards (Fig. 14e); occipital carina low and weak, complete. Antenna 0.9 times as long as head and thorax together, clearly shorter than abdomen or vein C, pedicellum 1.2 times as long as broad, antennomere 3 about 1.9 times as long as antennomere 4, outer side of antennomeres 5–8 with distinct longitudinal furrow, subapical antennomeres clearly enlarged, antennomere 7 about 1.3 times as long as broad. Middle furrow of mesoscutal middle lobe deep; mesoscutellum weakly elevated, much broader than long (1.8: 1), without peak and carina, posterior slope almost flat; mesoscutellar appendage flat, without middle carina. Metapostnotum with a sharp middle ridge and merged with middle process of first tergite, about as high as metascutellum; middle of mesepisternum distinctly elevated, ventral spur absent. Legs slender, hind tibia 1.2 times longer than tarsus, apex as broad as breadth of apical sheath in lateral view, inner apical spur 0.5 times as long as metabasitarsus; metabasitarsus slender, 5 times as long as broad, as long as following 3 tarsomeres together; tarsal pulvilli developed, about half as long as apical breadth of each tarsomeres, distance between basal 2 pulvilli 1.8 times as long as second pulvillus (Fig. 14l). Forewing with vein R clearly shorter than free part of Sc and about 0.33 times length of vein R+M, vein R+M 2 times as long as first absccisa of vein Rs, vein 2r strongly curved, vein cu-a meeting cell 1M at basal 0.2, pterostigma 1.2 times as long as cell 1Rs; stalk of hind anal cell about 0.33 times length of vein cu-a. Sheath 1.25 times as long as middle tibia, apical sheath 1.5 times as long as basal sheath (Fig. 14k), setae on sheath long and dense, distinctly curved in dorsal view. Lancet (Figs 14m – n) with 14 serrulae, spiculella distinct, middle serrulae distinctly convex and with 18–22 small distal teeth, proximal basal tooth absent, cypsela clearly shorter than serrula. Male. Unknown. Primary type examined. 1♀, holotype (BMNH): “ Tibet, Tropde, 11000ft., 23.VI.1924, Maj. R. W. G. Hingston ”; “Everest Exp. Brit. Mus., 1924. 386”; “ Siobla iridipennis n. sp. (Type), Malaise det., 1933”; “Typus (coll. BMNH)”. Distribution. China (Tibet (Tropde)). Remarks. The species is rare found. Malaise (1934) simply described the species based on a female specimen in a key to Siobla species. After then there is no additional collecting record about the species. Malaise (1945) keyed the species again but the description was still simple and most characters were not mentioned. Siobla iridipennis is undoubtedly a species of Siobla as shown by the metabasitarsus distinctly compressed, the anterior margin of clypeus roundly protruding, anal cross vein of forewing oblique and at middle of the broad anal cell, and female lancet quite long and slender. However, Siobla iridipennis is remote from its congeners by the following characters: head subquadrate, temple distinctly elongated; sheath with dense setae, lancet with distinct spiculella; mesoscutellum quite flat and sparsely punctured; antenna stout and short; body yellow brown including head and antenna entirely, the ventrum of thorax black. Siobla iridipennis was placed in Siobla ferox group in the paper of Niu & Wei (2010a), based on the simple original description of Malaise (1934, 1945). However, it is unique in body color and structure as discussed above. It seems that S. iridipennis is possibly a basal lineage of the genus. So it is necessary to propose a new species group, Siobla iridipennis group, for the species. Funding The research was partly supported by the National Natural Science Foundation of China (31501885, 31672344, 31970447). Acknowledgements Our cordial thanks are due to Dr. David R. Smith of USNM, Dr. Natalie Dale-Skey of BMNH, Dr. Zombori of HNHM, Dr. Hege Vårdal, the curator of Hymenoptera Collection, Naturhistoriska riksmuseet, Sweden in lending us type material of Siobla species and their help when we visited the museums. Our thanks are also due to anonymous reviewers., Published as part of Niu, Gengyun & Wei, Meicai, 2020, Review of some Siobla (Hymenoptera: Tenthredinidae) species, pp. 208-224 in Zoological Systematics 46 (3) on pages 218-223, DOI: 10.11865/zs.2020303, http://zenodo.org/record/7175799, {"references":["Malaise, R. 1934. Schwedisch-chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas unter Leitung von Dr. Sven Hedin und Prof. Su Ping-Chang. Insekten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 23. Hymenoptera. 1. Arkiv for Zoologi, 27 [1934 - 1935] (2 [nr A 9]): 1 - 40.","Malaise, R. 1945. Tenthredinoidea of South-Eastern Asia with a general zoogeographical review. Opuscula Entomologica, Suppl. 4: 1 - 288."]}

Siobla ruficornis (Gimmerthal, 1834) (Figs1–3) Allantus ruficornis Gimmerthal, 1834: 125. Eriocampa ruficornis Cameron, 1876: 462. Siobla ruficornis var. albicornis Malaise, 1931: 121. Siobla sibirica var. bergmani Malaise, 1931: 122. Encarsioneura similis Mocsáry, 1909: 14–15. Type locality: Sibiria orientalis: Ussuri. syn. nov. Diagnosis. Only two species of Siobla are distributed in Europe, S. ruficornis and S. sturmii. The current species differs from the latter by the following: broad margin of clypeus and of pronotum reddish brown, microsculpures on the 2nd and 3rd abdominal tergites clear, hairs on dorsum of head pale brown, and slightly longer than the diameter of lateral ocellus. Within the genus, this species is similar to S. centralia, but differs from the latter by the following: the 2nd hind trochanter of both sexes black, the 7th to 9th abdominal tergites almost entirely black in female, at least the apical two tergites black in male; microsculptures clear in both sexes; sheath drastically narrow towards the apex. Distribution. China (Heilongjiang (Yichun), Jilin (Mt. Changbai), Qinghai (Huzhu), Hebei), Russia, Finland, Latvia. Primary type examined. 1♀, lectotype (HNHM, here designated): “Ussuri, Kasakewiisch, 1907, Koro”; “ Encarsioneura similis, det. Mocsáry” [red label]; “ Lectotype, Encarsioneura similis Mocsáry, 1909, Des. Niu G, 2010”; “DEI-GISHym 1001”; “ Siobla ruficornis (Gimmerthal, 1834), Det. Niu G, 2010”. 1♀, paralectotype (HNHM, here designated): “Ussuri, Kasakewiisch, 1907, Koro”; “ Encarsioneura similis, det. Mocsáry” [red label]; “ Paralectotype, Encarsioneura similis Mocsáry, 1909, Des. Niu G, 2010”; “DEI-GISHym 1002”; “ Siobla ruficornis (Gimmerthal, 1834), Det. Niu G, 2010”. Remarks. The types of Encarsioneura similis Mocsáry were carefully compared with specimens of Siobla ruficornis Gimmerthal (type missing). No distinct difference was found between Encarsioneura similis and S. ruficornis. The former is treated as a junior synonym of the latter here. The record of S. ruficornis from South Korea (Lee et al., 2000) is a misidentification of S. jucunda (Mocsáry, 1909) and Siobla sturmii (Klug, 1817)., Published as part of Niu, Gengyun & Wei, Meicai, 2020, Review of some Siobla (Hymenoptera: Tenthredinidae) species, pp. 208-224 in Zoological Systematics 46 (3) on pages 209-210, DOI: 10.11865/zs.2020303, http://zenodo.org/record/7175799, {"references":["Gimmerthal, B. A. 1834. Einige in Livland aufgefundene und benannte Sagewespen (Tenthredinae). Bulletin de la Societe Imperiale des Naturalistes de Moscou, 7: 122 - 128.","Cameron, P. 1876. VIII. Descriptions of new genera and species of Tenthredinidae and Siricidae, chiefly from the East Indies, in the Collection of the British Museum. Transactions of the Royal Entomological Society of London, 24 (3): 459 - 471.","Malaise, R. 1931. Blattwespen aus Wladiwostock und anderen Teilen Ostasiens. Entomologisk Tidskrift, 52: 97 - 159.","Mocsary, A. 1909. Chalastogastra nova in collectione Musei nationalis Hungarici. Annales Historico-Naturales Musei Nationalis Hungarici, 7: 1 - 39.","Lee, J. W., Ryu, S. M., Quan, Y. T., Jung, J. C. 2000. Economic Insects of Korea 2. Hymenoptera (Symphyta: Tenthredinidae). National Institute of Agricultural Science and Technology, Suwon. pp. 1 - 222."]}

Niu, Gengyun, Wei, Meicai (2020): Review of some Siobla (Hymenoptera: Tenthredinidae) species. Zoological Systematics 46 (3): 208-224, DOI: 10.11865/zs.2020303

Xiao, Wei, Niu, Gengyun, Wei, Meicai (2020): Two new species of the Tenthredo grahami group from China (Hymenoptera: Tenthredinidae), with a revised key to species. Zoological Systematics 45 (2): 118-125, DOI: 10.11865/zs.202016, URL: http://zoobank.org/30ef491f-21f8-4c42-b270-b7b3360c0a35

Tenthredo megamaculata sp. nov. (Figs 1 C–D, 3) Diagnosis. This new species is most similar to T. gulmargi Singh & Saini, 1987 from North India, but differs from the latter by the black spots on the 2nd to 8th abdominal tergites distinctly longer than half length of abdominal tergites, the 5th cypsella of lancet much shorter than 5th serrula, the male penis valve with narrow and long apical process, and the apex of parapenis almost truncate without inner process, while in T. gulmargi, the black spots on abdominal tergites about half length of the abdominal tergite, the 5th cypsella of lancet as long as 5th serrula, the male penis valve simple without apical process, and the apex of parapenis with a distinct inner process). Description. Female. Body length 10.5–11.5mm (Fig. 1C). Body and legs light green in alive by yellow brown in dried; following parts black: median fovea and connected broad lateral stripes of frons and postocellar area, ocellar basin, a small middle spot on postocellar area (Fig. 3B), antenna except apex of first antennomere, a narrow transverse stripe on bottom of pronotum, a wide middle stripe on mesoscutal middle lobe, dorsum of mesoscutal lateral lobe, bottom of parapsides, basal 1/3 of 1st abdominal tergite, middle strongly constricted basal bands on dorsum of 2nd to 8th abdominal tergites (Fig. 1C), slightly curved vertical narrow stripe on mesepisternum (Fig. 3D), a dorsal narrow stripe on all legs from trochanter to tarsi; most hairs on upper 2/5 of hind orbit and dorsum of head, thorax and abdomen black, setae on sheath brown, most hairs on lateral and ventral side of body silver. Wings hyaline, pterostigma and vein C yellow green, other venation brownish. Dorsal side of head (Fig. 3B) with fine but clear microsculpture, weakly shiny, microsculpture within black spots more distinct; mesonotum finely but densely sculptured, lateral lobe with sparse and minute punctures; anterior slope of mesoscutellum faintly microsculptured, posterior slope and lateral of appendage shallowly and sparsely punctured; upper half of mesopleuron finely microsculptured, shiny, ventral half sparsely punctured mixed with microsculpture; abdominal tergites finely sculptured, weakly shiny; sides of apical sheath with fine microsculpture, strongly shiny. Hairs on dorsum of head straight and distinctly inclined forward, 0.8 times as long as diameter of lateral ocellus; length of hairs on mesonotum, mesoscutellum and mesopleuron about 0.5, 0.75 and 1 times diameter of lateral ocellus, respectively. Apex of labrum round; clypeus large and flat, distinctly broader than distance between eyes below, anterior margin narrowly and shallowly incised to a depth about 0.25 times length of clypeus, lateral lobe truncate (Fig. 3A); malar space equal to diameter of lateral ocellus; distance between lower corners of eye 0.9 times longest axis of eye; supraantennal tubercles strongly elevated, free-standing and slightly divergent backwards, height 1.1 times its width, length about 2 times its height and as long as postocellar area, distance between posterior ends of tubercles 1.2 times its breadth, posterior end abruptly cut off from frontal ridges, bottom of median fovea without carina; interocellar furrow deep and straight, postocellar furrow deep; postocellar area hardly elevated, about 1.4 times as broad as long, without middle carina or middle furrow, lateral furrows weakly curved, distinctly divergent backward; in dorsal view length of temple 0.5 times eye length, lateral sides distinctly curved and narrowed backward (Fig. 3B); occipital carina low but complete, lower part without fold. Antenna filiform, not compressed, about as long as head and thorax together, 3rd antennomere 1.5 times as long as 4th antennomere, 8th antennomere 2.5 times as long as broad (Fig. 3I). Mesoscutellum roundly elevated, without longitudinal carina or peak but with faint transversal ridge, mesoscutellar appendage with short longitudinal carina; distance between cenchri 2.2 times longest axis of a cenchrus; mesepisternum weakly elevated at middle, ventral thorn absent (Fig. 3D). Hind tibia 0.8 times as long as tarsus, inner tibial spur 0.55 times as long as basitarsus; basitarsus slender, 0.8 times as long as following 3 tarsomeres together; claw with a short and obtuse basal lobe, inner tooth distinctly shorter than outer tooth. Vein cu-a in forewing joining cell 1M at basal 0.35, cell 2Rs distinctly longer than cell 1Rs; hind anal cell sessile (Fig. 1C). Lancet narrow and long with 17 serrulae (Fig. 3C), serrulae distinctly oblique, middle serrulae each with 1 inner subbasal tooth and 11–13 distal subbasal teeth, annular sutures strongly oblique, annular spines bands very narrow, annular spines very short and sparse; 5 th to 7 th serrulae as in Figure 3H. Male. Body length 8.5–9.5 mm (Fig. 1D). Body color and structure similar to female but hind coxa with narrow black dorsal and ventral stripes, malar space 0.5 times diameter of lateral ocellus, distance between eyes below about 0.7 times longest axis of eye, in dorsal view head shorter and distinctly narrowed, basal lobe of claw distinct; subgenital plate 1.2 times as long as broad, apex roundly and narrowly convex; harpe longer than broad, apex of parapenis almost truncate without inner process (Fig. 3F); penis valve broad with a long and narrow apical process (Fig. 3G). Material examined. Holotype. ♀, China, Sichuan Province, Mt. Emei, Leidongping (29°55′N, 103°33′E; elev. 2350 m), 7 July 2009, Meicai Wei leg. Paratypes. 1♀, Sichuan Province, Mt. Emei, Leidongping (29°55′N, 103°33′E; elev. 2350 m), 7 July 2009, Meicai Wei leg.; 2♀ 2♂, Sichuan Province, Mt. Emei, Leidongping (29°55′N, 103°33′E; elev. 2350 m), 8 July 2009, Meicai Wei leg.; 1♀ 1♂, Sichuan Province, Mt. Emei, Leidongping (29°55′N, 103°33′E; elev. 2350 m), 8 July 2009, Gengyun Niu leg.; 2♀ 3♂, Sichuan Province, Mt. Emei, Leidongping (29°55′N, 103°33′E; elev. 2350 m), 7 July 2009, Zejian Li leg.; 1♀ 2♂, Sichuan Province, Mt. Emei, Jinding (29°31.37′N, 103°21.22′E; elev. 3073 m), 20 July 2011, Junzhe Xue & Hu Ping leg.; 1♀, Tibet, Chaya County, G241 (30°42.59′N, 97°19.56′ E; elev. 3350 m), 24 June 2009, Zejian Li leg. Distribution. China (Sichuan, Tibet). Etymology: The specific epithet is composed of a Latin prefix mega- and maculata, referring to the abdomen with large black maculae.

Tenthredo megamaculata sp. nov. (Figs 1 C���D, 3) Diagnosis. This new species is most similar to T. gulmargi Singh & Saini, 1987 from North India, but differs from the latter by the black spots on the 2nd to 8th abdominal tergites distinctly longer than half length of abdominal tergites, the 5th cypsella of lancet much shorter than 5th serrula, the male penis valve with narrow and long apical process, and the apex of parapenis almost truncate without inner process, while in T. gulmargi, the black spots on abdominal tergites about half length of the abdominal tergite, the 5th cypsella of lancet as long as 5th serrula, the male penis valve simple without apical process, and the apex of parapenis with a distinct inner process). Description. Female. Body length 10.5���11.5mm (Fig. 1C). Body and legs light green in alive by yellow brown in dried; following parts black: median fovea and connected broad lateral stripes of frons and postocellar area, ocellar basin, a small middle spot on postocellar area (Fig. 3B), antenna except apex of first antennomere, a narrow transverse stripe on bottom of pronotum, a wide middle stripe on mesoscutal middle lobe, dorsum of mesoscutal lateral lobe, bottom of parapsides, basal 1/3 of 1st abdominal tergite, middle strongly constricted basal bands on dorsum of 2nd to 8th abdominal tergites (Fig. 1C), slightly curved vertical narrow stripe on mesepisternum (Fig. 3D), a dorsal narrow stripe on all legs from trochanter to tarsi; most hairs on upper 2/5 of hind orbit and dorsum of head, thorax and abdomen black, setae on sheath brown, most hairs on lateral and ventral side of body silver. Wings hyaline, pterostigma and vein C yellow green, other venation brownish. Dorsal side of head (Fig. 3B) with fine but clear microsculpture, weakly shiny, microsculpture within black spots more distinct; mesonotum finely but densely sculptured, lateral lobe with sparse and minute punctures; anterior slope of mesoscutellum faintly microsculptured, posterior slope and lateral of appendage shallowly and sparsely punctured; upper half of mesopleuron finely microsculptured, shiny, ventral half sparsely punctured mixed with microsculpture; abdominal tergites finely sculptured, weakly shiny; sides of apical sheath with fine microsculpture, strongly shiny. Hairs on dorsum of head straight and distinctly inclined forward, 0.8 times as long as diameter of lateral ocellus; length of hairs on mesonotum, mesoscutellum and mesopleuron about 0.5, 0.75 and 1 times diameter of lateral ocellus, respectively. Apex of labrum round; clypeus large and flat, distinctly broader than distance between eyes below, anterior margin narrowly and shallowly incised to a depth about 0.25 times length of clypeus, lateral lobe truncate (Fig. 3A); malar space equal to diameter of lateral ocellus; distance between lower corners of eye 0.9 times longest axis of eye; supraantennal tubercles strongly elevated, free-standing and slightly divergent backwards, height 1.1 times its width, length about 2 times its height and as long as postocellar area, distance between posterior ends of tubercles 1.2 times its breadth, posterior end abruptly cut off from frontal ridges, bottom of median fovea without carina; interocellar furrow deep and straight, postocellar furrow deep; postocellar area hardly elevated, about 1.4 times as broad as long, without middle carina or middle furrow, lateral furrows weakly curved, distinctly divergent backward; in dorsal view length of temple 0.5 times eye length, lateral sides distinctly curved and narrowed backward (Fig. 3B); occipital carina low but complete, lower part without fold. Antenna filiform, not compressed, about as long as head and thorax together, 3rd antennomere 1.5 times as long as 4th antennomere, 8th antennomere 2.5 times as long as broad (Fig. 3I). Mesoscutellum roundly elevated, without longitudinal carina or peak but with faint transversal ridge, mesoscutellar appendage with short longitudinal carina; distance between cenchri 2.2 times longest axis of a cenchrus; mesepisternum weakly elevated at middle, ventral thorn absent (Fig. 3D). Hind tibia 0.8 times as long as tarsus, inner tibial spur 0.55 times as long as basitarsus; basitarsus slender, 0.8 times as long as following 3 tarsomeres together; claw with a short and obtuse basal lobe, inner tooth distinctly shorter than outer tooth. Vein cu-a in forewing joining cell 1M at basal 0.35, cell 2Rs distinctly longer than cell 1Rs; hind anal cell sessile (Fig. 1C). Lancet narrow and long with 17 serrulae (Fig. 3C), serrulae distinctly oblique, middle serrulae each with 1 inner subbasal tooth and 11���13 distal subbasal teeth, annular sutures strongly oblique, annular spines bands very narrow, annular spines very short and sparse; 5 th to 7 th serrulae as in Figure 3H. Male. Body length 8.5���9.5 mm (Fig. 1D). Body color and structure similar to female but hind coxa with narrow black dorsal and ventral stripes, malar space 0.5 times diameter of lateral ocellus, distance between eyes below about 0.7 times longest axis of eye, in dorsal view head shorter and distinctly narrowed, basal lobe of claw distinct; subgenital plate 1.2 times as long as broad, apex roundly and narrowly convex; harpe longer than broad, apex of parapenis almost truncate without inner process (Fig. 3F); penis valve broad with a long and narrow apical process (Fig. 3G). Material examined. Holotype. ♀, China, Sichuan Province, Mt. Emei, Leidongping (29��55���N, 103��33���E; elev. 2350 m), 7 July 2009, Meicai Wei leg. Paratypes. 1♀, Sichuan Province, Mt. Emei, Leidongping (29��55���N, 103��33���E; elev. 2350 m), 7 July 2009, Meicai Wei leg.; 2♀ 2♂, Sichuan Province, Mt. Emei, Leidongping (29��55���N, 103��33���E; elev. 2350 m), 8 July 2009, Meicai Wei leg.; 1♀ 1♂, Sichuan Province, Mt. Emei, Leidongping (29��55���N, 103��33���E; elev. 2350 m), 8 July 2009, Gengyun Niu leg.; 2♀ 3♂, Sichuan Province, Mt. Emei, Leidongping (29��55���N, 103��33���E; elev. 2350 m), 7 July 2009, Zejian Li leg.; 1♀ 2♂, Sichuan Province, Mt. Emei, Jinding (29��31.37���N, 103��21.22���E; elev. 3073 m), 20 July 2011, Junzhe Xue & Hu Ping leg.; 1♀, Tibet, Chaya County, G241 (30��42.59���N, 97��19.56��� E; elev. 3350 m), 24 June 2009, Zejian Li leg. Distribution. China (Sichuan, Tibet). Etymology: The specific epithet is composed of a Latin prefix mega- and maculata, referring to the abdomen with large black maculae., Published as part of Xiao, Wei, Niu, Gengyun & Wei, Meicai, 2020, Two new species of the Tenthredo grahami group from China (Hymenoptera: Tenthredinidae), with a revised key to species, pp. 118-125 in Zoological Systematics 45 (2) on pages 122-123, DOI: 10.11865/zs.202016, http://zenodo.org/record/4617204

Tenthredo nigrobullifera sp. nov. (Figs 1 A–B, 2) Diagnosis. This new species is similar to T. bullifera Malaise, 1945, but differs from the latter by the body having much more black maculae, the hairs on dorsum of head much longer and apically curved, the pterostigma black, and the male penis valve with a broad triangular apical process. Description. Female. Body length 12.0–14.0 mm (Fig. 1A). Head and thorax black, following parts yellowish green in alive: palpi, basal 2/5 of mandible, labrum, clypeus except narrow basal margin, triangular supraclypeal area (Fig. 2A), supraantennal tubercles, narrow inner orbit and connected subtriangular macula on temple, lateral stripes of postocellar area (Fig. 2B), lower 3/5 of hind orbit, lower anterior lobe and lateral corner of pronotum, tegula, a V-shaped stripe on posterior of mesoscutal middle lobe, mesoscutellum except narrow hind margin, mesoscutellar appendage, metascutellum (Fig. 1A), narrow base of metapostnotum, two big maculae on mesepisternum and entire metepisternum (Fig. 2D). Abdomen yellowish green, broad basal band of each segment black (Fig. 1A), upper margin of basal plate black, ovipositor apical sheath brown (Fig. 2E). Leg black, outer stripe on each coxa, anterior side of fore femur largely, narrow dorsal stripe on middle femur and apex of hind femur, middle 3/5 of each tibia, 1st to 4th tarsomeres except apex and 5th tarsomere entirely yellow green. Hairs on head and thorax black brown, pubescence on labrum, margin of clypeus and most of tibia and tarsi light brown. Wings hyaline, apex of vein C and pterostigma black, vein C except apex and R1 pale brown. Dorsum of head (Fig. 2B) strongly shiny, black part with fine microsculptures, impunctate; mesonotum with fine and dense microsculpture, weakly shiny, margin of mesonotum with small punctures; mesoscutellum largely shining, posterior margin of mesoscutellum and lateral sides of mesoscutellar appendage with fine microsculpture; upper half of mesopleuron weakly microsculptured, lower half finely and densely microsculptured, punctures indistinct; metapleuron largely polished; all abdominal tergites distinctly microsculptured, weakly shiny; lateral of apical sheath weakly microsculptured, weakly shiny. Length of hairs on dorsum of head, mesonotum, mesoscutellum and mesopleuron about 2, 1, 1.5 and 1.3 times diameter of lateral ocellus, respectively, apex of hairs on dorsum of head distinctly curved. Apex of labrum round; clypeus flat and much broader than distance between eyes below, anterior margin emarginated to a depth of 0.25 times length of clypeus (Fig. 2B); malar space 1.2 times diameter of lateral ocellus; distance between eyes below about 0.8 times longest axis of an eye; supraantennal tubercles strongly elevated, free-standing and parallel to each other, height 0.9 times its width, length about 2 times its height and 1.5 times the length of postocellar area, distance between tubercles 0.9 times its height, posterior end abruptly cut off from frontal ridges, bottom of median fovea with distinct carina; interocellar furrow deep and straight, postocellar furrow shallow; postocellar area flat, without middle carina or middle furrow, width 1.5 times its length; lateral furrows slightly curved, distinctly divergent backwards; temple in dorsal view 0.8 times length of eyes, both sides obtusely convex (Fig. 2B); occipital carina sharp and complete, low part without fold. Antenna as long as head and thorax together, 3rd antennomere 1.65 times length of 4th antennomere, subapical antennomeres not dilated, 8th antennomere twice as long as board (Fig. 2I). Mesoscutellum obtusely elevated, slightly higher than mesonotum, without middle longitudinal and transversal carina; mesoscutellar appendage with low and short middle carina; distance between cenchri 2.7 times longest axis of a cenchrus; middle of mesepisternum weakly and subtriangularly elevated, without sharp peak or carina, ventral thorn absent. Hind tibia 0.85 times as long as tarsus, basitarsus slightly shorter than following three tarsomeres together, inner tibial spur about 0.65 times length of basitarsus, basitarsus slender, tarsal pulvillus developed, distance between basal two pulvilli 2 times length of 1st pulvillus; claw with a small basal lobe, inner tooth shorter than outer tooth. Vein cu-a in fore wing joining cell 1M at basal 0.33, cell 2Rs longer than 1Rs; hind anal cell sessile. Ovipositor sheath slightly bent ventrally, apical sheath narrow, about 2 times length of basal sheath, apical margin round (Fig. 2E). Setae on sheath mostly straight. Lancet very narrow and long with 15 oblique serrulae, annular suture strongly oblique with indistinct annular spine band, ctenidia weakly developed (Fig. 2C); 5th–7th serrulae as Figure 2H, each with about 14 small outer subbasal teeth, spiculella small but distinct. Male. Body length 11.0–12.0 mm (Fig. 1B). Body color and structure similar to female, but sometimes mesosternum with a middle triangular green spot, malar space about 0.5 times diameter of lateral ocellus, in dorsal view temple 0.5 times as long as eyes; subgenital plate slightly longer than broad, apex narrowly truncate; gonoforcep as in Figure 2F, harpe longer than broad, apex round; penis valve as in Figure 2G, valviceps with a short and triangular apical lobe. Material examined. Holotype. ♀, China, Tibet, Motuo County, 24K (29°47′N, 95°42′E; elev. 3663 m), 21 June 2009, Meicai Wei leg. Paratypes. 5♀ 3♂, Tibet, Motuo County, Lage (29°34′N, 94°59′E; elev. 3740 m), 15 June 2009, Zejian Li leg.; 6♀ 1♂, Tibet, Motuo County, Hanmi (29°22′N, 95°7′E; elev. 2180 m), 16 June 2009, Zejian Li leg.; 1♀, Tibet, Motuo County, 2K (29°45′N, 95°43′E; elev. 3272 m), 17 June 2009, Gengyun Niu leg.; 2♂, Tibet, Motuo County, 2K (29°45′N, 95°43′E; elev. 3272 m), 17 June 2009, Meicai Wei leg.; 7♀ 11♂, Tibet, Motuo County, 24K (29°47′N, 95°42′E; elev. 3663 m), 21 June 2009, Meicai Wei leg.; 3♀ 5♂, Tibet, Motuo County, 24K (29°47′N, 95°42′E; elev. 3663 m), 21 June 2009, Gengyun Niu leg.; 1♀, Tibet, Motuo County, 30K (29°45′N, 95°42′E; elev. 3808 m), 20 June 2009, Gengyun Niu leg.; 1♀, Tibet, Motuo County, 30K (29°45′N, 95°42′E; elev. 3808 m), 20 June 2009, Meicai Wei leg. Distribution. China (Tibet). Etymology. This new species is named after its body color as it has the richest black spots in the Tenthredo grahami group.

Key to species of Tenthredo grahami group. 1. Frons including frontal walls entirely black.........................................................................................................................................2 Frontal wall green.............................................................................................................................................................................. 10 2. First antennomere yellow green; penis valve without apical process................................................................................................... 3 First antennomere largely or entirely black; penis valve with or without apical process.....................................................................4 3. Mesepisternum green without black stripe; interspaces between punctures on mesonotum not smooth; microsculptured; mesepisternum flat; black frontal macula not extending backward along lateral of postocellar area (Distribution. China (Jilin), Northeastern Asia)............................................................................................................................... T. pseudolivacea Malaise, 1931 Mesepisternum green with a distinct black stripe; interspaces between punctures on mesonotum smooth, without distinct microsculpture; mesepisternum triangularly elevated at middle; black frontal macula much extending backward along lateral of postocellar area (Distribution. Kashmir, Hymalayas)............................................................................ T. nigromaculata Smith, 1878 4. Thorax mainly black, mesepisternum with 2 large yellow green maculae; middle and hind femora entirely black, each tibia yellow green with black base and apex; pterostigma black; postocellar area with a black middle macula (Distribution. China (Tibet))............................................................................................................................................................................... T. nigrobullifera sp. nov. Thorax green, mesepisternum with a narrow black stripe or entirely green; legs green with longitudinal black stripe; pterostigma green or pale brown.............................................................................................................................................................................. 5 5. Supraantennal tubercles quite low, clearly broader than height; black frontal macula transverse elliptical, not extending backward, bottom of middle fovea entirely green; postocellar area more than 2 times as long as broad; mesepisternum flat and entirely green; penis valve with a narrow but not very long apical process (Distribution. China (Gansu, Shaanxi, Henan)).......................................................................................................................................................................................................... T. flatopectalina Wei, 2002 Supraantennal tubercles high, not broader than height; black frontal macula not transverse elliptical, distinctly extending backward, bottom of middle fovea partly or entirely black; postocellar area less than 2 times as long as broad; mesepisternum elevated at middle or with a distinct black stripe; penis valve with or without apical process...........................................................................................6 6. Dorsum of head with an ��- like black macula ending with a hook near posterior corner of postocellar area; ratio of length and breadth of supraantennal tubercles not less than 3; penis valve without apical process (Distribution. Widespread in North and Central China, Russia Far East and Japan)......................................................................................................................... T. omega (Takeuchi, 1936) Dorsum of head with a subquadrate black macula and without a hook near posterior corner of postocellar area; ratio of length and breadth of supraantennal tubercles less than 3; penis valve usually with a distinct apical process......................................................7 7. Anterior incision of clypeus round, deeper than 0.3 times clypeus length; a transverse black band present between clypeus and supraclypeus; each abdominal tergite with a sickle like black band; mesepisternum flat; body length 10 mm. Liaoning...................................................................................................................................................................................................... T. tombi Mallach, 1936 Anterior incision of clypeus shallow, less than 0.25 times clypeus length; a transverse black band absent between clypeus and supraclypeus; 2nd abdominal tergite with a black band much longer than black bands on other tergites; body length 12.0��� 13.5mm........ 8 8. Bottom of middle fovea black; mesepisternum flat and smooth; basal black band on each of abdominal tergite narrow, black band on 2nd tergite clearly narrower than half length of the tergite; ovipositor sheath very narrow; penis valve with a very long and slender apical process (Distribution. China (Hubei, Sichuan, Tibet))....................................................................... T. bullifera Malaise, 1945 Bottom of middle fovea green; mesepisternum distinctly elevated at middle and with fine microsculptures; basal black band on each of abdominal tergite broad, black band on 2nd tergite longer than half length of the tergite; ovipositor sheath broad; penis valve with a short and stout apical process (Distribution. Widespread in North and Central China)............. T. pseudobullifera Wei & Liu, 2013 9. Supraantennal tubercles clearly higher than broad; black bands on abdominal tergites 2���4 large and clearly separated, clearly larger than black bands on tergites 6���8; apical process of penis valve very long and slender (Distribution. China (Henan, Shanxi, Shaanxi and Sichuan)).......................................................................................................................................... T. pseudograhami Wei, 2002 Supraantennal tubercles as high as broad; black bands on abdominal tergites 2���4 very narrow and indistinct, or quite large and not separated at middle; black bands on tergites 6���7 not smaller than black bands on tergites 3���4, or penis valve without apical process..........................................................................................................................................................................................................10 10. Each black band on abdominal tergites not shorter than half length of each tergite........................................................................... 11 Black bands on tergites 1���4 clearly shorter than half length of tergite...............................................................................................12 11. Black bands on abdominal tergites 2���8 as long as half-length of tergite; penis valve round at apex and without process (Distribution. India)................................................................................................................................................. T. gulmargi Singh & Saini, 1987 Black bands on abdominal tergites 2���8 longer than 0.7 times the length of tergite; penis valve with a long apical process (Distribution. China (Sichuan, Tibet))................................................................................................................................ T. megamaculata sp. nov. 12. Black bands on abdominal tergites 2���8 very narrow and about equal in length; penis valve round at apex, without process (Distribution. China (Shanxi, Henan, Sichuan))................................................................................................................. T. grahami Malaise, 1945 Black bands on abdominal tergites 6���7 broad, tergites 2���5 almost absent; penis valve with a very long and slender apical process (Distribution. China (Tibet))............................................................................................................ T. paragrahami Wei & Liu, 2013, Published as part of Xiao, Wei, Niu, Gengyun & Wei, Meicai, 2020, Two new species of the Tenthredo grahami group from China (Hymenoptera: Tenthredinidae), with a revised key to species, pp. 118-125 in Zoological Systematics 45 (2) on page 124, DOI: 10.11865/zs.202016, http://zenodo.org/record/4617204, {"references":["Malaise, R. 1945. Tenthredinoidea of South-Eastern Asia with a general zoogeographical review. Opuscula Entomologica, Lund Supplementum, 4: 1 - 288.","Liu, M. M., Wei, M. C. 2013. Two new species of grahami group of Tenthredo Linnaeus (Hymenoptera, Tenthredinidae) from China. Acta Zootaxonomica Sinica, 38 (4): 841 - 848."]}

Key to the Chinese species of Gilpinia Benson (females; examined species are marked with *) 1. Inner spur of hind tibia scale like......................................................................... 2 - Inner spur of hind tibia simple........................................................................... 4 2. Annuli 2 and 3 divergent downwards (Fig. 2B); pronotum entirely yellow; only posterior margin of mesoscutellum black. China (Heilongjiang), Finland....................................................... * G. fennica (Forsius, 1911) - Annuli 2 and 3 parallel (Figs. 2A, C); pronotum yellow with black marks; both anterior and posterior margins of mesoscutellum black............................................................................................ 3 3. Serrulae 3���5 broader than the ctenidium bear them (Fig. 2A); head with a dark band on ocellar area, not reaching inner margins of eyes; apical margin of clypeus straight; ocellar area with 3 small brownish marks; pronotum pale with anterior margin black. China (Inner Mongolia)............................................... G. baiyinaobaoa G. Xiao & X. Huang, 1985 - Serrulae 3���5 equal to the ctenidium bear them (Fig. 2C); head with a dark band on ocellar area, reaching inner margins of eyes; apical margin of clypeus slightly incised; ocellar area without brownish marks; pronotum black with lateral corners pale. China (Heilongjiang, Jilin), Siberia, Europe.................................................... * G. virens (Klug, 1812) 4. Head black at least between ocellar area and toruli........................................................... 5 - Head pale with black at most on ocellar area above toruli..................................................... 10 5. Head black at least above toruli, but dorsally more or less pale.................................................. 6 - Head black with labrum, clypeus, supraclypeal area and dorsal margin of head more or less pale....................... 8 6. Scape, pedicel and flagellomere 1 black and each basally pale; lateral mesoscutal lobe black with anterior and lateral margins pale; mesoscutellum pale with a central longitudinal black line; legs whitish yellow. Lancet in Fig. 2E. China (Sichuan, Yunnan)................................................................ G. yongrenica G. Xiao & X. Huang, 1984 - Scape and pedicel pale, flagellomere 1 black; lateral mesoscutal lobe black with lateral margins pale; mesoscutellum pale with posterior margin black; legs yellow with black marks or dirty yellow............................................. 7 7. Pectus, lower anterior margin of mesepisternum, lower mesepimeron, and most of metepimeron black; legs yellow with black marks; lancet with basal annuli straight, subparallel (Smith 1974: Fig. 1). China (Fujian), Thailand................................................................................................... * G. marshalli (Forsius, 1931) - Pectus, mesepisternum, mesepimeron, metepimeron pale; legs dirty yellow. Lancet with basal annuli curved, annuli 1 and 2 distinctly divergent (Fig. 2F). China (Heilongjiang)............................. G. pinicola G. Xiao & X. Huang, 1985 8. Body entirely black; wings strongly smoky; lancet with 11 distinct annuli; length of basal ctendium to lancet apex about 2.0�� maximum width of lancet; maximum length of annulus 2 about 2.1�� minimum; annulus 1 narrow and about 0.4�� length of annulus 2 (Fig. 2H). China (Jiangxi).......................................... * G. infuscalae Wang & Wei, sp. nov. - Body black with pale marks; wings transparent; lancet with 9 or 10 distinct annuli; length of basal ctendium to lancet apex at least 2.5�� maximum width of lancet; maximum length of annulus 2 at most 1.6�� minimum; annulus 1 about at least 0.7�� length of annulus 2 (Fig. 2D, G).......................................................................... 9 9. Head black with labrum and ventral margin of clypeus yellow; median mesoscutal lobe and mesoscutellum black; legs yellow, coxae, trochanters and femora basally dark brown, tibiae basally more or less whitish; annuli 2 and 3 divergent downwards, annulus 1 about 0.7�� length of annulus 2 (Fig. 2G). China (Heilongjiang), Japan (Hokkaido)......... G. tohi Takeuchi, 1940 - Head black with labrum, clypeus, supraclypeal area and dorsal margin of head pale; median mesoscutal lobe black with posterior lateral margin pale, mesoscutellum black with a central horizontal pale band; coxae and trochanters black, tarsi, hind femur and apex of tibiae reddish brown, tibiae basally yellow; annulus 2 parallel with 3, annulus 1 about 1.1�� length of annulus 2 (Fig. 2D). China (Anhui)................................................... * G. massoniana G. Xiao, 1992 10. Annulus 1 with a distinct serrula (Fig. 2 I)................................................................. 11 - Annulus 1 without serrula (Figs. 2 J-M)................................................................... 12 11. Body reddish brown; ocellar area with a short dark band; mesoscutellum, central part of pronotum and most of lateral mesoscutal lobes black (median mesoscutal lobes sometimes with black triangular marks); tergum 1 black. Lancet in Fig. 2 I. China (Gansu).................................................................... * G. tabulaeformis G. Xiao, 1992 - Body yellowish brown; ocellar area pale without black marks; pronotum, median mesoscutal lobe, lateral mesoscutal lobe and tergum 1 pale; mesoscutellum black. China (Hebei).............................. G. funingensis Wen, Sun & Li, 1991 12. Ctenidium 1 broken at middle (Fig. 2M); lateral mesoscutal lobe entirely pale. China (Anhui, Guangxi).......................................................................................* G. lipuensis G. Xiao & X. Huang, 1985 - Ctenidium 1 entire; lateral mesoscutal lobe pale with dark marks or entirely black................................. 13 13. Annuli 1 and 2 straight and parallel (Fig. 2J); body yellow; mesoscutellum entirely pale; sheath with lateral margin of scopa slenderly protruding. China (Yunnan).................................................. * G. hebedentata Xu, 1997 - Annuli 1 and 2 curved and divergent downwards (Fig. 2K, L); body reddish brown, mesoscutellum entirely black or pale with posterior margin black; sheath with lateral margin of scopa more or less flat...................................... 14 14. Annuli 1 and 2 distinctly divergent ventrally, basal serrulae acute (Fig. 2K); labrum black; flagellomeres dorsally reddish brown, ventrally black; dorsal mesonotum mostly pale, median mesoscutal lobe pale, both lateral mesoscutal lobes with central longitudinal dark stripes; mesoscutellum with posterior margin black. China (Yunnan).................................................................................................. G. jinghongensis G. Xiao & X. Huang, 1984 - Annuli 1 and 2 weakly divergent ventrally, basal serrulae truncate (Fig. 2L); labrum reddish brown; flagellomeres black; dorsal mesonotum mostly black; both median mesoscutal lobe with posterior dark marks; lateral mesoscutal lobes and mesoscutellum entirely black. China (Yunnan, Guizhou)......................................* G. jingxii G. Xiao & X. Huang, 1984, Published as part of Wang, Hannan, Smith, David R., Xiao, Wei, Niu, Gengyun & Wei, Meicai, 2019, Gilpinia infuscalae Wang & Wei, sp. nov. and a key to the Chinese Gilpinia species (Hymenoptera: Diprionidae), pp. 589-596 in Zootaxa 4571 (4) on page 591, DOI: 10.11646/zootaxa.4571.4.11, {"references":["Xiao, G. R., Huang, X. Y. & Zhou, S. Z. (1985) The Chinese sawflies of the family Diprionidae (Hymenoptera, Symphyta). Scientia Silvae Sinicae, 21 (1), 30 - 43.","Klug, J. C. F. (1812) Die Blattwespen nach ihren Gattungen und Arten zusammengestellt. Der Gesellschaft Naturforschender Freunde zu Berlin Magazin fur die neuesten Entdeckungen in der gesamten Naturkunde, Berlin, 14 pp. [pp. 49 - 62]","Xiao, G. R., Zhou, S. Z. & Huang, X. Y. (1984) Seven new species of conifer sawflies from China (Hymenoptera: Diprionidae). Entomotaxonomia, 6 (2 - 3), 141 - 150.","Smith, D. R. (1974) Diprionidae in Thailand (Hymenoptera: Symphyta). Proceedings of the Entomological Society of Washington, 76, 212 - 216.","1911), 178 - 183. Forsius, R. (1931) A new Diprion from China. Notulae Entomologicae, 11 (1), 26 - 27. Hara, H. & Shinohara, A. (2015) The Gilpinia abieticola species group (Hymenoptera, Diprionidae). Bulletin of the National","Takeuchi, K. (1940) A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire (III). Tentharedo. Acta Entomologica, 3 (2), 187 - 199.","Wen, X. J., Sun, C. H., Li, W. G., E, X. Q. & Han, Y. S. (1991) Study on Gilpinia funingensis. Forest Pest and Disease, 1991 (1), 14 - 15.","Xu, Z. H. (1997) Two new species and one new combination of the conifer sawfly family Diprionidae (Hymenoptera: Symphyta) from Yunnan, China. Zoological Research, 18 (2), 171 - 176."]}

Wang, Hannan, Smith, David R., Xiao, Wei, Niu, Gengyun, Wei, Meicai (2019): Gilpinia infuscalae Wang & Wei, sp. nov. and a key to the Chinese Gilpinia species (Hymenoptera: Diprionidae). Zootaxa 4571 (4): 589-596, DOI: https://doi.org/10.11646/zootaxa.4571.4.11

Gilpinia infuscalae Wang & Wei, sp. nov. (Figs. 1 A���M, 2H, 3) Female. Body length 8.2 mm (8.0��� 9.6 mm). Wing expanse 16.7 mm (16.2���19.2 mm). Mainly black (Fig. 1A); mandible basally black, apically reddish brown; palpi yellowish brown; antenna black with scape and pedicel dark brown; cenchri brownish; terga 2���6 with faint lateral brownish spots (Fig. 1E). Legs black with followings dark brown: claws, spurs, inner sides of fore and middle tibiae. Wings smoky with dark veins and stigma, center of stigma whitish. Head and dorsal mesonotum densely punctured. Posterior and lateral margins of head and ocellar area with dense punctures about 0.02 mm, interspaces between punctures less than diameter of a puncture; clypeus (apical margin and central part shiny without punctures), malar space and lower paraantennal field with dense punctures about 0.07 mm, interspaces between punctures less than diameter of a puncture; anterior and posterior margins of median mesoscutal lobe, lateral side of lateral mesoscutal lobe with punctures about 0.03 mm (Fig. 1D); mesoscutellum and center of mesepisternum with punctures about 0.1 mm (Fig. 1F); katepimeron and metepisternum feebly shiny and wrinkled; postspiracular sclerite, metascutellum, anepimeron, metepimeron and center of tergum 1 (Fig. 1E) with dense punctures about 0.07 mm; terga wrinkled with few shallow punctures; ventral terga also with punctures, gradually denser toward lateral margin; tergum 10 sparsely punctured, gradually dense and distinct to the posterior end (Fig. 1H); sterna with dense punctures. Head and mesonotum covered with dense whitish hair 0.9�� diameter of front ocellus. Clypeus concave apically; frontal field and frontal pit distinct; postocellar area rounded, frontal postocellar furrow concave, lateral postocellar furrows faint; distances between eye and lateral ocellus, between lateral ocelli, and between lateral ocellus and posterior margin of head about 1.0: 1.0: 1.0 (Fig. 1C), distance between eye and torulus 1.7���1.8�� distance between toruli; malar space 1.4���1.5�� diameter of front ocellus; antenna serrate with 19 distinct antennomeres, apex obtuse, 8���11th rami longer than stem of each flagellomere, length of pedicel equal to diameter of front ocellus, length of ramus 1 equal to length of flagellomere 1. Mesoscutellum obtusely protruding with an anterior angle of 106��, its width 1.2���1.3�� length; distance between cenchri equal to length of a cenchrus. Inner hind tibial spur simple and slightly longer than outer spur, apex obliquely truncate, its length 0.4���0.5�� length of tarsomere 1 (including plantar lobe) and 0.3���0.4�� maximum width of hind tibia, total length of tarsomeres 2���5 about 3.0���3.1�� length of tarsomere 1; claw with an obvious denticle. Dorsal width of scopa 3.7���3.9�� width of a cercus (Fig. 1G), ventral length of a scopa 4.3���4.4�� length of a cercus (Fig. 1H); each scopa apically obtuse, slightly protruding beyond tergum 9 in lateral view; dorsal margin of lance convex at center in lateral view (Fig. 1I), apices of lances asymmetrical; lancet with 11 distinct annuli, gradually narrowing from annulus 2 toward apex, annulus 1 very narrow, about 0.4�� length of annulus 2 (Fig. 1J); annulus 1 without serrula, annulus 2 with longest serrula, close to and parallel with annulus 1, remote and distinctly divergent downwards with annulus 3; width of annulus 2 about 1.9�� width of annulus 3, total length of annuli 1���11 about 2.0�� maximum width of lancet, annuli 3���9 parallel (Fig. 1J); serrulae slightly concave ventrally (Fig. 1K). Male. Body length 6.7 mm (6.5���8.0 mm), wing expanse 14.1 mm (14.0���15.0 mm) (Fig. 1B). Mainly black, labrum and pterostigma brownish, cenchri dark brown, wings hyaline, apical 1/3 of fore femur, apical 1/4 of middle femur, all tarsi and claws orange brown. Punctation as in female, but generally denser and more distinct on head and mesonotum, shallower and sparser on ventral terga. Antenna with 21 distinct antennomeres, scape length 3.0�� length of pedicel, flagellomeres 3���19 pectinate, flagellomere 1 bears a long ramus. Distances between eye and lateral ocellus, between lateral ocelli, between lateral ocellus and posterior margin of head about 1.0:1.0:0.8. distance between eye and torulus 2.0���2.1�� distance between toruli; malar space 1.3���1.4�� diameter of front ocellus. Inner spur of hind tibia simple, 0.4���0.5�� length of tarsomere 1 (including plantar lobe), 0.5���0.6 �� maximum width of hind tibia, total length of tarsomeres 2���5 about 2.5���2.6�� length of tarsomere 1. Gonoforceps as Fig. 1L, parapenis long, gradually narrowing toward apex, inner margins of harpes slightly concave, valviceps simple with blunt apex and a multitoothed ventral margin (Fig. 1M), length of penis valve 4.0���4.1�� maximum width of valviceps. Larva (Fig. 3). Body yellow with head and thoracic legs black. Larvae found at 2012.11.0 6, gathered by collector from Pinus massoniana Lamb. (Pinaceae), reared in room temperature. Pupae to adult from 2013.03 to 2013.04. Host plant. Pinus massoniana Lamb. (Pinaceae). Type material. Holotype ♀, Jiangxi, Huichang, 2013.3.29, Maoling Sheng, pupa reared, Pinus massoniana Lamb. (Pinaceae). Paratypes. 18 ♀, 44 ♂: Jiangxi, Huichang, 2013.3.23���4.16, leg. Maoling Sheng, pupa reared, Pinus massoniana Lamb. (Pinaceae). Distribution. China (Jiangxi Province). Etymology. The specific name " infuscalae " is a noun, derived from Latin, " infusc -" (blacked) and " alae " (wings), referring to its largely smoky wings. Remarks. Gilpinia infuscalae is distinguished from other Gilpinia species by its entirely black color, simple inner hind tibial spur, and lancet which is triangular and with the second and third annuli distinctly divergent (Fig. 1J). It is the only entirely black Gilpinia species in China. Other Chinese species are mostly yellow, brown, or reddish brown and have hyaline wings. In color, G. infuscalae resembles G. nigra (Okutani, 1958) from Japan, also an entirely black species. However, G. nigra has more hyaline wings, the inner hind tibial spur is scale like, the anterior four tibiae and tarsi are brown, the cenchri are black, and the lancet is more slender, resembling the species in Fig. 2 A���E. Structurally, G. infuscalae is similar to those species with more triangular lancets, as in Figs. 2 I���M, but the second and third annuli are distinctly divergent in G. infuscalae. Gilpinia bodyarensis from India has smoky wings as well, but the wings are not as dark as in G. infuscalae. It also differs in having yellow marks on the head and with the first to third annuli almost parallel (Saini & Thind 1993: fig. 15)., Published as part of Wang, Hannan, Smith, David R., Xiao, Wei, Niu, Gengyun & Wei, Meicai, 2019, Gilpinia infuscalae Wang & Wei, sp. nov. and a key to the Chinese Gilpinia species (Hymenoptera: Diprionidae), pp. 589-596 in Zootaxa 4571 (4) on pages 592-594, DOI: 10.11646/zootaxa.4571.4.11, {"references":["Okutani, T. (1958) Three new species of Japanense Symphyta (Studies on Symphyta XI). Mushi, 32 (16), 143 - 147.","Saini, M. S. & Thind, A. S. (1993) Revision of genus Gilpinia Benson from India (Hymenoptera, Symphyta, Diprionidae). Deutsche entomologische Zeitschrift, 40 (2), 309 - 318. https: // doi. org / 10.1002 / mmnd. 19930400211"]}

Gilpinia infuscalae Wang & Wei, sp. nov. (Figs. 1 A–M, 2H, 3) Female. Body length 8.2 mm (8.0– 9.6 mm). Wing expanse 16.7 mm (16.2–19.2 mm). Mainly black (Fig. 1A); mandible basally black, apically reddish brown; palpi yellowish brown; antenna black with scape and pedicel dark brown; cenchri brownish; terga 2–6 with faint lateral brownish spots (Fig. 1E). Legs black with followings dark brown: claws, spurs, inner sides of fore and middle tibiae. Wings smoky with dark veins and stigma, center of stigma whitish. Head and dorsal mesonotum densely punctured. Posterior and lateral margins of head and ocellar area with dense punctures about 0.02 mm, interspaces between punctures less than diameter of a puncture; clypeus (apical margin and central part shiny without punctures), malar space and lower paraantennal field with dense punctures about 0.07 mm, interspaces between punctures less than diameter of a puncture; anterior and posterior margins of median mesoscutal lobe, lateral side of lateral mesoscutal lobe with punctures about 0.03 mm (Fig. 1D); mesoscutellum and center of mesepisternum with punctures about 0.1 mm (Fig. 1F); katepimeron and metepisternum feebly shiny and wrinkled; postspiracular sclerite, metascutellum, anepimeron, metepimeron and center of tergum 1 (Fig. 1E) with dense punctures about 0.07 mm; terga wrinkled with few shallow punctures; ventral terga also with punctures, gradually denser toward lateral margin; tergum 10 sparsely punctured, gradually dense and distinct to the posterior end (Fig. 1H); sterna with dense punctures. Head and mesonotum covered with dense whitish hair 0.9× diameter of front ocellus. Clypeus concave apically; frontal field and frontal pit distinct; postocellar area rounded, frontal postocellar furrow concave, lateral postocellar furrows faint; distances between eye and lateral ocellus, between lateral ocelli, and between lateral ocellus and posterior margin of head about 1.0: 1.0: 1.0 (Fig. 1C), distance between eye and torulus 1.7–1.8× distance between toruli; malar space 1.4–1.5× diameter of front ocellus; antenna serrate with 19 distinct antennomeres, apex obtuse, 8–11th rami longer than stem of each flagellomere, length of pedicel equal to diameter of front ocellus, length of ramus 1 equal to length of flagellomere 1. Mesoscutellum obtusely protruding with an anterior angle of 106°, its width 1.2–1.3× length; distance between cenchri equal to length of a cenchrus. Inner hind tibial spur simple and slightly longer than outer spur, apex obliquely truncate, its length 0.4–0.5× length of tarsomere 1 (including plantar lobe) and 0.3–0.4× maximum width of hind tibia, total length of tarsomeres 2–5 about 3.0–3.1× length of tarsomere 1; claw with an obvious denticle. Dorsal width of scopa 3.7–3.9× width of a cercus (Fig. 1G), ventral length of a scopa 4.3–4.4× length of a cercus (Fig. 1H); each scopa apically obtuse, slightly protruding beyond tergum 9 in lateral view; dorsal margin of lance convex at center in lateral view (Fig. 1I), apices of lances asymmetrical; lancet with 11 distinct annuli, gradually narrowing from annulus 2 toward apex, annulus 1 very narrow, about 0.4× length of annulus 2 (Fig. 1J); annulus 1 without serrula, annulus 2 with longest serrula, close to and parallel with annulus 1, remote and distinctly divergent downwards with annulus 3; width of annulus 2 about 1.9× width of annulus 3, total length of annuli 1–11 about 2.0× maximum width of lancet, annuli 3–9 parallel (Fig. 1J); serrulae slightly concave ventrally (Fig. 1K). Male. Body length 6.7 mm (6.5–8.0 mm), wing expanse 14.1 mm (14.0–15.0 mm) (Fig. 1B). Mainly black, labrum and pterostigma brownish, cenchri dark brown, wings hyaline, apical 1/3 of fore femur, apical 1/4 of middle femur, all tarsi and claws orange brown. Punctation as in female, but generally denser and more distinct on head and mesonotum, shallower and sparser on ventral terga. Antenna with 21 distinct antennomeres, scape length 3.0× length of pedicel, flagellomeres 3–19 pectinate, flagellomere 1 bears a long ramus. Distances between eye and lateral ocellus, between lateral ocelli, between lateral ocellus and posterior margin of head about 1.0:1.0:0.8. distance between eye and torulus 2.0–2.1× distance between toruli; malar space 1.3–1.4× diameter of front ocellus. Inner spur of hind tibia simple, 0.4–0.5× length of tarsomere 1 (including plantar lobe), 0.5–0.6 × maximum width of hind tibia, total length of tarsomeres 2–5 about 2.5–2.6× length of tarsomere 1. Gonoforceps as Fig. 1L, parapenis long, gradually narrowing toward apex, inner margins of harpes slightly concave, valviceps simple with blunt apex and a multitoothed ventral margin (Fig. 1M), length of penis valve 4.0–4.1× maximum width of valviceps. Larva (Fig. 3). Body yellow with head and thoracic legs black. Larvae found at 2012.11.0 6, gathered by collector from Pinus massoniana Lamb. (Pinaceae), reared in room temperature. Pupae to adult from 2013.03 to 2013.04. Host plant. Pinus massoniana Lamb. (Pinaceae). Type material. Holotype ♀, Jiangxi, Huichang, 2013.3.29, Maoling Sheng, pupa reared, Pinus massoniana Lamb. (Pinaceae). Paratypes. 18 ♀, 44 ♂: Jiangxi, Huichang, 2013.3.23–4.16, leg. Maoling Sheng, pupa reared, Pinus massoniana Lamb. (Pinaceae). Distribution. China (Jiangxi Province). Etymology. The specific name " infuscalae " is a noun, derived from Latin, " infusc -" (blacked) and " alae " (wings), referring to its largely smoky wings. Remarks. Gilpinia infuscalae is distinguished from other Gilpinia species by its entirely black color, simple inner hind tibial spur, and lancet which is triangular and with the second and third annuli distinctly divergent (Fig. 1J). It is the only entirely black Gilpinia species in China. Other Chinese species are mostly yellow, brown, or reddish brown and have hyaline wings. In color, G. infuscalae resembles G. nigra (Okutani, 1958) from Japan, also an entirely black species. However, G. nigra has more hyaline wings, the inner hind tibial spur is scale like, the anterior four tibiae and tarsi are brown, the cenchri are black, and the lancet is more slender, resembling the species in Fig. 2 A–E. Structurally, G. infuscalae is similar to those species with more triangular lancets, as in Figs. 2 I–M, but the second and third annuli are distinctly divergent in G. infuscalae. Gilpinia bodyarensis from India has smoky wings as well, but the wings are not as dark as in G. infuscalae. It also differs in having yellow marks on the head and with the first to third annuli almost parallel (Saini & Thind 1993: fig. 15).

Neocolochelyna (Neocolochelyna) acutilabria Niu & Wei, sp. nov. ( Figs 1 c���d, 3 d���i, 6 b���c, 7 b���c, 7 k���l, 8 d, 8 h, 8 l, 9 d, 9 h, 10 a���b, 11 b���c, 11 k���l ) Description. Female. Holotype (Fig. 1 c). Length 15 mm. Head brown to orange, labrum yellow brown, narrow posterior margin of clypeus, supraclypeal area, middle fovea and lateral foveae, lower 0.6 of inner orbit and postocellar furrow black (Figs 3 d���e), lower part of posterior side of head blackish; antenna entirely brown, basal part hardly paler than apex (Fig. 11 b). Prothorax black, narrow lateral margin and very broad posterior margin of pronotum brown; mesoscutal middle lobe and lateral lobes dark brown, centre of middle lobe and dorsal broad stripe on lateral lobe black brown to black, mesoscutellum pale brown, parapsis largely and mesoscutellar appendage black; metanotum pale brown, a dot lateral to metascutellum and most of metapostnotum blackish, cenchrus yellow (Fig. 6 b); mesopleuron and metapleuron black, a large elliptical macula near dorsal corner of mesepisternum, narrow posterior margin of mesepimeron, dorsal margin of metapleuron brown. About basal half of abdominal tergites 1���8 black, a narrow stripe covering spiracle and about apical half of tergites 1���8, tergites 9��� 10 entirely, cerci and ovipositor sheath pale brown, about basal 0.7 of each sternite black brown. Body hairs brown to pale brown. Legs orange, ventral side of fore and middle coxae, hind coxa except for lateral stripe, black, hind tarsus yellow brown. Wings yellowish infuscate in anterior part and gradually paler toward posterior margin, vein C and pterostigma pale brown, posterior of vein Sc+R and most of vein M+Cu blackish; hind wing weakly infuscate, slightly paler than forewing. Punctures on labrum small and sparse, broad interspaces between punctures smooth, shiny; punctures on clypeus dense and large, narrow interspaces between punctures smooth, shiny; frons and inner orbit coarsely punctured, rugose, mat (Fig. 3 e); hind orbit densely punctured, interspaces linear, weakly microsculptured, less shiny; punctures on temple and postocellar area sparser than inner orbit, interspaces smaller than punctures, surface clearly but weakly microsculptured, less shiny (Fig. 3 f); malar space densely punctured; upper corner of pronotum densely punctured and almost dim, depressed part of pronotum coarsely punctured and microsculptured, almost mat, narrow lateral margin of pronotum shiny; mesoscutal middle lobe and lateral lobe densely punctured, interspaces very narrow, distinctly microsculptured, punctures on posterior part of lateral lobe smaller and much denser, almost dim; punctures on mesoscutellum large and dense, interspaces microsculptured, less shiny; mesoscutellar appendage weakly but clearly coriaceous, impunctate; parapsis densely microsculptured, metapostnotum strongly microsculptured (Fig. 6 b); punctures on upper half of mesepisternum large and very dense, interspaces very narrow, hardly shiny; punctures on lower half of mesepisternum small and much sparse, surface distinctly microsculptured, weakly shiny; mesepimeron and metapleuron strongly microsculptured mixed with some shallow and indistinct punctures, narrow anterior and posterior margins of mesepimeron shiny; abdominal tergite 1 strongly microsculptured, almost mat, center and lateral side with some shallow and large punctures, very narrow posterior margin weakly coriaceous, less shiny (Fig. 6 b); tergites 2���8 densely microsculptured with a few shallow and indistinct punctures laterally, mat; median part of tergite 8���10 weakly microsculptured, distinctly shiny; surface of basal sheath microsculptured mixed with some shallow punctures, apical sheath hardly coriaceous, shiny. Hairs on dorsum of head and mesopleuron dense, weakly curved at apex, about as long as diameter of median ocellus (Figs 3 d���e); hairs on mesoscutal middle and lateral lobes about 0.8 times as long as diameter of median ocellus; dorsum of abdominal tergites 1���8 distinctly pubescent, hairs on tergite 1 not longer and denser than other tergites; mesoscutellar appendage and intertergites of abdomen glabrous (Fig. 6 b). Labrum triangular, apically acute; clypeus with a low middle ridge, anterior margin almost truncate, with a very short but distinct middle tooth; malar space as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.5 times as long as longest axis of eye (Fig. 3 e); mandibles as Figs 11 k���l; hind orbit with a distinct occipital carina in lower half; middle fovea roundish, shallow, narrowly open to frons; lateral fovea indistinct, with a small tubercle; frontal ridge broad and round; POL: OOL: OCL= 15: 35: 31; interocellar furrow short and very deep, postocellar furrow narrow and distinct, triangularly bent forward at middle; postocellar area almost flat, 1.35 times as broad as long, posterior margin elevated; lateral furrows very shallow and indistinct in anterior 0.7, replaced by a low carina in posterior 0.3, hardly curved, distinctly divergent backwards; head roundly dilated behind eyes in dorsal view (Fig. 3 d). Antenna as long as thorax, antennomere 3 about 1.2 times as long as antennomeres 4 and 5 together and 1.1 times as long as antennomeres 6���9 together, antennomere 7 about as long as broad (Fig. 11 b). Mesoscutal middle lobe with narrow but distinct middle furrow; mesoscutellum roundly elevated, anterior margin almost truncate, hardly protruding, transverse anterior furrow deep; mesoscutellar appendage short with middle length about 1.1 times as long as diameter of median ocellus; mesoscutellum 4 times as long as mesoscutellar appendage; cenchrus transverse, much broader than long, CD= 1.3 (Fig. 6 b); mesepisternum roundly and weakly elevated at middle, without distinct ridge. Venation as Fig. 1 c, cell 2 Rs about 0.75 times as long as cells 1 R 1 + 1 Rs, cell A of forewing weakly constricted at basal 0.25, where distance between 1 A and 2 A+ 3 A 4 times as broad as vein 2 A+ 3 A; vein 1 M as long as vein 1 m-cu; vein 2 r 1 meeting cell 2 Rs at apical 0.4. Inner tibial spur of fore leg as Fig. 7 k, apical incision very shallow; claw as Fig. 7 b, inner tooth clearly longer than apical tooth. Triangular middle lobe of sternite 7 large and broad, much broader than lateral incision of sternite 7 (Fig. 8 l). Ovipositor sheath 0.9 times as long as front tibia and 0.7 times as long as middle tibia, apical sheath 1.1 times as long as basal sheath, narrowly truncate at apex in lateral view. Lancet (paratype ♀) as Fig. 8 d, with 25 serrulae, middle and apical segments as Figs 9 d, 9 h; annular spines and surface sculptures of lancet as Fig. 9 h; basal sensilla as Fig. 8 h. Male. Paratype (Fig. 1 d). Body length 13.5 mm; color, punctation and structure similar to female except for following: posterior 0.7 of clypeus, frons, upper inner orbits largely, postocellar area, lower half of hind orbit, mesoscutal middle and lateral lobes black; malar space 0.8 times as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.3 times as long as longest axis of eye (Fig. 3 h); dorsum of head and mesonotum coarsely rugose, punctures irregular, not clearly defined (Fig. 3 g, 3 i); subgenital plate 1.1 times as broad as long, apical margin round; harpes, volsella and gonolacinia as Fig. 10 b; penis valve as Fig. 10 a, dorsal apical hook very long and strongly tapering toward apex. Variation. In a paratype female, the CD is about 1.5 and the vein 2 r 1 meets cell 2 Rs at middle. Distribution. China (Xizang) (Fig. 12). Etymology. This species is named after the triangular labrum which is acute at apex. The name is to be treated as a noun. Material examined. Holotype: ♀, CHINA: XIZANG: Yigong, 2300 m, Sichuan Prov. [Xizang], 28. iv. 1986, T. Naito leg.; 86. 4. 28, 2, Xizang, Yigong; (CSCS, lancet dissected and missing, possibly kept in KUK). Paratypes: 1 ♀, data same as the holotype (KUK); 1 ��, data same as the holotype (KUK). Host plant. Unknown. Remarks. Neocolochelyna acutilabria is similar to N. montana Konow. See the above key for the differences between the two species. Two females and one male of Neocolochelyna acutilabria are known. The female specimen with lancet missing was selected as holotype because the specimen is in good state, the second female and the male were partly broken., Published as part of Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko & Naito, Tikahiko, 2016, Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae), pp. 432-456 in Zootaxa 4127 (3) on pages 450-453, DOI: 10.11646/zootaxa.4127.3.2, http://zenodo.org/record/264340

Distribution of Colochela The distribution of Colochela is shown in Fig. 6. The genus is endemic to southeastern Asia at present. The distribution of the three known species overlaps in Sichuan, Chongqing, Shaanxi and Hubei provinces. Within the generic distribution, C. nigrata is distributed in the southeastern area, C. zhongi distributed in the north and C. rufidorsata distributed in the southwestern area. It is possible that the genus is originated and differentiated in the Qinba Mountains of China. Further research on the molecular biogeography of the genus is needed to test this.

Neocolochelyna (Curvatapenis) wui Niu & Wei, sp. nov. (Figs 1 h, 5 a–c, 6g, 7g, 7 p, 10 i –j, 11g) Description. Male. Holotype (Fig. 1 h). Length 12 mm. Body black; labrum brown, palpi of mouthparts dark brown; antenna yellow brown, scape entirely, basal 0.7 of pedicellum and basal half of antennomere 3 black (Fig. 11 g); narrow posterior margin of pronotum, tegula and cenchrus yellow brown; extreme narrow posterior margin of abdominal tergite 1, posterior 0.15 of tergites 3–7, posterior 0.3 of tergite 8, tergite 10 and cerci entirely, posterior 0.3 of sternites 3–7, subgenital plate entirely, yellow brown; tergite 2 entirely, sternite 2 entirely and sternite 3 largely yellow white. Body hairs yellow brown. Coxa entirely, fore and middle trochanters largely black; femora blackish brown, anterior side with broad brownish stripe; fore and middle tibiae entirely, basal 0.3 of hind tibia and all tarsi yellow white, apical 0.7 of hind tibia reddish brown to orange. Wings yellowish infuscate in anterior part and gradually paler toward posterior margin, vein C and pterostigma pale brown, posterior of vein Sc+R and most of vein M+Cu blackish; hind wing weakly infuscate, slightly paler than forewing. Labrum coarsely punctured and distinctly microsculptured; punctures on clypeus very dense and large, interspaces between punctures linear, almost smooth, weakly shiny; head otherwise coarsely and irregularly punctured, rugose, with some short and irregular carinae, interspaces indistinct, strongly microsculptured, mat (Figs 5 a–c); pronotum densely punctured and weakly microsculptured, weakly shiny; mesoscutal middle and lateral lobes densely punctured, interspaces linear, strongly microsculptured, mat, punctures on posterior part of lateral lobe smaller than punctures on anterior part but much denser; mesoscutellum coarsely punctured without interspaces, mat; mesoscutellar appendage smooth, impunctate, shiny; parapsis and metapostnotum strongly microsculptured (Fig. 6 g); punctures on upper half of mesepisternum large and very dense, interspaces linear, hardly shiny; punctures on lower half of mesepisternum small and much sparse, surface hardly microsculptured, strongly shiny; mesepimeron weakly microsculptured in anterior half, posterior half densely punctured; depressed part of metepimeron weakly microsculptured, almost smooth, other part of metapleuron densely punctured, interspaces strongly microsculptured; abdominal tergites 1 distinctly punctured at middle and lateral sides, intermedian area microsculptured, not mat; tergite 2 distinctly but weakly microsculptured (Fig. 6 g), tergites 3–9 strongly microsculptured, rugose, surface mat. Hairs on dorsum of head and mesopleuron dense and distinctly curved at apex, about 1.3 times as long as diameter of median ocellus (Figs 5 a–b); hairs on mesonotum dense, about as long as diameter of median ocellus; hairs on tergite 1 sparse and about 0.8 times as long as diameter of median ocellus; tergite 2 largely glabrous (Fig. 6 g); hairs on tergites 3–8 dense, about 0.7 times as long as diameter of median ocellus; mesoscutellar appendage and intertergites of abdomen glabrous. Labrum short and broad, anterior margin round, not acute; anterior margin of clypeus truncate, without middle tooth; malar space 0.6 times as long as diameter of median ocellus; hind orbit round, occipital carina present in lower 0.3; distance between eyes at level of anterior tentorial pits 1.1 times as long as longest axis of eye (Fig. 5 b); middle fovea round, deep, open to frons; lateral fovea small; frons short, frontal ridge broad; POL: OOL: OCL= 14: 35: 31; interocellar furrow short and very deep, postocellar furrow fine but distinct; postocellar area weakly elevated, about 1.6 times as broad as long; lateral furrow broad and distinct, weakly curved, distinctly divergent backwards; head weakly and roundly narrowed behind eyes in dorsal view (Fig. 5 a). Antenna stout, about as long as thorax, antennomere 3 about 1.1 times as long as antennomeres 4 and 5 together and 0.95 times as long as antennomeres 6–9 together, antennomere 7 as long as broad (Fig. 11 g), outer side of antennomeres 4–9 without longitudinal furrow. Mesoscutal middle lobe without middle furrow, with a fine middle carina; mesoscutellum roundly elevated, anterior margin roundly and distinctly protruding, transverse anterior furrow deep; mesoscutellar appendage short with middle length about as long as diameter of median ocellus; cenchrus transverse, much broader than long, CD= 1.3 (Fig. 6 g). Mesepisternum hardly elevated at middle, without blunt ridge. Venation as Fig. 1 h, cell 2 Rs about 1.2 times as long as cells 1 R 1 + 1 Rs, cell A of forewing strongly constricted at basal 0.25, where distance between 1 A and 2 A+ 3 A 3 times as broad as vein 2 A+ 3 A, first abscissa of vein M 0.9 times as long as 1 m-cu, vein 2 r 1 meeting cell 2 Rs at apical 0.2. Inner tibial spur of fore leg as Fig. 7 p, apex deeply incised; claw as Fig. 7 g, inner tooth clearly shorter than apical tooth. Subgenital plate about as long as broad, apical margin round; harpes, volsella and gonolacinia as Fig. 10 j; penis valve as Fig. 10 i, dorsal apical hook short, connected with caudal corner by membrane. Female. Unknown. Etymology. The species is named after the collector of the holotype. Material examined. Holotype: Ƌ, CHINA: GANSU: Mt. Woniu, Tan'gelinchang, Mt. Xiaolong, 34.4895 °N, 104.7961 °E, 2200m, 2.vii. 2009, Wu Xingyu (CSCS). Distribution. China (Gansu) (Fig. 12). Host plant. Unknown. Remarks. This new species is similar to N. (C.) hyalina sp. nov. See the above key for the differences between them and other species of the genus. The dorsal apical hook of penis valve is not conspicuous as it is connected with the caudal corner of valviceps by membrane. FIGURE 10. Neocolochelyna species. a, c, e, g, i. Penis valve. b, d, f, h, j. Gonoforceps. a–b. N. acutilabria, Ƌ, paratype; c–d. N. testaceoa, Ƌ, paratype; e–f. N. itoi, Ƌ, Minokoshi; g–h. N. hyalina, Ƌ, paratype; i–j. N. wui, Ƌ, holotype. Scale bar = 100 µm (The longer scale applies only to the larger figures of valviceps in the first line. The shorter scale applies to the smaller figures of whole penis valve in first line and the figures of second line).

Neocolochelyna (Curvatapenis) wui Niu & Wei, sp. nov. (Figs 1 h, 5 a���c, 6g, 7g, 7 p, 10 i ���j, 11g) Description. Male. Holotype (Fig. 1 h). Length 12 mm. Body black; labrum brown, palpi of mouthparts dark brown; antenna yellow brown, scape entirely, basal 0.7 of pedicellum and basal half of antennomere 3 black (Fig. 11 g); narrow posterior margin of pronotum, tegula and cenchrus yellow brown; extreme narrow posterior margin of abdominal tergite 1, posterior 0.15 of tergites 3���7, posterior 0.3 of tergite 8, tergite 10 and cerci entirely, posterior 0.3 of sternites 3���7, subgenital plate entirely, yellow brown; tergite 2 entirely, sternite 2 entirely and sternite 3 largely yellow white. Body hairs yellow brown. Coxa entirely, fore and middle trochanters largely black; femora blackish brown, anterior side with broad brownish stripe; fore and middle tibiae entirely, basal 0.3 of hind tibia and all tarsi yellow white, apical 0.7 of hind tibia reddish brown to orange. Wings yellowish infuscate in anterior part and gradually paler toward posterior margin, vein C and pterostigma pale brown, posterior of vein Sc+R and most of vein M+Cu blackish; hind wing weakly infuscate, slightly paler than forewing. Labrum coarsely punctured and distinctly microsculptured; punctures on clypeus very dense and large, interspaces between punctures linear, almost smooth, weakly shiny; head otherwise coarsely and irregularly punctured, rugose, with some short and irregular carinae, interspaces indistinct, strongly microsculptured, mat (Figs 5 a���c); pronotum densely punctured and weakly microsculptured, weakly shiny; mesoscutal middle and lateral lobes densely punctured, interspaces linear, strongly microsculptured, mat, punctures on posterior part of lateral lobe smaller than punctures on anterior part but much denser; mesoscutellum coarsely punctured without interspaces, mat; mesoscutellar appendage smooth, impunctate, shiny; parapsis and metapostnotum strongly microsculptured (Fig. 6 g); punctures on upper half of mesepisternum large and very dense, interspaces linear, hardly shiny; punctures on lower half of mesepisternum small and much sparse, surface hardly microsculptured, strongly shiny; mesepimeron weakly microsculptured in anterior half, posterior half densely punctured; depressed part of metepimeron weakly microsculptured, almost smooth, other part of metapleuron densely punctured, interspaces strongly microsculptured; abdominal tergites 1 distinctly punctured at middle and lateral sides, intermedian area microsculptured, not mat; tergite 2 distinctly but weakly microsculptured (Fig. 6 g), tergites 3���9 strongly microsculptured, rugose, surface mat. Hairs on dorsum of head and mesopleuron dense and distinctly curved at apex, about 1.3 times as long as diameter of median ocellus (Figs 5 a���b); hairs on mesonotum dense, about as long as diameter of median ocellus; hairs on tergite 1 sparse and about 0.8 times as long as diameter of median ocellus; tergite 2 largely glabrous (Fig. 6 g); hairs on tergites 3���8 dense, about 0.7 times as long as diameter of median ocellus; mesoscutellar appendage and intertergites of abdomen glabrous. Labrum short and broad, anterior margin round, not acute; anterior margin of clypeus truncate, without middle tooth; malar space 0.6 times as long as diameter of median ocellus; hind orbit round, occipital carina present in lower 0.3; distance between eyes at level of anterior tentorial pits 1.1 times as long as longest axis of eye (Fig. 5 b); middle fovea round, deep, open to frons; lateral fovea small; frons short, frontal ridge broad; POL: OOL: OCL= 14: 35: 31; interocellar furrow short and very deep, postocellar furrow fine but distinct; postocellar area weakly elevated, about 1.6 times as broad as long; lateral furrow broad and distinct, weakly curved, distinctly divergent backwards; head weakly and roundly narrowed behind eyes in dorsal view (Fig. 5 a). Antenna stout, about as long as thorax, antennomere 3 about 1.1 times as long as antennomeres 4 and 5 together and 0.95 times as long as antennomeres 6���9 together, antennomere 7 as long as broad (Fig. 11 g), outer side of antennomeres 4���9 without longitudinal furrow. Mesoscutal middle lobe without middle furrow, with a fine middle carina; mesoscutellum roundly elevated, anterior margin roundly and distinctly protruding, transverse anterior furrow deep; mesoscutellar appendage short with middle length about as long as diameter of median ocellus; cenchrus transverse, much broader than long, CD= 1.3 (Fig. 6 g). Mesepisternum hardly elevated at middle, without blunt ridge. Venation as Fig. 1 h, cell 2 Rs about 1.2 times as long as cells 1 R 1 + 1 Rs, cell A of forewing strongly constricted at basal 0.25, where distance between 1 A and 2 A+ 3 A 3 times as broad as vein 2 A+ 3 A, first abscissa of vein M 0.9 times as long as 1 m-cu, vein 2 r 1 meeting cell 2 Rs at apical 0.2. Inner tibial spur of fore leg as Fig. 7 p, apex deeply incised; claw as Fig. 7 g, inner tooth clearly shorter than apical tooth. Subgenital plate about as long as broad, apical margin round; harpes, volsella and gonolacinia as Fig. 10 j; penis valve as Fig. 10 i, dorsal apical hook short, connected with caudal corner by membrane. Female. Unknown. Etymology. The species is named after the collector of the holotype. Material examined. Holotype: ��, CHINA: GANSU: Mt. Woniu, Tan'gelinchang, Mt. Xiaolong, 34.4895 ��N, 104.7961 ��E, 2200m, 2.vii. 2009, Wu Xingyu (CSCS). Distribution. China (Gansu) (Fig. 12). Host plant. Unknown. Remarks. This new species is similar to N. (C.) hyalina sp. nov. See the above key for the differences between them and other species of the genus. The dorsal apical hook of penis valve is not conspicuous as it is connected with the caudal corner of valviceps by membrane. FIGURE 10. Neocolochelyna species. a, c, e, g, i. Penis valve. b, d, f, h, j. Gonoforceps. a���b. N. acutilabria, ��, paratype; c���d. N. testaceoa, ��, paratype; e���f. N. itoi, ��, Minokoshi; g���h. N. hyalina, ��, paratype; i���j. N. wui, ��, holotype. Scale bar = 100 ��m (The longer scale applies only to the larger figures of valviceps in the first line. The shorter scale applies to the smaller figures of whole penis valve in first line and the figures of second line)., Published as part of Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko & Naito, Tikahiko, 2016, Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae), pp. 432-456 in Zootaxa 4127 (3) on pages 444-450, DOI: 10.11646/zootaxa.4127.3.2, http://zenodo.org/record/264340

Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko, Naito, Tikahiko (2016): Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae). Zootaxa 4127 (3): 432-456, DOI: http://doi.org/10.11646/zootaxa.4127.3.2

Neocolochelyna (Curvatapenis) itoi Takeuchi, 1951 (Figs 1 f, 1g, 2 c, 4 d���i, 6 e���f, 7 e���f, 7 n���o, 8 a, 8 e, 8 i, 9 a, 9 e, 10 e���f, 11 e���f, 11 o���p) Neocolochelyna itoi Takeuchi, 1951: 62; Takeuchi, 1955: 120; Okutani, 1956: 1; Maxwell, 1958: 19; Togashi, 1965: 246; Togashi, 1995: 870; Yasumatsu, 1957: 101; Okutani et al., 1992: 238; Naito et al., 2004: 50; Shikata & Itakura, 2009: 75. Neocolochelyna hakusana Togashi, 1995: 866; Taeger et al., 2010: 567. Syn. nov. Description. Female. Specimen from Mt. Hakusan (Fig. 1 f). Length 16 mm. Head brown (Figs 4 d���e), labrum and malar space yellowish brown, narrow posterior margin of clypeus, supraclypeal area, frons, lower 0.7 of inner orbit, ocellar area, narrow anterior margin of postocellar area and posterior side of head largely black; basal two antennomeres entirely, basal 0.4 of antennomere 3 yellow brown, apical 0.6 of antennomere 3 and remaining antennomeres dark brown, antennomere 3 more or less black at middle (Fig. 11 e). Thorax black, broad posterior margin of pronotum, tegula, lateral stripe on mesoscutal middle lobe, broad anterior margin and narrow posterior margin of parapsis, mesoscutellum (excluding appendage), metascutellum and cenchrus pale brown. Abdomen brown to pale brown, basal 0.8 (middle) to 0.6 (lateral) of tergite 1 (Fig. 6 e), basal 0.3���0.4 of tergites 3���7, short lateral macula and narrow anterior margin of tergite 2, about basal 0.4���0.7 of each sternite black; ovipositor sheath largely brown. Body hairs brown to pale brown. Legs black, hind trochanter entirely, broad anterior stripe and broad apex of fore and middle femora, fore and middle tibiae entirely pale brown, hind tibia brown to dark brown, all tarsi yellow brown. Wings yellowish infuscate in anterior part and gradually paler toward posterior margin, vein C and pterostigma pale brown, posterior of vein Sc+R and most of vein M+Cu blackish; hind wing weakly infuscate, slightly paler than forewing. Punctures on labrum small and dense, narrow interspaces between punctures weakly microsculptured; punctures on clypeus dense and large, interspaces between punctures smooth, shiny (Fig. 3 e); inner orbit and hind orbit densely punctured, interspaces linear, weakly microsculptured, less shiny; frons densely and coarsely punctured, interspaces very narrow and microsculptured, almost dim; punctures on temple and postocellar area much sparser than frons, surface very weakly microsculptured, moderately shiny (Figs 4 f); malar space densely punctured, shiny; upper corner of pronotum moderately punctured and shiny, depressed part of pronotum coarsely punctured and microsculptured, almost mat; mesoscutal middle lobe and lateral lobe densely punctured, interspaces very narrow, distinctly microsculptured, almost dim; mesoscutellum densely punctured with narrow smooth interspaces, less shiny; mesoscutellar appendage weakly coriaceous, impunctate; parapsis and metapostnotum densely microsculptured (Fig. 6 e); punctures on upper half of mesepisternum large and very dense, interspaces very narrow, hardly shiny; punctures on lower half of mesepisternum small and much sparse, surface indistinctly microsculptured, distinctly shiny; mesepimeron and metapleuron strongly microsculptured mixed with some large but shallow punctures; abdominal tergite 1 densely microsculptured, almost mat, narrow lateral side densely punctured, very narrow posterior margin weakly coriaceous, less shiny; tergite 2 hardly punctured, densely microsculptured (Fig. 6 e), other tergites densely microsculptured mixed with dense or sparse punctures, almost mat; median part of tergite 8���9 weakly microsculptured, feebly shiny; surface of ovipositor sheath weakly microsculptured mixed with some shallow punctures. Hairs on dorsum of head and mesopleuron dense and almost straight, about 0.9 times as long as diameter of median ocellus (Figs 4 d���e); hairs on mesonotum dense and about 0.6 times as long as diameter of median ocellus; dorsum of abdominal tergites 1���3 largely glabrous except for narrow middle stripe (Fig. 6 e), other tergites evenly pubescent; mesoscutellar appendage and intertergites of abdomen glabrous. Labrum broad, anterior margin round, not acute; anterior margin of clypeus very weakly incised in entire length, almost truncate; malar space about as long as diameter of median ocellus (Fig. 4 e); mandibles as Figs 11 o��� p; hind orbit round, without occipital carina; distance between eyes at level of anterior tentorial pits 1.4 times as long as longest axis of eye (Fig. 4 e); middle fovea round, deep, closed; lateral fovea distinct; frons small, very short, frontal ridge broad and round; POL: OOL: OCL= 15: 25: 26; interocellar furrow broad and very deep, postocellar furrow fine and shallow, hardly bent forward; postocellar area roundly elevated, about 1.3 times as broad as long; lateral furrow deep and very broad, weakly curved, distinctly divergent backwards; head roundly dilated behind eyes in dorsal view (Fig. 4 d). Antenna as long as thorax, antennomere 3 about 1.1 times as long as antennomeres 4 and 5 together and 0.8 times as long as antennomeres 6���9 together, antennomere 7 about as long as broad (Fig. 11 e). Mesoscutal middle lobe with narrow but distinct middle furrow; mesoscutellum roundly elevated, anterior margin almost truncate, hardly protruding, transverse anterior furrow deep; mesoscutellar appendage short with middle length about as long as diameter of median ocellus; cenchrus transverse, much broader than long, CD= 1.6 (Fig. 6 e); mesepisternum roundly and weakly elevated at middle, without blunt ridge. Venation as Fig. 1 f, cell 2 Rs about 0.9 times as long as cells 1 R 1 + 1 Rs, cell A of forewing strongly constricted at basal 0.25, where distance between 1 A and 2 A+ 3 A 2.5 times as broad as vein 2 A+ 3 A, first abscissa of vein M 0.9 times as long as 1 m-cu, vein 2 r 1 meeting cell 2 Rs at apical 0.35���0.45. Inner tibial spur of fore leg as Fig. 7 n; claw as Fig. 7 e, inner tooth clearly shorter than apical tooth. Triangular middle lobe of sternite 7 very large and broad, about as broad as lateral straight posterior margin of sternite 7 (Fig. 8 i). Ovipositor sheath 1.1 times as long as front tibia and 0.9 times as long as middle tibia, apical sheath as long as basal sheath, narrowly truncate at apex in lateral view. Lancet as Fig. 8 a, with 26 serrulae, middle and apical segments as Figs 9 a, 9 e, annular spines and surface sculptures of middle of lancet as Fig. 9 e; basal sensilla as Fig. 8 e. Male. Specimen from Minokoshi (Fig. 1 g). Body length 14 mm. Color, punctation and structure similar to female except for following: malar space 0.5 times as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.2 times as long as longest axis of eye (Fig. 4 h); head hardly dilated behind eyes in dorsal view, POL: OOL: OCL= 15: 29: 30 (Fig. 4 g); punctures on temple denser (Fig. 4 i); antenna as Fig. 11 f; subgenital plate about as long as broad, apical margin broadly and obtusely truncate; harpes, volsella and gonolacinia as Fig. 10 f; penis valve as Fig. 10 e, middle tubercle of valviceps long and weakly convex. Variation. Black macula on head sometimes reduced; mesonotum sometimes largely brown to orange; upper of mesepisternum sometimes with a large or small brown macula; abdominal tergite 3 sometimes largely brown, basal black band almost absent; anterior yellow shade of forewing varying from quite deep to light. Material examined. Types. JAPAN: Neocolochelyna itoi Takeuchi, 1951 Holotype: ♀, ���Sikoku, Iya-Otiai, 5.VI. 1951, Issiki et Ito���; ��� Neocolochelyna itoi Take., Holotype ���; ��� Curvatapenis itoi (Takeuchi, 1951), det. M. Wei, 2014 ���(OPU). Paratypes and allotype: 1 ♀, Sikoku, Iya-Nagoro, 2. vi. 1950, Issiki & Ito; Neocolochelyna itoi Tak., Paratype. (OPU). 1 ��, Sikoku, Iya-Nagoro, 2. vi. 1950, Issiki & Ito; Neocolochelyna itoi Tak., Allotype. (OPU). 1 ��, Sikoku, Iya-Nagoro, 2. vi. 1950, Issiki & Ito; Neocolochelyna itoi Tak., Paratype. (OPU). 1 ��, Sikoku, Iya- Nagoro, 5. vi. 1950, Issiki & Ito; Neocolochelyna itoi Tak., Paratype. (OPU). Neocolochelyna hakusana Togashi, 1995: Holotype ♀, ��� 1300-1500m, Mt. Hakusan, Ishikawa Pref., 30.VI. 1994, I. Togashi���; ���NSMT-HYM 62142 ���; ��� Neocolochelyna itoi hakusana sub sp. nov. det. Togashi, 1994 ���; ��� Holotypus ��� [red]; ��� Curvatapenis itoi (Takeuchi), Det. M. C. Wei, 2014 ���; (NSMT). Paratypes: 1 ��, ���Arashimadake, Ohno City, 6.VI. 1982, T. Murota leg.���; ���NSMT-HYM 62143 ���; ��� Paratype ��� [red]; ��� Neocolochelyna hakusana sp. nov. det. Togashi, 1995 ���. (NSMT). 1 ♀, ���Mt. Hakusan (alt. 1300���1500 m), Ishikawa Pref., 30. VI. 1994, I. Togashi leg.���; ��� Neocolochelyna itoi hakusana sub sp. nov., det. Togashi, 1994 ���; ��� Paratype ���[red]; ���NSMT-HYM 2061977 ���; ��� Curvatapenis itoi (Takeuchi, 1951), det. M. Wei, 2012 ���. (USNM). Other specimens examined. JAPAN: HONSHU: Gunma Pref.: 1 ♀, Mt. Kurotakiyama, Minamimakimura, 20.v. 1987, T. Matsumoto (NSMT). Tokyo Met.: 1 ♀, Sanjokousen, Kumotoriyama, 3.vi. 1952, T. Okada (NSMT). Kanagawa Pref.: 1 ♀, Mizuhinoatama, Mt. Ooyama, 18.v. 1997, S. Tsuyuki (NSMT). Nara Pref.: 1 ♀, 4 ��, Mt. Odaigahara, 28-29.v. 1977, K. Mizuno (NSMT); 1 ♀, My��jindaira, Mt. Kunimiyama, 5.vi. 1988, K. Mizuno (NSMT); 3 ♀, 3 ��, Mt. Wasamata, Mts. Ohmine, 7.vi. 1981, K. Mizuno (NSMT). Kyoto Pref.: 1 ♀, Omi, Ohara, Sakyo-ku, Kyoto, 21.vi. 1984, T. Matsumoto (NSMT). SHIKOKU: Ehime Pref,: 1 ♀, Mt. Ishizuchi, 18.vi. 1978, N. Yashiro (NSMT); 3 ♀, 112 ��, Nanokawagoe, Tsuchigoya, Ishizuchiyama Mts., 33.75 ��N, 133.15 ��E, 1450m, 2- 4.vi. 2003, A. Shinohara (NSMT); 1 ♀, Nanokawagoe, Tsuchigoya, Ishizuchiyama Mts., 33.75 ��N, 133.15 ��E, 1450m, 2-4.vi. 2003, A. Shinohara, (CSCS); 14 ��,Nanokawagoe, Ishizuchiyama Mts., 33.75 ��N, 133.15 ��E, 1450m, 8-10.vi. 2005, A. Shinohara (NSMT); 1 ♀, Tsuchigoya, Omogo-mura, 12.vi. 1997, H. Urushihara (NSMT). Tokushima Pref.: 1 ♀, ��� Japan, AWA; Turugiyama, 1. vi. 1950, Issiki & Ito��� ��� Neocolochelyna itoi Tak., Paratype, det. Takeuchi��� (OPU) [this specimen was not mentioned in the original description]; 1 ��, ��� 30. v. 1950, Mt. Turugi, Awa, Takeuchi��� (OPU); 9 ♀, 20 ��,Minokoshi, Tsurugisan Mts., 33.52 ��N 134.05 ��E, 1500m, 5.vi. 2003, T. Kurihara (NSMT); 2 ♀, 3 ��, Minokoshi, Tsurugisan Mts., 33.52 ��N 134.05 ��E, 1500m, 5.vi. 2003, T. Kurihara (CSCS); 2 ♀, 18 ��, Minokoshi, Tsurugisan Mts., 33.52 ��N 134.05 ��E, 1500m, 5.vi. 2003, A. Shinohara (NSMT); 1 ��, Meotoike, Tsurugisan Mts., 33.87 ��N, 134.07 ��E, 1400m, 6-7.vi. 2003, A. Shinohara (NSMT); 2 ��, Kasutorisawa, Tsurugisan Mts., 33.88 ��N, 134.05 ��E, 1250m, 6.vi. 2003, T. Kurihara (NSMT). KYUSHU: Oita Pref.: 1 ♀, Chojabaru, Kuju, 14- 15.v. 1986, A. Shinohara (NSMT). Fukuoka Pref.: 1 ♀, ���[Kyushu], Hikosan, 15. v. 1955, K. Yasumatsu��� (OPU); 1 ♀, Hikosan, 23.v. 1950, N. Fukuhara (NSMT); 1 ♀, Hikosan, 13-15.V. 1964, T. Okada, (NSMT); 1 ♀, Mt. Hiko, 3.vi. 1971 (NSMT). Distribution. Japan (Fig. 12). Host plant. Actinidia arguta Plauch (Okutani 1956; Yasumatsu 1957; Togashi 1995; Shikata & Itakura 2009) Remarks. This species is endemic to Japan. It is similar to N. testaceoa. See above key for the differences between the species and its congeners. No distinct difference has been confirmed between N. hakusana and N. itoi while comparing the holotypes of the two species. The differences between the two species pointed by Togashi (1995) fall into the variation range of N. itoi. Maxwell (1958) studied the internal anatomy of the larva of this species. Togashi (1995) described the larva and reported some biological characters of this species under the name of N. hakusana. Takeuchi (1951) stated that there was one more paratype male with the same data as the holotype but it was not found in OPU when the first two authors visited the university in March, 2014., Published as part of Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko & Naito, Tikahiko, 2016, Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae), pp. 432-456 in Zootaxa 4127 (3) on pages 437-443, DOI: 10.11646/zootaxa.4127.3.2, http://zenodo.org/record/264340, {"references":["Takeuchi, K. (1951) New and unrecorded sawflies from Shikoku, Japan 1). (Hymenoptera: Symphyta). Transactions of the Shikoku Entomological Society, 2 (4), 57 - 62.","Takeuchi, K. (1955) Coloured Illustrations of the Insects of Japan, Volume II. Hoikusha, Osaka. 190 pp., 68 plates. [In Japanese]","Okutani, T. (1956) Some records on the food-plants for Japanese sawflies with a note on an egg-laying habit of Macrophya apicalis (Studies on Symphyta IV). The Science Reports of the Hyogo University of Agriculture, Agricultural Biology, 2 (2), 1 - 2.","Maxwell, D. E. (1958) Comparative internal larval anatomy. Results in the Tenthredinidae based upon six Japanese species. Mushi, 31 (2), 15 - 23.","Togashi, I. (1965) Tenthredinidae. In: Asahina, S., Ishihara, T. & Yasumatsu, K. (Eds.), Iconographia Insectorum Japonicorum Colore naturali edita, Volumen III, pp. 245 - 254, pls. 123 - 126. Hokuryukan, Tokyo. [In Japanese]","Togashi, I. (1995) Description of Neocolochelyna hakusana, sp. nov., and its larva (Hymenoptera: Tenthredinidae) from Japan. Proceedings of the Entomological Society of Washington, 97 (4), 866 - 871.","Yasumatsu, K. (1957) On the host-plant of Neocolochelyna itoi Takeuchi. Kontyu, 25 (3), 101. [In Japanese]","Okutani, T., Tsuruta, K. & Shinohara, A. (1992) Records of Neocolochelyna itoi (Hymenoptera, Tenthredinidae) in Honshu. Japanese Journal of Entomology, 60 (1), 238.","Naito, T., Yoshida, H., Nakamine, H., Morita, T., Ikeda, T., Suzuki, H. & Nakanishi, A. (2004) Species diversity of sawflies in Hyogo Prefecture, central Japan. Museum of Nature and Human Activities, Hyogo, Monograph of Natural History and Environmental Science, (1), [1 - 2] + [pl. 1 - 10] + 1 - 85. [In Japanese]","Shikata, K. & Itakura, T. (2009) First record of Neocolochelyna itoi in Nagano Prefecture. Natural History Reports of Inadani, 10, 75 - 77. [In Japanese]","Taeger, A., Blank, S. M. & Liston, A. D. (2010) World Catalog of Symphyta (Hymenoptera). Zootaxa, 2580, 1 - 1064."]}

Distribution of Colochela The distribution of Colochela is shown in Fig. 6. The genus is endemic to southeastern Asia at present. The distribution of the three known species overlaps in Sichuan, Chongqing, Shaanxi and Hubei provinces. Within the generic distribution, C. nigrata is distributed in the southeastern area, C. zhongi distributed in the north and C. rufidorsata distributed in the southwestern area. It is possible that the genus is originated and differentiated in the Qinba Mountains of China. Further research on the molecular biogeography of the genus is needed to test this., Published as part of Niu, Gengyun & Wei, Meicai, 2016, Revision of Colochela (Hymenoptera, Tenthredinidae), pp. 457-470 in Zootaxa 4127 (3) on page 469, DOI: 10.11646/zootaxa.4127.3.3, http://zenodo.org/record/259643

Key to species of Neocolochelyna Konow 1 Claw with inner tooth clearly shorter than outer tooth (Figs 7 d���i); occipital carina present in lower 0.3 or absent; vein 1 M as long as vein 1 m-cu; cell A of forewing strongly constricted at basal 0.25, where distance between veins 1 A and 2 A+ 3 A about 2���3 times as broad as 2 A+ 3 A at same position; apex of labrum broadly round (Figs 4 b, e, h; Figs 5 b, e, h). Subgenus Curvatapenis Wei........................................................................................... 2 - Claw with inner tooth clearly longer than outer tooth (Figs 7 a���c); occipital carina present in lower half of hind orbit; vein 1 M clearly shorter than vein 1 m-cu; cell A of forewing weakly constricted at basal 0.25, where distance between veins 1 A and 2 A+ 3 A about 4���5 times as broad as 2 A+ 3 A at same position; apex of labrum subtriangular (Figs 3 b, e, h). Subgenus Neocolochelyna Konow....................................................................................... 5 2 Mesoscutellum coarsely punctured without interspaces, mat (Figs 6 g���i); head coarsely punctured or densely rugose, mat, narrow smooth interspaces absent (Figs 5 a���i); hairs on dorsum of head and mesopleuron very dense, longer than diameter of median ocellus or distinctly curved at apex; at least basal 0.3 of antenna black; distance between antennal sockets as long as distance between antennal socket and inner margin of eye; abdominal tergites 3���7 largely black brown to black, pale posterior margin less than 0.3 times length of tergite, tergite 1 almost entirely black; body length about 13 mm in female and about 10��� 12 mm in male........................................................................................ 3 - Mesoscutellum and head densely punctured with narrow but distinct smooth and shiny interspaces, not mat (Figs 3���4); hairs on dorsum of head and mesopleuron straight, not very dense, shorter than diameter of median ocellus, apex not curved; basal 0.25 of antenna distinctly paler than apical 0.75; distance between antennal sockets clearly shorter than distance between antennal socket and inner margin of eye; abdominal tergites 3���7 narrowly black brown at base, pale posterior stripe not less than 0.6 times length of tergite, broad posterior margin of tergite 1 pale brown; body length: female about 16 mm, male 13���14 mm.. 4 3 Eyes small, distance between them at level of lateral corner of clypeus about 1.3���1.4 times as long as longest axis of eye (Figs 5 e, 5 h); antenna largely or entirely black, outer side of antennomeres 4���9 with shallow but distinct longitudinal furrow; abdominal tergite 1 with long and dense hairs, surface strongly and densely microsculptured mixed with small and indistinct punctures, mat (Figs 6 h, i); pale posterior margin of tergites 3���7 about 0.3 times length of tergite; wings almost entirely hyaline; middle fovea indistinct; mesepisternum strongly elevated at middle with a blunt but distinct ridge; ventral half of mesepisternum strongly microsculptured, mat; male: dorsum of abdominal tergite 2 with a large middle black spot (Fig. 6 i); male: dorsum of abdominal tergite 2 with a large middle black spot (Fig. 6 i); female: dorsum of abdominal tergite 2 yellow brown without large black spot (Fig. 6 h), pronotum largely yellow white. China: Hubei, Sichuan.................................................................................................... N. (C.) hyalina Niu & Wei, sp. nov. - Eyes medium sized, distance between them at level of lateral corner of clypeus about 1.1 times as long as longest axis of eye (Fig. 5 b); apical 0.75 of antenna entirely yellow brown, outer side of antennomeres 4���9 without longitudinal furrow; abdominal tergite 1 with short and sparse hairs, middle area and lateral with some large and deep punctures mixed with microsculpture, surface weakly shiny, not mat (Fig. 6 g); pale posterior margin of tergites 3���7 less than 0.15 times length of tergite; anterior half of forewing strongly infuscate; middle fovea distinct; mesepisternum hardly elevated at middle, without distinct ridge; ventral half of mesepisternum hardly microsculptured, distinctly shiny; male tergite 2 entirely yellow brown, without black macula dorsally and ventrally (Fig. 6 g); female unknown. China: Gansu............ N. (C.) wui Niu & Wei, sp. nov. 4 Antennomere 3 1.2 times as long as apical 4 antennomeres together (in male, female unknown); postocellar area flat, lateral furrows of postocellar area shallow and obscure; POL: OOL: OCL= 15: 40: 35 (Fig. 4 a); dorsal hook of penis valve almost straight, middle tubercle of valviceps short and distinctly convex (Fig. 10 c). China: Shaanxi, Henan.................................................................................................. N. (C.) testaceoa (Wei, 2002) - Antennomere 3 0.9 times (in male) or 0.8 times (in female) as long as apical 4 antennomeres together; postocellar area distinctly elevated, lateral furrows of postocellar area deep and broad; POL: OOL: OCL= 15: 25: 26 (female) or 15: 29: 30 (male) (Figs 4 d, 4g); dorsal hook of penis valve distinctly curved, middle tubercle of valviceps long and weakly convex (Fig. 10 e). Japan.......................................................................... N. (C.) itoi Takeuchi, 1951 5 Clypeus with a blunt middle ridge, anterior margin with a distinct middle tooth (Figs 3 e, 3 h); mesoscutellum about 4 times as long as median length of mesoscutellar appendage, anterior corner obtusely truncate, weakly protruding (Figs 6 b���c); apex of labrum acute (Figs 3 e, 3 h); antennomere 3 about 1.1 times as long as antennomeres 6���9 together; eyes small, distance between eyes at level of anterior tentorial pits 1.5 times as long as longest axis of eye in female (Fig. 3 e); posterior half of frons and upper inner orbit in female yellow brown (Fig. 3 d); mesepisternum in both sexes black with a distinct yellow brown macula near upper corner; lateral furrows of postocellar area distinctly curved, subparallel in posterior 0.6 (Figs 3 d, 3g); posterior margin of postocellar area elevated; apex of inner tibial spur of fore leg very shallowly incised (Figs 7 k���l); triangular middle lobe of sternite 7 broad and large, much broader than lateral deeply concaved posterior margin of sternite 7 (Fig. 8 l); lancet as Fig. 8 d; apical margin of subgenital plate round. China: eastern Tibet.................. N. (N.) acutilabria Niu & Wei, sp. nov. - Clypeus flat, anterior margin without distinct middle tooth (Fig. 3 b); mesoscutellum about 6 times as long as median length of mesoscutellar appendage, anterior corner roundly and strongly protruding (Figs 6 a); apex of labrum narrowly obtuse (Fig. 3 b); antennomere 3 about 0.9 times as long as antennomeres 6���9 together; eyes large, distance between eyes at level of anterior tentorial pits 1.2 times as long as longest axis of eye in female(Fig. 3 b); frons and inner orbit in female entirely black (Fig. 3 a); mesepisternum in female black, without yellow brown macula near upper corner; lateral furrows of postocellar area straight, strongly divergent backwards in entire length (Fig. 3 a); posterior margin of postocellar area flat; apex of inner tibial spur of fore leg deeply incised (Fig. 7 j); triangular middle lobe of sternite 7 small, much narrower than lateral straight posterior margin of sternite 7 (Fig. 8 k); lancet as Fig. 8 c; apical margin of subgenital plate subtruncate. India: Sikkim............................................................................................... N. (N.) montana (Konow, 1898), Published as part of Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko & Naito, Tikahiko, 2016, Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae), pp. 432-456 in Zootaxa 4127 (3) on pages 434-435, DOI: 10.11646/zootaxa.4127.3.2, http://zenodo.org/record/264340, {"references":["Wei, M. (2002) A new genus and a new species of Tenthredininae (Hymenoptera: Tenthredinidae). In: Shen, X. & Zhao, Y. (Eds.), Insects of the Mountains Taihang and Tongbai Regions. (The Fauna and Taxonomy of Insects in Henan, Vol. 5 [2003]), China Agricultural Science and Technology Press, pp. 253 - 256. [In Chinese, abstract in English]","Takeuchi, K. (1951) New and unrecorded sawflies from Shikoku, Japan 1). (Hymenoptera: Symphyta). Transactions of the Shikoku Entomological Society, 2 (4), 57 - 62.","Konow, F. W. (1898) Neue Chalastogastra-Gattungen und Arten. Entomologische Nachrichten, 24 (17 - 18), 268 - 282."]}

Niu, Gengyun, Wei, Meicai (2016): Revision of Colochela (Hymenoptera, Tenthredinidae). Zootaxa 4127 (3): 457-470, DOI: http://doi.org/10.11646/zootaxa.4127.3.3

Neocolochelyna (Curvatapenis) testaceoa (Wei, 2002) comb. nov. (Figs 1 e, 4 a���c, 6 d, 7 d, 7m, 10 c��� d, 11 d, 11m ���n) Curvatapenis testaceous Wei, 2002: 254; Wei et al., 2006: 523; Taeger et al., 2010: 549. Description. Male . Holotype (Fig. 1 e). Length 13 mm. Head brown (Figs 4 a���b), palpi of mouthparts, labrum and malar space yellowish brown, narrow posterior margin of clypeus, supraclypeal area, frons, lower 0.7 of inner orbit, ocellar area, anterior margin and middle stripe of postocellar area and posterior side of head largely black; basal two antennomeres entirely, basal 0.25 of antennomere 3 yellow brown, apical 0.75 of antennomere 3 and other antennomeres dark brown (Fig. 11 d). Thorax black, broad posterior margin of pronotum, tegula, lateral stripe on mesoscutal middle lobe, broad anterior margin and narrow posterior margin of parapsis, mesoscutellum (excluding appendage), metascutellum, cenchrus and upper margin of metapleuron pale brown. Abdomen brown to pale brown, tergite 2 and subgenital plate largely and harpes entirely yellow brown, basal 0.8 (middle) to 0.6 (lateral sides) of tergite 1, short lateral macula and narrow anterior margin of tergite 2 (Fig. 6 d), basal 0.3 of tergites 3���7 blackish brown, base of basal sternites more or less blackish brown. Body hairs brown to pale brown. Legs black, hind trochanter, broad anterior stripe and broad apex of fore and middle femora, fore and middle tibiae entirely pale brown, hind tibia brown to dark brown, all tarsi yellow brown. Wings yellowish infuscate in anterior part and gradually paler toward posterior margin, vein C and pterostigma pale brown, posterior of vein Sc+R and most of vein M+Cu blackish; hind wing weakly infuscate, slightly paler than forewing. Punctures on labrum small and dense, narrow interspaces between punctures weakly microsculptured; punctures on clypeus dense and large, interspaces between punctures almost smooth, shiny (Fig. 4 b); inner orbit and hind orbit densely punctured, interspaces linear, weakly microsculptured, less shiny; frons densely and coarsely punctured, interspaces narrow, hardly microsculptured, more or less shiny; punctures on temple and postocellar area slightly sparser than frons, surface very weakly microsculptured, moderately shiny (Fig. 4 a���c); malar space densely punctured, shiny; upper corner of pronotum moderately punctured and shiny, depressed part of pronotum coarsely punctured and microsculptured, feebly shiny; mesoscutal middle lobe and lateral lobe densely punctured, interspaces very narrow, distinctly microsculptured, almost dim; mesoscutellum densely punctured with narrow smooth interspaces, less shiny; mesoscutellar appendage polished, impunctate, strongly shiny; parapsis weakly microsculptured, less shiny; metapostnotum densely microsculptured (Fig. 6 d); punctures on upper half of mesepisternum large and very dense, interspaces very narrow, hardly shiny; punctures on lower half of mesepisternum smaller and much sparse, surface indistinctly microsculptured, shiny; mesepimeron and metapleuron strongly microsculptured mixed with some large but shallow punctures; abdominal tergite 1 densely microsculptured, almost mat, narrow lateral side densely punctured, very narrow posterior margin weakly coriaceous, less shiny; tergite 2 hardly punctured, densely microsculptured, other tergites strongly microsculptured mixed with dense but indistinct punctures in entire length, mat (Fig. 6 d); microsculpture on median part of tergite 8���9 hardly weaker than lateral part; sternites densely punctured except for apical half of subgenital plate. Hairs on dorsum of head and mesopleuron dense and almost straight, about 0.9 times as long as diameter of median ocellus (Fig. 4 a���b); hairs on mesonotum dense and about 0.6 times as long as diameter of median ocellus; dorsum of abdominal tergites 1���2 largely glabrous except for narrow middle stripe, other tergites densely pubescent; mesoscutellar appendage and intertergites of abdomen glabrous (Fig. 6 d). Labrum broad, anterior margin round, not acute; anterior margin of clypeus weakly incised in entire length, almost truncate; malar space about as long as radius of median ocellus (Fig. 4 b); hind orbit round, without occipital carina; distance between eyes at level of anterior tentorial pits 1.2 times as long as longest axis of eye (Fig. 4 b); mandibles as Figs 11 m���n; middle fovea roundish, shallow, closed; lateral fovea small; frons small, roundish, frontal ridge obtuse; POL: OOL: OCL= 15: 40: 35; interocellar furrow narrow and very deep, postocellar furrow fine and shallow, hardly bent forward; postocellar area flat, hardly elevated, about 1.3 times as broad as long; lateral furrow broad but very shallow, weakly curved, weakly divergent backwards; head hardly dilated behind eyes in dorsal view (Fig. 4 a). Antenna as long as thorax, antennomere 3 almost 1.3 times as long as antennomeres 4 and 5 together and 1.2 times as long as antennomeres 6���9 together, antennomere 7 about 0.9 times as long as broad (Fig. 11 d). Mesoscutal middle lobe with narrow but distinct middle furrow; mesoscutellum roundly elevated, anterior margin weakly and roundly protruding, transverse anterior furrow deep; mesoscutellar appendage short with middle length about as long as diameter of median ocellus; cenchrus transverse, much broader than long, CD= 1.7 (Fig. 6 d); mesepisternum roundly and weakly elevated at middle, without blunt ridge. Venation as Fig. 1 e, cell 2 Rs about 0.9 times as long as cells 1 R 1 + 1 Rs, cell A of forewing strongly constricted at basal 0.25, where distance between 1 A and 2 A+ 3 A about 3 times as broad as vein 2 A+ 3 A, first abscissa of vein M 0.9 times as long as 1 m-cu, vein 2 r 1 meeting cell 2 Rs at apical 0.45. Inner tibial spur of fore leg as Fig. 7 m; claw as Fig. 7 d, inner tooth clearly shorter than apical tooth. Subgenital plate about as long as broad, apical margin weakly and roundly protruding. Harpes, volsella and gonolacinia as Fig. 10 d; penis valve as Fig. 10 c, dorsal apical hook long and almost straight, strongly tapering toward apex, middle tubercle of valviceps short and distinctly convex. Female. Unknown. Material examined. Types. Holotype: ��, CHINA: HENAN: ���Baiyun Mountain, Songxian, Henan, 2001 -VI- 1, 1500 m, Zhong Yihai leg.���; ���CSCS-Holotype 00813001���[red]; ��� Curvatapenis testaceous Wei, 2002, det. M. Wei, 2002 ���; ��� Neocolochelyna testaceoa (Wei, 2002) ���. (CSCS). Paratype: 1 ��, CHINA: HENAN, ���Dakuaidi, Lushi, Henan, 2000 -V- 29, 1400 [m], Zhong Yihai leg.���; ��� Paratype ��� [yellow] (CSCS). Other specimen. 1 ��, CHINA: SHAANXI: Kaitianguan, Mt. Taibaishan, Qinling Mts., 34 ��00'N, 107 �� 51 'E, 2000m, 1. VI. 2005, A. Shinohara. (NSMT). Distribution. China (Shaanxi, Henan). The species is recorded from Shaanxi for the first time (Fig. 12). Host plant. Unknown. Remarks. As Curvatapenis merged with Neocolochelyna, Curvatapenis testaceous is transferred into Neocolochelyna. N. testaceoa is similar to N. itoi Takeuchi from Japan. See the above key for the differences between the two species., Published as part of Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko & Naito, Tikahiko, 2016, Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae), pp. 432-456 in Zootaxa 4127 (3) on pages 443-444, DOI: 10.11646/zootaxa.4127.3.2, http://zenodo.org/record/264340, {"references":["Wei, M. (2002) A new genus and a new species of Tenthredininae (Hymenoptera: Tenthredinidae). In: Shen, X. & Zhao, Y. (Eds.), Insects of the Mountains Taihang and Tongbai Regions. (The Fauna and Taxonomy of Insects in Henan, Vol. 5 [2003]), China Agricultural Science and Technology Press, pp. 253 - 256. [In Chinese, abstract in English]","Wei, M., Nie, H. & Taeger, A. (2006) Sawflies (Hymenoptera: Symphyta) of China - Checklist and review of research. In: Blank, S. M., Schmidt, S. & Taeger, A. (Eds.), Recent Sawfly Research - Synthesis and Prospects. Goecke & Evers, Keltern, pp. 505 - 574.","Taeger, A., Blank, S. M. & Liston, A. D. (2010) World Catalog of Symphyta (Hymenoptera). Zootaxa, 2580, 1 - 1064."]}

Neocolochelyna (Neocolochelyna) acutilabria Niu & Wei, sp. nov. ( Figs 1 c–d, 3 d–i, 6 b–c, 7 b–c, 7 k–l, 8 d, 8 h, 8 l, 9 d, 9 h, 10 a–b, 11 b–c, 11 k–l ) Description. Female. Holotype (Fig. 1 c). Length 15 mm. Head brown to orange, labrum yellow brown, narrow posterior margin of clypeus, supraclypeal area, middle fovea and lateral foveae, lower 0.6 of inner orbit and postocellar furrow black (Figs 3 d–e), lower part of posterior side of head blackish; antenna entirely brown, basal part hardly paler than apex (Fig. 11 b). Prothorax black, narrow lateral margin and very broad posterior margin of pronotum brown; mesoscutal middle lobe and lateral lobes dark brown, centre of middle lobe and dorsal broad stripe on lateral lobe black brown to black, mesoscutellum pale brown, parapsis largely and mesoscutellar appendage black; metanotum pale brown, a dot lateral to metascutellum and most of metapostnotum blackish, cenchrus yellow (Fig. 6 b); mesopleuron and metapleuron black, a large elliptical macula near dorsal corner of mesepisternum, narrow posterior margin of mesepimeron, dorsal margin of metapleuron brown. About basal half of abdominal tergites 1–8 black, a narrow stripe covering spiracle and about apical half of tergites 1–8, tergites 9– 10 entirely, cerci and ovipositor sheath pale brown, about basal 0.7 of each sternite black brown. Body hairs brown to pale brown. Legs orange, ventral side of fore and middle coxae, hind coxa except for lateral stripe, black, hind tarsus yellow brown. Wings yellowish infuscate in anterior part and gradually paler toward posterior margin, vein C and pterostigma pale brown, posterior of vein Sc+R and most of vein M+Cu blackish; hind wing weakly infuscate, slightly paler than forewing. Punctures on labrum small and sparse, broad interspaces between punctures smooth, shiny; punctures on clypeus dense and large, narrow interspaces between punctures smooth, shiny; frons and inner orbit coarsely punctured, rugose, mat (Fig. 3 e); hind orbit densely punctured, interspaces linear, weakly microsculptured, less shiny; punctures on temple and postocellar area sparser than inner orbit, interspaces smaller than punctures, surface clearly but weakly microsculptured, less shiny (Fig. 3 f); malar space densely punctured; upper corner of pronotum densely punctured and almost dim, depressed part of pronotum coarsely punctured and microsculptured, almost mat, narrow lateral margin of pronotum shiny; mesoscutal middle lobe and lateral lobe densely punctured, interspaces very narrow, distinctly microsculptured, punctures on posterior part of lateral lobe smaller and much denser, almost dim; punctures on mesoscutellum large and dense, interspaces microsculptured, less shiny; mesoscutellar appendage weakly but clearly coriaceous, impunctate; parapsis densely microsculptured, metapostnotum strongly microsculptured (Fig. 6 b); punctures on upper half of mesepisternum large and very dense, interspaces very narrow, hardly shiny; punctures on lower half of mesepisternum small and much sparse, surface distinctly microsculptured, weakly shiny; mesepimeron and metapleuron strongly microsculptured mixed with some shallow and indistinct punctures, narrow anterior and posterior margins of mesepimeron shiny; abdominal tergite 1 strongly microsculptured, almost mat, center and lateral side with some shallow and large punctures, very narrow posterior margin weakly coriaceous, less shiny (Fig. 6 b); tergites 2–8 densely microsculptured with a few shallow and indistinct punctures laterally, mat; median part of tergite 8–10 weakly microsculptured, distinctly shiny; surface of basal sheath microsculptured mixed with some shallow punctures, apical sheath hardly coriaceous, shiny. Hairs on dorsum of head and mesopleuron dense, weakly curved at apex, about as long as diameter of median ocellus (Figs 3 d–e); hairs on mesoscutal middle and lateral lobes about 0.8 times as long as diameter of median ocellus; dorsum of abdominal tergites 1–8 distinctly pubescent, hairs on tergite 1 not longer and denser than other tergites; mesoscutellar appendage and intertergites of abdomen glabrous (Fig. 6 b). Labrum triangular, apically acute; clypeus with a low middle ridge, anterior margin almost truncate, with a very short but distinct middle tooth; malar space as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.5 times as long as longest axis of eye (Fig. 3 e); mandibles as Figs 11 k–l; hind orbit with a distinct occipital carina in lower half; middle fovea roundish, shallow, narrowly open to frons; lateral fovea indistinct, with a small tubercle; frontal ridge broad and round; POL: OOL: OCL= 15: 35: 31; interocellar furrow short and very deep, postocellar furrow narrow and distinct, triangularly bent forward at middle; postocellar area almost flat, 1.35 times as broad as long, posterior margin elevated; lateral furrows very shallow and indistinct in anterior 0.7, replaced by a low carina in posterior 0.3, hardly curved, distinctly divergent backwards; head roundly dilated behind eyes in dorsal view (Fig. 3 d). Antenna as long as thorax, antennomere 3 about 1.2 times as long as antennomeres 4 and 5 together and 1.1 times as long as antennomeres 6–9 together, antennomere 7 about as long as broad (Fig. 11 b). Mesoscutal middle lobe with narrow but distinct middle furrow; mesoscutellum roundly elevated, anterior margin almost truncate, hardly protruding, transverse anterior furrow deep; mesoscutellar appendage short with middle length about 1.1 times as long as diameter of median ocellus; mesoscutellum 4 times as long as mesoscutellar appendage; cenchrus transverse, much broader than long, CD= 1.3 (Fig. 6 b); mesepisternum roundly and weakly elevated at middle, without distinct ridge. Venation as Fig. 1 c, cell 2 Rs about 0.75 times as long as cells 1 R 1 + 1 Rs, cell A of forewing weakly constricted at basal 0.25, where distance between 1 A and 2 A+ 3 A 4 times as broad as vein 2 A+ 3 A; vein 1 M as long as vein 1 m-cu; vein 2 r 1 meeting cell 2 Rs at apical 0.4. Inner tibial spur of fore leg as Fig. 7 k, apical incision very shallow; claw as Fig. 7 b, inner tooth clearly longer than apical tooth. Triangular middle lobe of sternite 7 large and broad, much broader than lateral incision of sternite 7 (Fig. 8 l). Ovipositor sheath 0.9 times as long as front tibia and 0.7 times as long as middle tibia, apical sheath 1.1 times as long as basal sheath, narrowly truncate at apex in lateral view. Lancet (paratype ♀) as Fig. 8 d, with 25 serrulae, middle and apical segments as Figs 9 d, 9 h; annular spines and surface sculptures of lancet as Fig. 9 h; basal sensilla as Fig. 8 h. Male. Paratype (Fig. 1 d). Body length 13.5 mm; color, punctation and structure similar to female except for following: posterior 0.7 of clypeus, frons, upper inner orbits largely, postocellar area, lower half of hind orbit, mesoscutal middle and lateral lobes black; malar space 0.8 times as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.3 times as long as longest axis of eye (Fig. 3 h); dorsum of head and mesonotum coarsely rugose, punctures irregular, not clearly defined (Fig. 3 g, 3 i); subgenital plate 1.1 times as broad as long, apical margin round; harpes, volsella and gonolacinia as Fig. 10 b; penis valve as Fig. 10 a, dorsal apical hook very long and strongly tapering toward apex. Variation. In a paratype female, the CD is about 1.5 and the vein 2 r 1 meets cell 2 Rs at middle. Distribution. China (Xizang) (Fig. 12). Etymology. This species is named after the triangular labrum which is acute at apex. The name is to be treated as a noun. Material examined. Holotype: ♀, CHINA: XIZANG: Yigong, 2300 m, Sichuan Prov. [Xizang], 28. iv. 1986, T. Naito leg.; 86. 4. 28, 2, Xizang, Yigong; (CSCS, lancet dissected and missing, possibly kept in KUK). Paratypes: 1 ♀, data same as the holotype (KUK); 1 Ƌ, data same as the holotype (KUK). Host plant. Unknown. Remarks. Neocolochelyna acutilabria is similar to N. montana Konow. See the above key for the differences between the two species. Two females and one male of Neocolochelyna acutilabria are known. The female specimen with lancet missing was selected as holotype because the specimen is in good state, the second female and the male were partly broken.

Colochela zhongi Wei & Niu, sp. nov. (Figs 1 f���k, 2 c, f, i, l, o, 3 a���d, g, j, 4 c, f, i, m, n, 5 c, h, i, p���r) urn:lsid:zoobank.org:act:CA 215 D 40 -04B 3-4655 - 8854 -DB 2724 B 86 B 34 Description. Holotype: ♀. Length 17.5 mm (Fig. 1 f). Body deep black; mesoscutal middle lobe entirely, lateral lobe except for narrow lateral margin, mesoscutellum except for narrow posterior margin, red; body hairs black brown, hairs on red part of mesonotum golden. Wings strongly and evenly infuscate, vein R 1 largely brown, pterostigma and other veins black; hind wing slightly paler than forewing. Clypeus densely punctured, interspaces between punctures narrow and smooth, shiny; inner orbit, temple and postocellar area with fine surface microsculpture, weakly shiny; frons densely microsculptured with some minute punctures; malar space densely microsculptured, hind orbit with dense and minute punctures; mesoscutal middle lobe and lateral lobe indistinctly punctured, mesoscutellum minutely punctured in anterior 0.75 and densely punctured in posterior margin; mesoscutellar appendage (posttergite) smooth and glabrous, strongly shiny; punctures minute and rather dense on upper corner of mesepisternum, and gradually larger and sparser toward ventral side, surface indistinctly microsculptured, shiny; depressed area of mesepimeron and of metapleuron densely microsculptured, broad posterior margin of mesepimeron and of metepimeron smooth and strongly shiny; abdominal tergites 1���2 sparsely and minutely punctured, interspaces microsculptured, shiny; tergites 3���10 densely and minutely punctured, interspaces microsculptured, weakly shiny; intertergites smooth, strongly shiny. Hairs on dorsum of head and mesopleuron dense and slightly shorter than diameter of median ocellus; hairs on mesonotum extreme dense and about 0.7 times as long as diameter of median ocellus; abdominal tergites densely pubescent; intertergites glabrous. Anterior margin of clypeus almost truncate; malar space 0.5 times as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.2 times as long as longest axis of eye (Fig. 2 f); labium as Fig. 3 b, maxilla as Fig. 3 c; middle fovea round, very deep; frontal ridge indistinct; POL: OOL: OCL = 11: 20: 18; interocellar furrow deep and broad, postocellar furrow shallow, hardly bent forward; postocellar area roundly elevated, about 1.8 times as broad as long; lateral furrow deep, hardly curved, distinctly divergent backward; head hardly dilated behind eyes in dorsal view (Fig. 2 c). Antenna as long as head and thorax together, not incrassated beyond middle, 5 th ��� 9 th antennomeres with shallow but distinct longitudinal furrow in both sides, third antennomere about 0.9 times as long as following two antennomeres together, 7 th antennomere about 2.3 times as long as broad (Fig. 3 g). Mesoscutellum strongly and triangularly protruding; mesoscutellar appendage flat, without middle carina, posterior half with a short and shallow middle furrow; cenchrus transverse and elliptical, much broader than long, CD = 1.9. Venation as Fig. 1 f, anal cross vein present; petiole of hind anal cell about 0.4 times length of cu-a. Claw as Fig. 3 d. Ovipositor sheath 1.1 times as long as front tibia, apical sheath as long as basal sheath, distinctly tapering toward apex, apex narrowly round (Fig. 2 o); sheath and setae in dorsal view as Fig. 2 l, sheath distinctly broadened toward apex, about 4 times as broad as breadth of cercus; cercus slender, about 4 times as long as broad. Lancet as Fig. 4 c, with 17 serrulae, middle and apical segments of lancet as Figs 4 i, f, annular spines long and dense; surface sculpture of lancet as Figs 4 m, n. Male. Body length 15 mm (Fig. 1 g); color, punctation and structure similar to female except for following: malar space 0.3 times as long as diameter of median ocellus (Fig. 2 i); distance between eyes at level of anterior tentorial pits as long as longest axis of eye; head distinctly narrowed behind eyes in dorsal view; subgenital plate broader than long, apical margin broadly truncate; harpes 1.9 times as long as broad (Fig. 5 c); volsella and gonolacinia as Figs 5 h, i; penis valve as Figs 5 p���r. Variation. The anal cross vein in forewing is sometimes absent in a few specimens. Etymology. The species is named after Dr. Yihai Zhong, an early collector of some type specimens of the species. Specimens examined. Holotype. ♀, CHINA: Shaanxi: Kaitianguan, Mt. Taibai, Meixian, 34.00��N, 107.8579 ��E, 1852m, 7.vi. 2014, Wei Meicai, (CSCS). Paratypes. CHINA: Hubei: Dalongtan, Shennongjia, 31.4949 ��N, 110.2962 ��E, 2180m, 1 ♂, 9.vii. 2011, Li Zejian, Liu Yao (CSCS); Shaanxi: Kaitianguan, Mt. Taibai, Meixian, 34.00��N, 107.8579 ��E, 1852m, 1 ♂, 5.vi. 2014, Qi Liwei, Kang Weinan, CH 3 COOC 2 H 5 (CSCS), 1 ♀, 7.vi. 2014, Wei Meicai, KCN (CSCS), 1 ♀, 9.vi. 2014, Wei Meicai, KCN (CSCS); Kaitianguan, Mt. Taibaishan, Qinling Mts, 34.00��N, 107.85 ��E, 2000m, 1 ♀, 2.vi. 2004, A. Shinohara (NMNS), 1 ♂, 27.v. 2005, A. Shinohara (NMNS), 1 ♂, 5.vi. 2004, A. Shinohara (NMNS), 1 ♂, 8.vi. 2006, A. Shinohara (NMNS); Liangfengyading, Mt. Taibai, Meixian, 33.6853 ��N, 107.8541 ��E, 2128m, 1 ♀, 17.vi. 2014, Qi Liwei, Kang Weinan, CH 3 COOC 2 H 5 (CSCS), 1 ♀, 17.vi. 2014, Liu Mengmeng, Liu Ting, CH 3 COOC 2 H 5 (CSCS); Zhuiwoping, Mt. Taibai, Meixian, 34.03 ��N, 107.8672 ��E, 1601m, 4 ♀, 4.vi. 2014, Qi Liwei, Kang Weinan, CH 3 COOC 2 H 5 (CSCS), 6 ♀, 6.vi. 2014, Qi Liwei, Kang Weinan, CH 3 COOC 2 H 5 (CSCS), 2 ♀, 6.vi. 2014, Liu Mengmeng, Liu Ting, CH 3 COOC 2 H 5 (CSCS), 2 ♀, 6.vi. 2014, Wei Meicai, KCN (CSCS), 1 ♂, 6.vi. 2014, Wei Meicai, KCN (CSCS); Sichuan: Leidongping, Mt. Emei, 29.546 ��N, 103.327 ��E, 2350m, 1 ♀, 2 ♂, 3.vii. 2009, Zhong Yihai, KCN (CSCS), Yajiageng, Luding, 29.8812 ��N, 102.0199 ��E, 3646m, 1 ♀, 4.vii. 2009, Wei Meicai, KCN (CSCS); Emei Shan: Jieyindian, 29.537 ��N + 103.331 ��E., 2400���2500m, 1 ♀, 0 6.07. 2009, leg. S.M. Blank, A.D. Liston & A. Taeger, DEI-GISHym 17846 (SDEI), a specimen with barcode (BOLD: ABV 5340, Genbank accession number KC 974810); 1 ♂ same data, without barcode (SDEI); 1 ♀, paratype of Colochela rufidorsata Malaise: ���Szechuen, CHINA, DC Graham���; ��� 30 Mi N. of Tatsienlu���; ��� Paratypus ���; ��� Colochela rufidorsata n gen. n. sp., det. Malaise, 1936 ���; ��� paratype ���; ��� Colochela zhongi Wei, sp. nov., det. M. Wei, 2013 ���(USNM). Distribution. China (Shaanxi, Sichuan, Hubei) (Fig. 6). Remarks. The new species is very close to C. rufidorsata, but the female and male genitalia are much different. See the above key for detailed differences between the two species., Published as part of Niu, Gengyun & Wei, Meicai, 2016, Revision of Colochela (Hymenoptera, Tenthredinidae), pp. 457-470 in Zootaxa 4127 (3) on pages 465-469, DOI: 10.11646/zootaxa.4127.3.3, http://zenodo.org/record/259643

Colochela rufidorsata (Malaise, 1937) (Figs 1 c���e, 2 b, e, h, k, n, 3 f, i, 4 b, e, h, l, 5 b, f, g, m���o) Colochela rufidorsata: Malaise, 1937: 48. (Type locality: Moupin [48 km N. Kangding]. 1m #, 2 f #, holotype in Mus. Paris, allotype and paratype in Mus. Washington.) Colochela rufidorsata: Malaise, 1945: 103; Singh & Saini, 1990: 271; Neocolochelyna rufidorsata (Malaise, 1937): Lacourt, 1997: 389. Description. Non-type: ♀. Length 17mm (Figs 1 c, d). Body black; mesoscutal middle lobe entirely, lateral lobe except for narrow lateral margin, mesoscutellum except for narrow posterior margin, red; body hairs black brown, hairs on red part of mesonotum golden. Wings strongly and evenly infuscate, vein R 1 largely brown, pterostigma and other veins black; hind wing slightly paler than forewing. Punctures on clypeus dense and large, interspaces between punctures smooth, shiny; inner orbit, frons, temple and postocellar area with very fine surface microsculpture, moderately shiny; malar space densely microsculptured, hind orbit with sparse and minute punctures, shiny; mesoscutal middle lobe and lateral lobe indistinctly punctured, mesoscutellum minutely and sparsely punctured in anterior 0.8 and densely punctured in posterior margin; mesoscutellar appendage (posttergite) smooth, strongly shiny; punctures on upper half of mesepisternum extremely minute, indistinct, narrow dorsal margin almost impunctate; punctures on lower half of mesepisternum large and shallow, sparse, surface indistinctly microsculptured, shiny; anterior half of mesepimeron, depressed area and posterior corner of metapleuron smooth and shiny; broad posterior margin of mesepimeron and other part of metepimeron sparsely punctured; abdominal tergite 1 uniformly and minutely punctured, interspaces microsculptured, shiny; tergite 9 smooth with few punctures, tergites 2���8 and 10 moderately punctured near posterior margin and sparsely punctured on other part, shiny; intertergites smooth, strongly shiny. Hairs on dorsum of head and mesopleuron dense and about 0.7 times as long as diameter of median ocellus; hairs on mesonotum extreme dense and about 0.5 times as long as diameter of median ocellus; abdominal tergites densely pubescent; mesoscutellar appendage and intertergites of abdomen glabrous. Anterior margin of clypeus almost truncate; malar space 0.4 times as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.1 times as long as longest axis of eye (Fig. 2 e); middle fovea round, very deep, closed; frons small and flat, frontal ridge indistinct; POL: OOL: OCL = 13: 26: 28; interocellar furrow deep and broad, postocellar furrow broad and shallow, hardly bent forward; postocellar area roundly elevated, about 1.7 times as broad as long; lateral furrow deep, hardly curved, weakly divergent backwards; head slightly and roundly dilated behind eyes in dorsal view (Fig. 2 b). Antenna about 0.8 times as long as head and thorax together, distinctly incrassated beyond middle, 5 th ��� 9 th antennomeres with indistinct longitudinal furrow in both sides, third antennomere about 1.2 times as long as following two antennomeres together, seventh antennomere about 1.3 times as long as broad (Fig. 3 f). Mesoscutellar appendage flat, without middle carina; cenchrus subtriangular, much broader than long, CD = 2.0. Venation as Fig. 1 c, anal cross vein absent; petiole of hind anal cell about 0.4 times length of cu-a. Ovipositor sheath 1.2 times as long as front tibia, apical sheath as long as basal sheath, roundly tapering toward apex in lateral view (Fig. 2 n); sheath and setae in dorsal view as Fig. 2 k, sheath distinctly broadened toward apex. about 3.5 times as broad as breadth of cercus; cercus slender, about 4 times as long as broad. Lancet as Fig. 4 b, with 15 serrulae, middle and apical segments as Figs 4 h, 4 e, annular spines absent; surface sculpture of lancet as Fig. 4 l. Male. Body length 16.5 mm (Fig. 1 e); color, punctation and structure similar to female except for following: malar space 0.5 times as long as diameter of median ocellus (Fig. 2 h); distance between eyes at level of anterior tentorial pits 1.2 times as long as longest axis of eye; head distinctly narrowed behind eyes in dorsal view; subgenital plate broader than long, apical margin roundish with a very shallow and narrow middle incision; harpes about 2 times as long as broad (Fig. 5 b); volsella and gonolacinia as Figs 5 f���g; penis valve as Figs m���o. Variation. The basal anal cross vein is present in the right wing of a specimen collected from Hubei, China. The petiole of hind anal cell is about 0.2 times length of cu-a in one femalehttp://www.morphbank.net/?id= 850767. Types. Holotype: ♀: ���MUSEUM PARIS, THIBET Mou-Pin, A. David, 1689 - 70 ���; ��� Typus ���; ��� Neocolochelyna sp. n., det. Malaise, 1935 ���; ��� Colochela rufidorsata Malaise, 1937 ���; ��� MNHN, Holotype, Colochela rufidorsata Malaise, 1937 ��� (MNHN). Paratypes see under C. zhongi. Other material examined. CHINA: Hubei: Fenshuiling, Hefeng, 1100m, 1 ♀, 29.vii. 1989, Wang Shuyong (IZAS), Pingbaying, Xianfeng, 1500m, 1 ♀, 21.vii. 1999, Deng Tiejun (CSCS); No data, 1 ♀ (CSCS), Shayuan, Hefeng, 1400m, 1 ♀, 31.vii. 1989, Wang Shuyong (IZAS); Yunnan: Hetouzhai, Jinping, 1700m, 1 ♂, 16.v. 1956, Huang Keren (IZAS), Mengzhe, Xishuangbanna, 1750m, 1 ♂, 23.vi. 1958, Pu Fuji (IZAS), Yongsheng, Liude, 2300m, 1 ♀, 9.vii. 1984, Wang Shuyong (IZAS). INDIA: Meghalaya: Shillong, 1600m, 1 ♀, 20.v. 1986, M. S. Saini (USNM). NEPAL: Gandaki: Marsyandi Valley, Bagarchap, Latamarang, 2100-2400m, 1 ♂, 22.vi. 1988 (SEHU) [0000051161 Sys. Ent. Hokkaido Univ. Japan]. THAILAND: Maeo Khun: Klang, Doi Inthanon, 1300m, 1 ♀, 21.X. 1983, M. Tomokuni (NMNS). Distribution. China (Sichuan, Hubei, Yunnan); India (Meghalaya); Laos (Haris 2014); Nepal (Gandaki); Thailand (Chiang Mai) (Fig. 6). This species is recorded from Nepal and Thailand for the first time. Remarks. The redescription of the species is based on the holotype kept in MNHN. The pictures of the female lancet were taken from a female collected from Hubei, China, which was compared in detail with the holotype. No distinct difference has been found between them including genitalia characters. C. rufidorsata is close to C. zhongi sp. nov. See the above key for the differences between the two species. Among the three species of Colochela, C. rufidorsata has the weakest body sculpture and punctation. Malaise (1937) stated in the original description of Colochela that the anal cell in forewing is with a short and perpendicular cross-vein at basal third. The examination of the holotype in MNHN (Paris) revealed that the anal cell in forewing of C. rufidorsata is without a cross-vein (Fig. 1 c). The female paratype of C. rufidorsata kept in USNM (Washington D. C.) does have a short anal cross-vein in both forewings (Figs 1 h���j) but the specimen is a female of C. zhongi sp. nov. We examined the male paratype of C. rufidorsata kept in USNM but overlooked the state of the basal anal cross-vein, so we do not sure whether the forewing of the male has basal anal cross-vein or not. In CNCI, a male specimen collected from Nepal during the Canadian Nepal expedition in 1967 refers to C. rufidorsata Malaise. The abdominal tergites are not microsculptured and are strongly shiny. The data of the specimen is: Nepal, Ktmd [Katmandu], Godavari, 5000 ��� [about 1,600 m], 31.VII. 1967, Can. Nepal Exped. (Dr. H. Goulet, personal communication)., Published as part of Niu, Gengyun & Wei, Meicai, 2016, Revision of Colochela (Hymenoptera, Tenthredinidae), pp. 457-470 in Zootaxa 4127 (3) on pages 461-465, DOI: 10.11646/zootaxa.4127.3.3, http://zenodo.org/record/259643, {"references":["Malaise, R. (1937) New Tenthredinidae mainly from the Paris Museum. Revue francaise d'Entomologie, 4, 43 - 53.","Malaise, R. (1945) Tenthredinoidea of South-Eastern Asia with a general zoogeographical review. Opuscula Entomologica, Lund Supplementum, 4, 1 - 288, 20 plates.","Singh, D. & Saini, M. S. (1990) First record of the genus Colochela (Hymenoptera, Tenthredinidae) from India. Entomon, 15, 271 - 272.","Lacourt, J. (1997) Contribution a une revision mondiale de la sous-famille des Tenthredininae (Hymenoptera: Tenthredinidae). Annales de la Societe Entomologique de France (N. S.), 32 [1996] (4), 363 - 402.","Haris, A. (2014) The subfamily Tenthredininae from Laos (Hymenoptera, Tenthredinidae). Natura Somogyiensis, 25, 73 - 80."]}

Colochela nigrata Wei & Niu, sp. nov. (Figs 1 a���b, 2 a, d, g, j, m, 3 e, h, 4 a, d, g, j, k, 5 a, d, e, j���l) urn: lsid:zoobank.org:act: 34 DA 4 F 27 - 2 B00- 4 F 77 - 9 F 34 -C 4 E 1 B 5 FFB 0E0 Description. Holotype: ♀. Length 14.5 mm (Fig. 1 a). Body black, body hairs black brown entirely. Wings strongly and evenly infuscate, vein R 1 largely brown, pterostigma and other veins black; hind wing slightly paler than forewing. Clypeus densely punctured except for narrow anterior margin, interspaces between punctures very narrow and smooth, hardly microsculptured; inner orbit, temple and postocellar area minutely and densely punctured, interspaces smooth, weakly shiny; punctures on frons extreme dense and fine, interspaces microsculptured; malar space densely microsculptured; punctures on hind orbit sparse and shallow; punctures on pronotum, mesoscutal middle lobe and lateral lobe, mesoscutellum, upper part of mesopleuron and metapleuron entirely much denser than punctures on head, interspaces hardly recognizable, almost mat; punctures on ventral part of mesepisternum gradually sparser and larger, interspaces hardly microsculptured, weakly shiny; punctures on mesoscutellar appendage sparse and large, surface smooth, shiny; metascutellum sparsely punctured, shiny; abdominal tergites 1��� 10 densely and minutely punctured, interspaces strongly microsculptured, matt; narrow intertergites smooth, strongly shiny. Hairs on dorsum of head and mesopleuron about 0.6 times as long as diameter of median ocellus; hairs on mesonotum extreme dense and about 0.4 times as long as diameter of median ocellus; abdominal tergites densely pubescent, extremely short, about 0.2 times as long as diameter of median ocellus; mesoscutellar appendage and intertergites of abdomen glabrous. Anterior margin of clypeus weakly incised, almost truncate; malar space 0.3 times as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits as long as longest axis of eye (Fig. 2 d); middle fovea small, round and deep, closed; frons flat, frontal ridge indistinct; POL: OOL: OCL = 13: 20: 22; interocellar furrow deep and broad, postocellar furrow shallow, weakly bent forward; postocellar area roundly elevated, about 1.5 times as broad as long; lateral furrow narrow and deep, weakly curved, hardly divergent backward; head slightly narrowed behind eyes in dorsal view (Fig. 2 a). Antenna 0.8 times as long as head and thorax together, distinctly incrassated beyond middle, 5 th ��� 9 th antennomeres with indistinct longitudinal furrow in both sides, third antennomere as long as following two antennomeres together, seventh antennomere about 1.5 times as long as broad (Fig. 3 e). Mesoscutellar appendage flat, with a distinct middle carina; cenchrus roundish, slightly broader than long, CD = 2. Venation as Fig. 1 a, basal anal cross vein present; petiole of hind anal cell about 0.3 times length of cu-a. Ovipositor sheath 1.5 times as long as front tibia, apical sheath as long as basal sheath, tapering toward apex (Fig. 2 m); sheath and setae in dorsal view as Fig. 2 j, sheath clearly tapering toward apex, about 2 times as broad as breadth of cercus; cercus small, about 2 times as long as broad. Lancet as Fig. 4 a, with about 27 serrulae, apical segment of lancet as Fig. 4 d, middle segments of lancet as Fig. 4 g, annular spines short and sparse, surface sculpture of lancet as Figs 4 j, k. Male. Body length 13.5 mm (Fig. 1 b); color, punctation and structure similar to female except for following: malar space 0.2 times as long as diameter of median ocellus (Fig. 2 g); distance between eyes at level of anterior tentorial pits about 0.9 times as long as longest axis of eye; head distinctly narrowed behind eyes in dorsal view; subgenital plate broader than long, apical margin round; harpes almost as long as broad (Fig. 5 a); volsella and gonolacinia as Figs 5 d���e; penis valve as Figs 5 j���l. Variation. The head of one female collected from Shaanxi Province is without lateral furrows. Etymology. The specific epithet refers to the entirely black body of the new species. Types: Holotype. ♀, CHINA: ANHUI: Lu���an, Central Forest Park, 75m, 1 ♀, 28.iv. 2015, Genbank accession number KT 833356 (CSCS). Paratypes: No data, 1 ♂, (CSCS); HUBEI: Baoji'an [Wuchang], 11.iv. 1954; SHAANXI: Xixiang, 560m, 1 ♀, 2.v. 1988 (CSCS); CHONGQING: 1 ♀, 20.vi. 1997, Wei Meicai (CSCS); ZHEJIANG: 1 ♂, 6.v. 1992, Wu Hong (CSCS). Other material examined. CHINA: Guangxi: Yangshuo, 1 ♀, 3.iv. 1938 (IZAS). Distribution. China (Shaanxi, Anhui, Hubei, Zhejiang, Chongqing, Guangxi) (Fig. 6). Remarks. See the above key for the differences between C. nigrata and the other two species of the genus. As to the specimen collected from Shaanxi that has the lateral furrows absent, no other distinct difference could be found between the specimen and others., Published as part of Niu, Gengyun & Wei, Meicai, 2016, Revision of Colochela (Hymenoptera, Tenthredinidae), pp. 457-470 in Zootaxa 4127 (3) on pages 460-461, DOI: 10.11646/zootaxa.4127.3.3, http://zenodo.org/record/259643

Neocolochelyna Malaise, 1937 Neocolochelyna Malaise, 1937: 47; Abe & Smith, 1991: 55; Lacourt, 1997: 389 [pt.]; Wei & Nie, 1998: 27 [pt.]; Wei, 2002: 254; Saini et al, 2006: 589 [pt.]; Saini, 2007: 53; Taeger et al., 2010: 566. Type species: Colochela montana Konow, 1898, by original designation. Curvatapenis Wei, 2002: Wei et al., 2006: 523; Taeger et al., 2010: 549. Type species: Curvatapenis testaceous Wei, 2002, by original designation. Treated here as a subgenus. Description. Body large to medium sized; black with brown maculae; wings hyaline with yellowish tinge, usually with infuscate shade in anterior part. Head and thorax strongly punctured, abdominal tergites strongly microsculptured. Clypeus less than two times as broad as long, anterior margin truncate to subtruncate (Figs 3 b, 3 e, 3 h, 4 b, 4 e, 4 h, 5 b, 5 e, 5 h); labrum small; mandibles short and stout, subsymmetrically tridentate, apical dent hardly bent (Figs 11 j���r); palpi distinctly reduced (Figs 3 a���c); malar space 0.6 ���1.0 times as long as diameter of median ocellus; eyes small or medium sized, inner margins weakly convex, distinctly convergent downwards, distance between them at level of lateral corner of clypeus about 1.1���1.4 times as long as longest axis of eye; distance between antennal sockets broader than distance between antennal socket and inner margin of eye (Figs 3 b, 3 e, 3 h, 4 b, 4 e, 4 h, 5 b, 5 e, 5 h); frons small, not depressed, frontal wall indistinct; postorbit round, occipital carina absent or present in lower half; postocellar area broader than long; inner orbit hardly oblique; supra-antennal tubercle absent; antenna short and stout, about as long as thorax, antennomere 2 slightly longer than broad, antennomere 3 longer than following 2 antennomeres together (Figs 11 a���i); anterior lobe of pronotum short, about 2 times as long as diameter of lateral ocellus, marginal carina absent; ventral margins of propleuron narrowly and roundly meeting; mesoscutellar appendage very short; anterior marginal carina of mesepisternum strongly elevated; distance between cenchri about 1.3���1.9 times as broad as a cenchrus; metapostnotum strongly oblique, divided into two narrow triangular plates by protruding middle corner of first abdominal tergite (Figs 6 a���i); dorsal apical lobe of metepimeron narrow elliptical with a short but distinct middle petiole. Hind coxa not strongly enlarged; apex of inner tibial spur of fore leg shallowly incised or distinctly bifurcate (Figs 7 j���r); hind tibia slightly longer than hind tarsus, hind tibial spur as long as apical width of tibia and less than 0.3 times of metabasitarsus; metabasitarsus distinctly compressed laterally, longer than following 3 tarsomeres combined but shorter than following 4 tarsomeres combined, tarsal pulvillus small; claw small, not robust, without basal lobe, inner tooth longer or shorter than outer tooth (Figs 7 a���i). Fore wing narrow and long, cell 2 Rs 0.75���1.2 times as long as 1 R 1 + 1 Rs, anal cell without cross vein; crossvein cu-a meeting cell 1 M at about basal 0.15, vein 2 A+ 3 A remote from vein 1 A at basal 0.25 with distance between them about 3���5 times as broad as width of vein 1 A at that point; hind wing with cell Rs and M closed, hind anal cell with a short petiole (Figs 1 d, 1g ���i). Setae on sheath short and almost straight, sheath in dorsal view tapering toward apex. Lancet soft, not distinctly sclerotized (Figs 8 a���d), serrulae almost flat with many fine subbasal teeth, annular spines sparse (Figs 9 a���h). Penis valve weakly sclerotized, with long or short dorsal hook (Figs 10 a, c, e, g, i); harpes much longer than broad, apical margin broadly roundish (Figs 10 b, d, f, h, j). Distribution. Eastern Asia (China, N. India, Japan). Host plant. Actinidia arguta Plauch of Actinidiaceae (known only for N. itoi: Okutani 1956; Yasumatsu 1957; Togashi 1995; Shikata & Itakura 2009). Remarks. Niu & Wei (2015) provided a key to separate Neocolochelyna and its relatives. Neocolochelyna can be easily recognized by the peculiar black and brown body color, the distinctly reduced palpi of mouthparts, the tridentate mandibles, the short antenna, the anal cell of forewing without cross vein, the upper hind orbit round without occipital carina, the long and soft lancet with many fine subbasal teeth, and the penis valve with a dorsal apical hook. Comparing the type species of Neocolochelyna and Curvatapenis, the two genus -group taxa can be separated clearly by several characters (see the key below). Here Curvatapenis is treated as a subgenus of the former because: A. Curvatapenis and Neocolochelyna share several important characters, such as the subsymmetrically tridentate mandibles, the reduced palpi of mouthparts, the short and stout antenna, the characteristic body color, the small claw, the soft lancet and peculiar penis valve; B. Among the 6 genera of Siobla group, i.e. Colochelyna, Colochela, Neocolochelyna, Curvatapenis, Corymbas, Neocorymbas (Sioblini of Takeuchi, 1952; Siobla + Neocolochelyni of Wei & Nie, 1998), Neocolochelyna and Curvatapenis are a pair of closely related taxa. Another similar pair of genera in the group is the pair of Corymbas and Neocorymbas. It is better to treat Curvatapenis and Neocorymbas as a subgenus of Neocolochelyna and Corymbas respectively, not only for understanding the evolution and differentiation within the Siobla group (this will be discussed in detail in a separate paper), but also for the practice of classification: both of them have only a few species at present and besides, it seems that they are possibly in the early stage of generic differentiation as shown by the characters differentiation. The generic status and species of Corymbas and Neocorymbas will be discussed and revised in a separate paper in near future. Actinidia is a genus of plants native to temperate eastern Asia, occurring throughout most of China, Korea and Japan, and extending north to southeast Siberia and south into Indochina (Wu et al. 2003). About 100 species and varieties have been found in China. Based on the previous collecting records, the adults of Neocolochelyna appear from late April to early June in mountains of central and west China, where collecting of sawflies have been made mainly from early June to early August. More species of the genus will probably be found in the future if sawfly collecting activity there shifts to an earlier season., Published as part of Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko & Naito, Tikahiko, 2016, Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae), pp. 432-456 in Zootaxa 4127 (3) on pages 433-434, DOI: 10.11646/zootaxa.4127.3.2, http://zenodo.org/record/264340, {"references":["Malaise, R. (1937) New Tenthredinidae mainly from the Paris Museum. Revue francaise d'Entomologie, 4, 43 - 53.","Abe, M. & Smith, D. R. (1991) The genus-group names of Symphyta (Hymenoptera) and their type species. Esakia, 31, 1 - 115.","Lacourt, J. (1997) Contribution a une revision mondiale de la sous-famille des Tenthredininae (Hymenoptera: Tenthredinidae). Annales de la Societe Entomologique de France (N. S.), 32 [1996] (4), 363 - 402.","Wei, M. (2002) A new genus and a new species of Tenthredininae (Hymenoptera: Tenthredinidae). In: Shen, X. & Zhao, Y. (Eds.), Insects of the Mountains Taihang and Tongbai Regions. (The Fauna and Taxonomy of Insects in Henan, Vol. 5 [2003]), China Agricultural Science and Technology Press, pp. 253 - 256. [In Chinese, abstract in English]","Saini, M. S., Blank, S. M. & Smith, D. R. (2006) Checklist of the sawflies (Hymenoptera: Symphyta) of India. In: Blank, S. M., Schmidt, S. & Taeger, A. (Eds.), Recent Sawfly Research - Synthesis and Prospects. Goecke & Evers, Keltern, pp. 575 - 612.","Saini, M. S. (2007) Subfamily Tenthredininae Sans Genus Tenthredo. In: Indian Sawflies Biodiversity. Keys, Catalogue & Illustrations. Vol. 2. Bishen Singh Mahendra Pal Singh, Dehra Dun, pp. [1 - 8] + 1 - 234.","Taeger, A., Blank, S. M. & Liston, A. D. (2010) World Catalog of Symphyta (Hymenoptera). Zootaxa, 2580, 1 - 1064.","Konow, F. W. (1898) Neue Chalastogastra-Gattungen und Arten. Entomologische Nachrichten, 24 (17 - 18), 268 - 282.","Wei, M., Nie, H. & Taeger, A. (2006) Sawflies (Hymenoptera: Symphyta) of China - Checklist and review of research. In: Blank, S. M., Schmidt, S. & Taeger, A. (Eds.), Recent Sawfly Research - Synthesis and Prospects. Goecke & Evers, Keltern, pp. 505 - 574.","Okutani, T. (1956) Some records on the food-plants for Japanese sawflies with a note on an egg-laying habit of Macrophya apicalis (Studies on Symphyta IV). The Science Reports of the Hyogo University of Agriculture, Agricultural Biology, 2 (2), 1 - 2.","Yasumatsu, K. (1957) On the host-plant of Neocolochelyna itoi Takeuchi. Kontyu, 25 (3), 101. [In Japanese]","Togashi, I. (1995) Description of Neocolochelyna hakusana, sp. nov., and its larva (Hymenoptera: Tenthredinidae) from Japan. Proceedings of the Entomological Society of Washington, 97 (4), 866 - 871.","Shikata, K. & Itakura, T. (2009) First record of Neocolochelyna itoi in Nagano Prefecture. Natural History Reports of Inadani, 10, 75 - 77. [In Japanese]","Niu, G. & Wei, M. (2015) Revision of Colochela (Hymenoptera, Tenthredinidae). Zootaxa, in press.","Takeuchi, K. (1952) A Generic Classification of the Japanese Tenthredinidae (Hymenoptera: Symphyta). Kyoto, 1 - 90.","Wu, Z., Lu, A., Tang, Y., Chen Z. & Li, D. (2003) The families and genera of Angiosperms in China, a comprehensive analysis. Science Press, Beijing, 1 - 1209, pls. 16 color plates."]}

Key to Colochela, Colochelyna and Neocolochelyna 1 Anal cell in forewing with a long and suberect cross vein at about middle; occipital carina present; mandibles asymmetric, left mandible bidentate, right mandible simple or with an indistinct inner tooth; penis valve narrow and simple, without distinct lateral lobe or hook; wings strongly yellowish; claw with inner tooth much longer than outer tooth; lancet very narrow, more or less sclerotized; clypeus narrow and long, about 1.3–1.4 times as broad as long; anterior-lateral lobe of pronotum (before transversal furrow) with fine but distinct marginal or submarginal carina; head small, about 0.7 times as broad as thorax. [Cell 2 Rs in forewing much longer than 1 R 1 + 1 Rs]........................................... Colochelyna (3 known species) - Anal cell in forewing with a very short cross vein at basal 0.25 or without cross vein; occipital carina absent; mandibles subsymmetric bidentate or tridentate; penis valve with distinct lateral or dorsal lobe or hook; wings blackly infuscate, or claw with inner tooth shorter than outer tooth and lancet not very narrow and not sclerotized; clypeus about 1.9–2 times as broad as long; anterior-lateral lobe of pronotum (before transversal furrow) without marginal or submarginal carina; head about 0.9 times as broad as thorax....................................................................................... 2 2 Wings strongly infuscate, almost blackish, without longitudinal shade along anterior margin; forewing with cell 2 Rs much longer than 1 R 1 + 1 Rs together; basal third of anal cell strongly constricted, distance between veins 1 A and 2 A+ 3 A about as broad as 1 A at same position, with or without cross vein; mandibles bidentate; lancet distinctly sclerotized; basal and middle serrula without distinct subbasal tooth, apical serrulae sometimes with a few subbasal teeth; penis valve without dorsal hook but with a ventral apical lobe, or apical margin broadly rolling up laterally; body largely or entirely black, at most mesonotum red; antenna longer than thorax; palpi of mouthparts normal, hardly reduced; cerci slender, about 2–4.5 times as long as broad; distance between antennal sockets broader than distance between antennal socket and inner margin of eye at same level; setae on sheath long and strongly curved; claw quite large, inner tooth much longer than outer tooth..... Colochela (3 known species) - Wings brown, with distinct longitudinal shade along anterior margin; forewing with cell 2 Rs shorter than 1 R 1 + 1 Rs together; basal third of anal cell weakly constricted, distance between veins 1 A and 2 A+ 3 A at least 2 times as broad as 1 A at same position, without anal cross vein; mandibles tridentate with a small third tooth; lancet not sclerotized, serrulae each with many minute subbasal teeth; penis valve with a long dorsal hook, apical margin not rolling up laterally; body yellow brown or ferruginous with black maculae; antenna not longer than thorax; palpi of mouthparts much reduced; cerci very short, about as long as broad; distance between antennal sockets clearly shorter than distance between antennal socket and inner margin of eye at same level; setae on sheath short and straight; claw small with inner tooth shorter than outer tooth in subgenus Curvatapenis, or small or medium-sized with inner tooth longer and stronger than outer tooth in nominal subgenus.................................................................................................... Neocolochelyna (6 species)

Key to Colochela, Colochelyna and Neocolochelyna 1 Anal cell in forewing with a long and suberect cross vein at about middle; occipital carina present; mandibles asymmetric, left mandible bidentate, right mandible simple or with an indistinct inner tooth; penis valve narrow and simple, without distinct lateral lobe or hook; wings strongly yellowish; claw with inner tooth much longer than outer tooth; lancet very narrow, more or less sclerotized; clypeus narrow and long, about 1.3���1.4 times as broad as long; anterior-lateral lobe of pronotum (before transversal furrow) with fine but distinct marginal or submarginal carina; head small, about 0.7 times as broad as thorax. [Cell 2 Rs in forewing much longer than 1 R 1 + 1 Rs]........................................... Colochelyna (3 known species) - Anal cell in forewing with a very short cross vein at basal 0.25 or without cross vein; occipital carina absent; mandibles subsymmetric bidentate or tridentate; penis valve with distinct lateral or dorsal lobe or hook; wings blackly infuscate, or claw with inner tooth shorter than outer tooth and lancet not very narrow and not sclerotized; clypeus about 1.9���2 times as broad as long; anterior-lateral lobe of pronotum (before transversal furrow) without marginal or submarginal carina; head about 0.9 times as broad as thorax....................................................................................... 2 2 Wings strongly infuscate, almost blackish, without longitudinal shade along anterior margin; forewing with cell 2 Rs much longer than 1 R 1 + 1 Rs together; basal third of anal cell strongly constricted, distance between veins 1 A and 2 A+ 3 A about as broad as 1 A at same position, with or without cross vein; mandibles bidentate; lancet distinctly sclerotized; basal and middle serrula without distinct subbasal tooth, apical serrulae sometimes with a few subbasal teeth; penis valve without dorsal hook but with a ventral apical lobe, or apical margin broadly rolling up laterally; body largely or entirely black, at most mesonotum red; antenna longer than thorax; palpi of mouthparts normal, hardly reduced; cerci slender, about 2���4.5 times as long as broad; distance between antennal sockets broader than distance between antennal socket and inner margin of eye at same level; setae on sheath long and strongly curved; claw quite large, inner tooth much longer than outer tooth..... Colochela (3 known species) - Wings brown, with distinct longitudinal shade along anterior margin; forewing with cell 2 Rs shorter than 1 R 1 + 1 Rs together; basal third of anal cell weakly constricted, distance between veins 1 A and 2 A+ 3 A at least 2 times as broad as 1 A at same position, without anal cross vein; mandibles tridentate with a small third tooth; lancet not sclerotized, serrulae each with many minute subbasal teeth; penis valve with a long dorsal hook, apical margin not rolling up laterally; body yellow brown or ferruginous with black maculae; antenna not longer than thorax; palpi of mouthparts much reduced; cerci very short, about as long as broad; distance between antennal sockets clearly shorter than distance between antennal socket and inner margin of eye at same level; setae on sheath short and straight; claw small with inner tooth shorter than outer tooth in subgenus Curvatapenis, or small or medium-sized with inner tooth longer and stronger than outer tooth in nominal subgenus.................................................................................................... Neocolochelyna (6 species), Published as part of Niu, Gengyun & Wei, Meicai, 2016, Revision of Colochela (Hymenoptera, Tenthredinidae), pp. 457-470 in Zootaxa 4127 (3) on page 459, DOI: 10.11646/zootaxa.4127.3.3, http://zenodo.org/record/259643

Neocolochelyna (Neocolochelyna) montana (Konow, 1898) (Figs 1 a, 3 a���c, 6 a, 7 a, 7 j, 8 c, 8g, 8 j, 9 c, 9g, 11 a, 11 j) Colochelyna montana Konow, 1898: 281; Konow, 1905: 118; Malaise, 1937: 47. Neocolochelyna montana: Malaise, 1937: 47; Malaise, 1945: 114; Saini et al., 2006: 589; Saini, 2007: 54. Description. Female. Holotype (Fig. 1 a). Length 16 mm. Head brown with a large black macula covering narrow posterior margin of clypeus, supraclypeal area, inner orbit, frons, ocellar area, anterior of temple and anterior 0.6 of postocellar furrow; labrum yellow brown (Fig. 3 a), lower part of posterior side of head blackish; antenna entirely brown, basal part hardly paler than apex (Fig. 11 a). Thorax black, narrow lateral margin and very broad posterior margin of pronotum brown; mesoscutellum (not including mesoscutellar appendage), a short stripe on posterior margin of mesoscutal lateral lobe, anterior and posterior margins of parapsis, metascutellum, cenchri, postspiracular sclerite, prepectus (not epicnemium) and dorsal margin of metapleuron yellow brown. Abdomen black, narrow posterior margin of tergite 1 (Fig. 6 a), about apical half of tergites 2���10 and apical 0.25 of each sternite dark reddish brown to yellow brown, cerci and ovipositor sheath pale brown. Body hairs brown to pale brown. Legs orange, ventral side of each coxa and first trochanter of each leg black, outer side of hind femur dark brown, all tarsi largely yellow brown. Wings yellowish infuscate in anterior part and gradually paler toward posterior margin and apex, vein C and pterostigma pale brown, posterior of vein Sc+R and most of vein M+Cu blackish; hind wing weakly infuscate, clearly paler than forewing. Punctures on labrum large and shallow, surface smooth, shiny; punctures on clypeus dense and large, narrow interspaces between punctures hardly microsculptured, shiny; frons and inner orbit coarsely and irregularly punctured, narrow interspaces microsculptured, mat; punctures on temple and postocellar area sparser than inner orbit, interspaces smaller than punctures, surface clearly but weakly microsculptured, less shiny; malar space densely punctured (Figs 3 a���c); hind orbit densely punctured, interspaces linear, microsculptured, hardly shiny; upper corner of pronotum densely punctured and almost dim, depressed part of pronotum coarsely punctured and microsculptured, almost mat, narrow lateral margin of pronotum shiny; mesoscutal middle lobe and lateral lobe densely punctured, interspaces very narrow, distinctly microsculptured, punctures on posterior part of lateral lobe much denser, almost dim; punctures on mesoscutellum large and dense, interspaces narrow but flat, weakly microsculptured, less shiny; mesoscutellar appendage distinctly coriaceous in lateral side and smooth in middle part, impunctate; parapsis and metapostnotum strongly microsculptured (Fig. 6 a); punctures on upper half of mesepisternum large and very dense, interspaces very narrow, hardly shiny; punctures on lower half of mesepisternum small and much sparse, surface strongly microsculptured, almost dim; mesepimeron and metapleuron strongly microsculptured mixed with large and dense punctures, narrow anterior and posterior margins of mesepimeron and of metapleuron shiny; abdominal tergite 1 strongly microsculptured, almost mat, lateral side with some shallow and large punctures (Fig. 6 a); tergites 2���8 strongly microsculptured, mat, posterior margin of tergites 4���8 with some shallow and indistinct punctures; median part of tergite 10 weakly microsculptured, distinctly shiny; surface of basal sheath microsculptured mixed with some shallow punctures, apical sheath hardly coriaceous, ventral margin with a few shallow punctures, shiny. Hairs on dorsum of head and mesopleuron dense, hardly curved at apex, about 0.9���1 times diameter of median ocellus (Figs 3 a���b); hairs on mesoscutal middle and lateral lobes about 0.8 times as long as diameter of median ocellus; dorsum of abdominal tergites 1, 4��� 8 distinctly pubescent, hairs on tergite 1 not longer and denser than other tergites, tergites 2���3 largely, mesoscutellar appendage and intertergites of abdomen entirely glabrous (Fig. 6 a). Labrum subtriangular, apical corner distinctly angled but not very acute; clypeus flat, without middle ridge, anterior margin almost truncate, very shallowly and broadly incised; malar space as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits 1.2 times as long as longest axis of eye (Fig. 3 b); left mandible as Fig. 11 j; hind orbit with a distinct occipital carina in lower half; middle fovea roundish, deep, not open to frons; lateral fovea minute, with an indistinct tubercle; frons subquadrate, frontal ridge indistinct; POL: OOL: OCL= 15: 30: 24; interocellar furrow short, broad and deep, postocellar furrow narrow and indistinct, clearly bent forward at middle; postocellar area flat, clearly below temple, 1.7 times as broad as long, posterior margin not elevated; lateral furrows very shallow and indistinct in anterior 0.7, replaced by a low carina in posterior 0.3, straight, distinctly divergent backwards; head roundly dilated behind eyes in dorsal view (Fig. 4 a). Antenna as long as thorax, antennomere 3 about 1.1 times as long as antennomeres 4 and 5 together and 0.9 times as long as antennomeres 6���9 together, antennomere 7 about 1.2 times as long as broad (Fig. 11 a). Mesoscutal middle lobe with narrow but distinct middle furrow; mesoscutellum weakly and roundly elevated, anterior margin subtriangularly protruding, transverse anterior furrow shallow; mesoscutellar appendage short with middle length about 1.1 times as long as diameter of median ocellus; mesoscutellum 6 times as long as mesoscutellar appendage; cenchrus transverse, slightly broader than long, CD= 1.5 (Fig. 6 a); mesepisternum roundly and weakly elevated at middle, without distinct ridge. Venation as Fig. 1 a, cell 2 Rs about 0.8 times as long as cells 1 R 1 + 1 Rs, cell A of forewing weakly constricted at basal 0.25, where distance between 1 A and 2 A+ 3 A about 5 times as broad as vein 2 A+ 3 A; vein 1 M as long as vein 1 m-cu, vein 2 r 1 meeting cell 2 Rs at apical 0.35. Inner tibial spur of fore leg as Fig. 7 j, apical incision narrow, very deep; claw as Fig. 7 a, inner tooth clearly longer than apical tooth. Triangular middle lobe of sternite 7 small, much narrower than lateral straight posterior margin of sternite 7 (Fig. 8 j). Ovipositor sheath 0.9 times as long as front tibia and 0.7 times as long as middle tibia, apical sheath 1.1 times as long as basal sheath, narrowly truncate at apex in lateral view. Lancet as Fig. 8 c, with 26 serrulae, middle and apical segments as Figs 9 c, 9g, annular spines and surface sculptures of lancet as Fig. 9 g; basal sensilla as Fig. 8 g. Male. Body length 14 mm; color, punctation and structure similar to female except for following: head black, labrum and narrow posterior margin of temple brown; each femur largely dark brown; malar space 0.8 times as long as diameter of median ocellus; distance between eyes at level of anterior tentorial pits about 1.1 times as long as longest axis of eye; subgenital 1.1 times as long as broad, apical margin obtusely truncate; harpes about 2 times as long as broad; penis valve not dissected but similar to Fig. 10 a in outline, dorsal apical hook long and strongly tapering toward apex. Distribution. India (Sikkim) (Fig. 12). Material examined. Type. Holotype: ♀, ���Sikkim���; ���Coll. Konow���; ��� Colochelyna montana Knw., Sikkim.���; ��� Holotypus ���[red]; ��� Typus ���[red]; ���GBIF-GISHym 4192 ���. (SDEI). Other specimens. 1 ♀, MUSEUM PARIS, DARJILING, (MONTAGNES), HARMAND 1888; 3363, 86; Neocolochelyna montana Konow, Malaise det., 1935; Coll. Malaise; montana. (SNHM). 1 ��, ���SIKKIM, Runjit Valley, 4-94, BINGHAM COLL ���; ��� COLL. C. T. Bingham, 96 - 30 ���; ���British Museum���; ��� Colochelyna montana Knw, Malaise det.��� (BMNH). Host plant. Unknown. Remarks. N. montana is similar to N. acutilabria sp. nov.. See the above key for the differences between the two species. The holotype has a large hole on right side of mesonotum. Besides, the left prepectus (not epicnemium) in front of the postspiracular sclerite is much larger than the lateral ocellus. It should be abnormal because the prepectus on the right side is unrecognizable, and in N. acutilabria the prepectus is also very small and linear., Published as part of Niu, Gengyun, Wei, Meicai, Shinohara, Akihiko & Naito, Tikahiko, 2016, Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae), pp. 432-456 in Zootaxa 4127 (3) on pages 453-455, DOI: 10.11646/zootaxa.4127.3.2, http://zenodo.org/record/264340, {"references":["Konow, F. W. (1898) Neue Chalastogastra-Gattungen und Arten. Entomologische Nachrichten, 24 (17 - 18), 268 - 282.","Malaise, R. (1937) New Tenthredinidae mainly from the Paris Museum. Revue francaise d'Entomologie, 4, 43 - 53.","Malaise, R. (1945) Tenthredinoidea of South-Eastern Asia with a general zoogeographical review. Opuscula Entomologica, Suppl. 4, 1 - 288.","Saini, M. S., Blank, S. M. & Smith, D. R. (2006) Checklist of the sawflies (Hymenoptera: Symphyta) of India. In: Blank, S. M., Schmidt, S. & Taeger, A. (Eds.), Recent Sawfly Research - Synthesis and Prospects. Goecke & Evers, Keltern, pp. 575 - 612.","Saini, M. S. (2007) Subfamily Tenthredininae Sans Genus Tenthredo. In: Indian Sawflies Biodiversity. Keys, Catalogue & Illustrations. Vol. 2. Bishen Singh Mahendra Pal Singh, Dehra Dun, pp. [1 - 8] + 1 - 234."]}

Siobla pulchra Shinohara, Wei & Niu, sp. nov. (Figs 2 g–h, 5 o–p, 6g, 7g, 8g, p, 9 a–e, 10 a, e) urn:lsid:zoobank.org:act: 2 A07DDB 0-3 BA 3-4567 -BD 9 B- 5 F 4 D0A 67 CD 54 Siobla ferox: Naito et al., 2004: 56 (in part). Not Smith (1874). Siobla sp. 1: Tanaka & Tanaka, 2007: 109. Description. Female. Holotype (Figs 2 g, 5 o). Length 15.0 mm. Head and thorax black with following parts yellow white: clypeus, labrum, middle of mandible, antenna, posterior corner of pronotum, tegula, mesoscutellum, mesoscutellar appendage, metascutellum, minute stripe on middle of mesepimeron, and upper margin of metepimeron; palpi, narrow stripe on upper half of inner orbit, broad anterior stripe on upper half of hind orbit, and posterior part of postocellar area yellow brown. Abdomen yellow white with tergum 1 largely, small macula on tergum 3, most of terga 4 and 8 dorsally, terga 5–7 dorsally, and basal half of tergum 9 black. Legs yellow brown with base of fore coxa, middle and hind coxae, and apical 0.65 of hind femur black, and ventral side of hind tibia dark brown at apex. Body hairs yellow brown. Wings hyaline with weak yellow brown tinge, apical third distinctly infuscated, vein C and stigma yellow brown, other veins black brown. Apex of hind wing slightly grayish. Dorsum of head coarsely and densely punctured, interspaces between punctures on temple hardly recognizable, without microsculpture; clypeus shallowly and densely punctured, shiny; labrum shallowly and sparsely punctured; punctuation on dorsum of thorax similar to head; mesoscutellum matt, extremely densely punctured; mesoscutellar appendage polished, shiny, lateral corners microsculptured; metascutellum densely and coarsely punctured, anterior slope with linear interspaces between punctures; metapostnotum polished, shiny; mesepisternum feebly shiny, dorsal half coarsely and densely punctured and ventral half with small and shallow, dense, punctures; metepisternum with dorsal part densely punctured and ventral part sparsely punctured, shiny; middle and posterior part of metepimeron shiny, dorsal part of metepimeron coarsely and densely punctured; abdominal terga 1–3 strongly shiny, without microsculpture, other terga weakly microsculptured, terga 4–9 laterally and outer side of hind coxa shallowly and sparsely punctured. Hairs on temple straight, 1 × diameter of median ocellus. Clypeus 2.2 × as broad as long, with anterior margin truncate; malar space 1 × diameter of median ocellus; lower interocular distance 1.2 × eye height (Fig. 10 a); anterior margin of supraantennal tubercle elevated, small, posteriorly confluent with frontal ridge; middle fovea round and broad, deep; lateral fovea small and deep; interocellar furrow narrow and deep; postocellar furrows broad and shallow; postocellar area weakly elevated, as high as top of ocelli, with sharp middle carina, about 1.4 × as broad as long; lateral furrows deep, distinctly curved outwards in anterior half and subparallel posteriorly; head behind eyes slightly shorter than eyes in dorsal view, convex at basal 0.4 and distinctly narrowing posteriorly (Fig. 9 a). Antenna shorter than vein C of forewing, about as long as head, thorax and abdominal tergum 1 together; pedicel slightly longer than broad, antennomere 3 1.5 × as long as antennomere 4, subapical antennomeres slightly enlarged, antennomere 7 2 × as long as broad (Fig. 9 e). Middle furrow on pronotum deep; mesoscutellum weakly elevated, anterior slope quite flat, much longer than posterior slope, without peak and carina; mesoscutellar appendage almost flat, without middle carina; mesepisternum roundly elevated, ventral spur absent. Apex of hind tibia distinctly enlarged; metabasitarsus 3.8 × as long as broad, 1.1 × length of following 3 tarsomeres combined; hind inner tibial spur 0.5 × length of metabasitarsus; pulvilli small, distance between basal two pulvilli 2.5 × length of second pulvillus. Hind wing with petiole of anal cell 0.7 × length of cu-a. Abdominal tergum 1 with distinctly elongated carina just above spiracle; sternum 7 with each lateral part roundly incised; ovipositor sheath 0.93 × length of middle tibia, apical sheath subequal in length to basal sheath (Fig. 9 d); lancet as in Fig. 6 g, middle serrulae as in Fig. 7 g. Male. Paratype from Nakayama-toge. Length 14.0 mm (Figs 2 h, 5 p). Head and thorax black, labrum, mesoscutellum, mesoscutellar appendage, metascutellum and dorsum of metepimeron yellow white; palpi and tegula yellow brown; antenna yellow brown, scape entirely and pedicel largely black; abdomen yellow brown, basal 0.65 of tergum 1 and dorsum of terga 5–8 black. Legs yellow brown; each coxa, broad and long inner stripe and outer apical 0.35 of hind femur black; dorsal stripe on fore and middle femora and ventral half of hind tibia black brown. Punctuation and structure similar to female except as follows: eyes large, shortest distance between eyes almost as long as height of eye; malar space 0.8 × diameter of lateral ocellus (Fig. 9 c); lateral side of temples strongly narrowed backward in dorsal view (Fig. 9 b); pulvilli minute; subgenital plate longer than broad, apex round; penis valve as in Fig. 8 p; gonoforceps as in Fig. 8 g, harpe broad, less than 2 × as long as broad. Etymology. The specific epithet, meaning beautiful, refers to the appearance of the new species. Distribution. Japan (Hokkaido, Honshu). Holotype (Figs 2 g, 5 o): ♀, [JAPAN: Honshu], Minoto, 1850m, Yatsugatake Mts., Nagano Pref. 25–29. VII. 2000, A. Shinohara, NSMT-I-HYM 68233, genitalia slide N 20131105 A (NSMT). Paratypes: 17 specimens (6 ♀ and 9 ♂, see Appendix). Variation. The length varies from 13.5 to 15.0 mm in the female and 12.5 to 14.0 mm in the male. The female specimens examined show little variation in colour pattern. The valvifer 2 (basal sheath) is subequal in length to or slightly longer than the valvifer 3 (apical sheath). Five males from Hyogo Prefecture, western Honshu, have the broad anterior margin of the clypeus pale yellow medially and the hind femur largely black. The ranges of some ratio variations are given in the key. Remarks. This new species belongs to the ferox group defined by Niu & Wei (2010) and is similar to S. fulvomarginata Wei & Nie, 1999, from Henan and Hunan Provinces, China. Siobla pulchra differs from S. fulvomarginata in the following features: head and thorax in both sexes largely black with a few yellow white maculae; eyes small, shortest distance between them longer than height of eye; postocellar area with distinct middle carina; dorsum of mesoscutellum flat, below top of mesoscutum; tergum 1 with distinct carina just lateral of spiracle; dorsum of head and thorax coarsely and densely punctured; apical 0.3 of forewing distinctly infuscate; tergum 1 and dorsum of terga 5–8 largely black and male subgenital plate longer than broad. Siobla pulchra differs from the Japanese congeners in the following characters. Female: clypeus, entire antenna, mesoscutellum, mesoscutellar appendage, metascutellum, and valvifer 2 (basal sheath) yellowish white; upper hind orbit and postocellar area marked with yellowish brown; postocellar area with distinct middle carina; forewing apically distinctly infuscated; mesoscutellum weakly raised; hind tibia mostly orange, with only obscure black spot near apex; tergum 1 largely black; abdominal segments 4 to 7 mostly whitish or pale orange ventrally; sternum 7 with each lateral part roundly incised; valvifer 2 (basal sheath) subequal in length to or slightly longer than valvifer 3 (apical sheath). Male: Clypeus entirely black or with broad anterior margin pale yellow medially; antenna yellowish white, with scape and most of pedicel black, and flagellomere 1 partly darkened; mesoscutellum, mesoscutellar appendage and metascutellum whitish yellow; mesoscutellum weakly swollen; forewing apically distinctly infuscated; hind tibia mostly orange, with only obscure black spot near apex; tergum 1 with basal half black. Tanaka & Tanaka (2007) noted that their Siobla sp. 1 “resembles S. ferox but the apex of the forewing is dark and there is a difference in hind tibia” (original in Japanese, p. 109; see also fig. 1 in p. 113). This species is probably S. pulchra.

Siobla atra Malaise, 1945 (Figs 3 a, 3 b, 5 e, 5 f, 7 j, k, 7 n, 7 p, q, 9 c, 9 p) Siobla atra Malaise, 1945: 123. ♀, type locality: Kambaiti (Myanmar). (Excluding males). Siobla rufipes Malaise, 1945: 124. ♂, type locality: Kambaiti (Myanmar). Syn. nov. Siobla rufipes var. infuscata Malaise, 1945: 124. ♂, infrasubspecific name. Description. Lectotype: ♀. Length, 11.5 mm (Figs 3 a, 5 e). Body and legs black; very narrow posterior margin of pronotum dark brown, 2 nd abdominal tergite with small white spots near spiracles; anterior of fore femur and most of tibia and of tarsus brown, middle tarsus black brown, hind trochanter black brown. Body hairs silver, hairs on dorsal side of head pale brown. Wings hyaline, vein C black, pterostigma black brown, other veins deep brown or blackish brown. Punctures on head large, dense and coarse without shiny interspaces, labrum sparsely punctured; prescutum, scutum, and anterior slope of mesoscutellum densely punctured, with narrow interspaces; lateral and posterior slope of mesoscutellum densely punctured, without interspaces; posttergite shiny with lateral corners microsculptured; parapsides with large and sparse punctures and dense microsculpture; metascutellum with anterior slope shiny with microsculpture and posterior slop densely and irregularly punctured; metapostnotum shiny with margins slightly microsculptured; middle and upper of mesepisternum densely and coarsely punctured, without shiny interspaces, venter with large and shiny interspaces between small and sparse punctures; anepimeron coarsely punctured, upper of katepimeron with large punctures, interspaces obviously longer than diameter of punctures, middle and lower of katepimeron microsculptured; upper of metepisternum with dense and small punctures, interspaces shiny, narrower than diameter of punctures, venter sparsely punctured, shiny; basin of metepimeron weakly microsculptured, dorsum with large punctures and posterior corner shiny. First abdominal tergite feebly microsculptured, shiny, lateral edges slightly punctured; other tergites obviously microsculptured, almost mat, lateral edges of both 3 rd and 4 th tergites, posterior of 4 th tergite, and most of 5 th to 8 th tergites obviously punctured. Hairs on dorsum of head straight, as long as diameter of lateral ocelli. Clypeus without a transverse carina, anterior margin truncate; malar space as long as transverse diameter of median ocellus; shortest distance between eyes below antennal sockets in front view 1.1 �� eye height; middle fovea broad with a short deep furrow on bottom; interocellar furrow deep, postocellar furrow broad and shallow; postocellar area 1.5 �� as broad as long, weakly elevated and clearly below top of ocelli, without middle ridge; lateral furrows weakly curved, deep in entire length, subparallel; head behind eyes 0.8 �� as long as eye in dorsal view; curved laterally and then convergent posteriorly; genal carina sharp, complete (Fig. 7 j). Antenna as long as head, thorax and abdominal tergites 1���2 together, as long as vein C, 2 nd antennomere 1.5 �� as long as broad, relative length of 3 rd to 4 th antennomeres as 1.7, 5 th to 8 th antennomeres hardly compressed, outer sides of 5 th to 8 th antennomeres with weak furrows, 7 th antennomere 2.6 �� as long as broad. Middle furrow of prescutum distinct; mesoscutellum distinctly elevated, as high as scutum. Metabasitarsus 5 �� as long as broad, 0.8 �� length of following 4 tarsomeres combined, hind inner tibial spur 0.45 �� length of metabasitarsus. Pulvilli of hind tarsus small. Ovipositor sheath as long as middle tibia in lateral view; exposed part of apical sheath about 2 �� as long as broad, apical sheath 1.5 �� as long as basal sheath, apical sheath narrow in dorsal view, exposed part 4 �� length of cerci, lateral setae on both sides of sheath straight, forming an angle about 50 degree. Lancet broadened before apex, with 16 serrulae (Fig. 9 c), middle serrulae each with 18���20 distal fine teeth (Fig. 9 p). Male. Length, 10 mm (Figs 3 b, 5 f). Body black; 2 nd to 4 th abdominal tergites, 2 nd to 5 th abdominal sternites, and most of 6 th sternite yellowish brown; legs yellowish brown to orange, each coxa, fore and middle trochanter, a small dot on apex of hind femur black; similar to female in structure, but malar space 0.5 �� transverse diameter of median ocellus (Fig. 7 q); shortest distance between eyes below antennal sockets in front view 1.05 �� eye height; head behind eyes 0.6 ��eyes in dorsal view, curved laterally and then strongly convergent posteriorly; genitalia as in Figs 10 u, 10 v. Variation. A female collected from Yunnan has the hind trochanter dark brown and first abdominal tergite less microsculptured. A male collected in Myanmar, which is named as S. rufipes var. infuscata Malaise, 1945, has the second tergite black. Specimens examined. MYANMAR: Lectotype (hereby designated), Siobla atra ♀, ���N. E. BURMA, Kambaiti, 7000ft, 24 / 5, R. MALAISE���; [red] ��� Typus ���; ��� Siobla atra n. sp., Malaise det. 1936 ���; ���NHRS-HYME 000005304���; ��� Lectotype, Siobla atra Malaise, 1945, Design Niu G. 2012 ��� (NRHS); Paralectotypes of S.atra: 2 ♀, same data as the lectotype, but 28 /5, 1 ♀ same data but 8 / 5. The specimens bear the ID numbers NHRS-HEVA 0 0 0 0 0 1608 - NHRS-HEVA 0 0 0 0 0 1610 (all NRHS). 1 ♀, same data as the lectotype, but 24 / 5 (CSCS); 2 ♀, same data (BMNH); 1 ♀, same data as the lectotype, but 28 / 5 (MNCN, teste Taeger). [Furthermore, 21 ♂ that belong to S. semipicta, see there]. Lectotype (hereby designated), Siobla rufipes ♂, ���N. E. BURMA, Kambaiti, 7000ft, 26 / 5, R. MALAISE���, ��� Typus ���, "NHRS-HYME 000005297���, Siobla rufipes n. sp., Malaise det. 1936; Lectotype, Siobla rufipes Malaise, 1945, Design Niu G. 2012; Siobla atra Malaise, 1945 Det. Niu G. (NRHS); Paralectotypes of S. rufipes: According to Taeger (pers. comm.), in addition to the lectotype there are 35 males at the NRHS. 14 are from "Kambaiti 7000 ft.", 17 from "Kambaiti 2000 m ", 3 "Kambaiti 1800m ", and one without data. Only 3 of these specimens got Malaise���s identification labels. The 33 specimens from 7000 ft. and 2000 m respectively, are considered to belong to the type series: 1 ♂ ���N. E. BURMA, Kambaiti, 7000ft, 23 / 4, R. MALAISE���; 3 ♂, same data but 24 / 4; 2 ♂, same data but 28 / 4; 2 ♂, same data but 29 / 4; 4 ♂, same data but 22 / 5; 1 ♂, same data but 26 / 5; 1 ♂, same data but 5 / 6; 1 ♂, same data but 2000 m and 4 / 4; 4 ♂, same data but 15 / 5; 12 ♂, same data but 1 / 6. The specimens bear the ID numbers NHRS-HEVA 0 0 0 0 0 1555 - NHRS-HEVA 0 0 0 0 0 1585. 1 ♂, ���N. E. BURMA, Kambaiti, 7000ft, 21 / 5, R. MALAISE���, ��� Typus ��� ���var. S. rufipes n. sp. infuscatus var., det. Malaise, 1936 ���, ��� Paralectotype, Siobla rufipes Malaise, 1945, Design Niu G. 2012 ��� (all NRHS). 13 ♂ same data as lectotype, but collecting days not recorded. (BMNH); 1 #m same data as lectotype but 2000m, 9. V. 1934 (HNHM); 1 #m same data as lectotype but 1. V. 1934 (CSCS); 1 ♂, same data as the lectotype, but 28 / 4 (MNCN, teste Taeger). Other material: CHINA: Yunnan:. 1 ♀, Heiwadi, Gongshan, Yunnan, 27.800 ��N, 98.590 �����E, alt. 2000m, 6 Jun. 2009, Zhong Yihai leg. (CSCS), 1 ♂, Gongshan 10km NW, +27.79700, +98.58500, 2150m, 0 6.06. 2009, Leg. S. M. Blank, A. D. Liston & A. Taeger; Siobla atra Malaise, 1945 Det. Niu G. (SDEI) barcode: BOLD:ABV 4520 (Taeger pers. comm.). Distribution. China (Yunnan); Myanmar. Remarks. This species is close to S. carinoclypea Niu &Wei, 2010 but differs from it in the anterior margin of clypeus truncate without a middle incision and without transverse carina (Fig. 7 n, 7 k); the postocellar area weakly elevated (Fig. 7 k), clearly below top of ocelli, and 1.5 �� as broad as long; the lateral furrows of postocellar area curved and deep in entire length (Fig. 7 j); the ovipositor sheath as long as middle tibia; the lancet with 16 serrulae and the middle serrulae each with about 18���20 distal fine teeth; the femora and tibia in male almost entirely orange; the shortest distance between eyes below antennal sockets in front view in male about as long as eye height. Taeger (pers. comm.) assumes that the male types of S. atra belong to S. semipicta, and S. rufipes is the male of S. atra. After careful study, we confirm that standpoint. Malaise (1945) recorded 30 rufipes specimens from Kambaiti at 2000 m. Already in the NHRS collection 33 specimens are to be found, and further specimens in other collections. Referring to Malaise's papers, it is obvious that he often estimated or rounded higher numbers of specimens (Taeger, pers. comm.). As there is no indication to exclude specimens with the data given by Malaise, 13 additional specimens of S. rufipes from the BMNH, one from HNHM, one from CSCS, and one from MNCN were labeled as paralectotypes., Published as part of Niu, Gengyun & Wei, Meicai, 2013, Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae), pp. 41-68 in Zootaxa 3746 (1) on pages 45-46, DOI: 10.11646/zootaxa.3746.1.2, http://zenodo.org/record/285307

Niu, Gengyun, Wei, Meicai (2013): Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae). Zootaxa 3746 (1): 41-68, DOI: http://dx.doi.org/10.11646/zootaxa.3746.1.2

Siobla rufopropodea Wei, 1998 (Figs 3 c, 3 d, 5g, 5 h, 9 d, 9 q, 10m, 10 n) Siobla rufopropodea Wei, 1998: 143. ♂, type locality: Luanchuan, Henan (China). Description. ♀. Length, 12 mm (Figs 3 c, 5g). Head black, labrum, palpi, posterior of mandible and antenna yellowish brown, apex of mandible dark reddish brown, prothorax and mesothorax black, posterior margin of pronotum (greatest width 1.5 �� length of lateral ocellus), tegula, posterior ridge of mesoparapsides and posterior margin of posttergite yellowish brown; metathorax yellowish brown, metascutellum black; abdomen black, 1 st to 4 th abdominal tergites, most of 5 th abdominal tergite, lateral edges of 8 th abdominal tergite, 9 th to 10 th abdominal tergites, 1 st to 5 th sternites and apical of sheath yellowish brown; legs yellowish brown, base of fore coxa, and apical 0.25 of hind femur black; body hairs yellowish brown; wings yellowish hyaline, pterostigma yellowish brown. Head densely and coarsely punctured, without any interspaces, mat; labrum shiny with minute and sparse punctures, clypeus punctured with shiny interspaces; prescutum and scutum densely punctured, interspaces narrow, microsculptured; mesoscutellum densely punctured without interspaces; middle of posttergite shiny, microsculptured laterally; parapsides with large and sparse punctures and dense microsculpture; metapostnotum shiny; middle and upper of mesepisternum densely and coarsely punctured, venter with large and shiny interspaces between small and sparse punctures; anepimeron coarsely punctured, katepimeron microsculptured, posterior shiny; metepisternum with anterior and posterior margin shiny; upper of metepisternum shallowly and densely punctured, venter sparsely punctured, shiny; basin and posterior corner of metepimeron shiny, dorsum with large punctures. Abdomen microsculptured, shiny, lateral edges of 1 st abdominal tergite and 6 th to 8 th abdominal tergites punctured. Hairs on dorsum of head straight, as long as diameter of lateral ocelli. Clypeus flat, anterior margin truncate; malar space as long as transverse diameter of median ocellus; shortest distance between eyes below antennal sockets in front view 1.1 �� as long as eye height; middle fovea broad and deep; interocellar furrow narrow and deep, postocellar furrow broad and shallow; postocellar area 1.7 �� as broad as long, obviously lower than top of ocelli; lateral furrows deep, curved, obviously divergent posteriorly; head behind eyes 0.8 �� length of eye in dorsal view; curved laterally and then convergent posteriorly; genal carina sharp, complete. Antenna as long as head, thorax, and 1 st abdominal tergite together, obviously short than vein C, 2 nd antennomere 1.2 �� as long as broad, relative length of 3 rd to 4 th antennomeres as 1.5, 5 th to 8 th antennomeres weakly compressed, not enlarged, inner and outer sides of 6 th to 8 th antennomeres with weak furrows, 7 th antennomere about 2 �� as long as broad. Middle furrow of prescutum distinct; mesoscutellum elevated, slightly higher than scutum. Metabasitarsus 5.5 �� as long as broad, about as long as following 3 tarsomeres combined, hind inner tibial spur 0.5 �� length of metabasitarsus. Pulvilli of hind tarsus weakly developed, 1 st pulvillus 0.5 �� basal breadth and 0.3 �� apical breadth of metabasitarsus, distance between basal two pulvilli 5 �� length of 2 nd pulvillus. Ovipositor sheath as long as middle tibia, apical sheath 1.3 �� as long as basal sheath. Lancet with 16 serrulae, (Fig. 9 d), middle serrulae each with 5���6 distal fine teeth (Fig. 9 q). Male. Length, 9.5 mm (Figs 3 d, 5 h). Similar to female in structure and color, but base of 1 st antennomere black; greatest width of pale posterior margin of pronotum 0.9 �� length of lateral ocellus; extreme anterior line of metapleuron black, 1 st to 5 th abdominal tergites and sternites yellowish brown. Hairs on dorsum of head 1.1 �� as long as diameter of lateral ocelli; shortest distance between eyes below antennal sockets in front view as long as eye height; postocellar area 2 �� as broad as long, malar space 0.5 �� transverse diameter of median ocellus; head in front view with lower interocular distance 0.9 �� eye height; head behind eyes 0.4 �� eyes in length in dorsal view, curved in middle and narrowing posteriorly. Genitalia as in Figs 10 m, 10 n. Variation. The anterior half of metapleuron of one male and entire 1 st antennomere black. Specimens examined. 4 ♀, 16 ♂. Henan: 1 ♂, ���Luanchuan, 1996 -VII- 13, Wei Meicai���; [red] ��� Holotype ���; ���CSCSHT 00810025��� (CSCS); 2 ♂, Luanchuan, 1996 -VII- 13, Wei Meicai leg.; [yellow] paratype; Hubei: 1 ♂, Guitouwan, Shennongjia, E. 110 ��08.872��, N. 31 �� 28.439��, 2150m, 2010 -V- 24, Li Zejian leg.; 2 ♂, CSCS 11021, Guitouwan, Shennongjia, Yichang, E. 110 ��08.872���, N. 31 �� 28.439 ���, 2150m, 2011. V. 19, Li Zejian leg.; 1 ♀ 5 ♂, CSCS 11022, Guitouwan, Shennongjia, Yichang, E. 110 ��08.872���, N. 31 �� 28.439 ���, 2150m, 2011. V. 25���28, Li Zejian leg.; 1 ♀, CSCS 11021, Guitouwan, Shennongjia, Yichang, E. 110 ��08.872���, N. 31 �� 28.439 ���, 2150m, 2011. V. 19, Li Zejian leg.; CSCS-Hym-M00681; 1 ♀ 4 ♂, Guitouwan 2150m, N 31 �� 28��, E. 110 ��09��, Shennongjia, 28.V. 2011, A. Shinohara leg.(NSMT); Shaanxi: 1 ♀, Kaitianguan 2000m, 34 00N 107 51 E, Mt. Taibaishan, Qinling Mts., 6. VI. 2007, A. Shinohara; NSMT-HYM, 62705 (NSMT); 1 ♂, nr. ropeway 2700m, Kaitianguan Tanguy, Mt. Taibaishan, Qinling Mts., 4. VI. 2007, A. Shinohara; NSMT-HYM, 62706 (NSMT). Distribution. China (Henan, Hubei, Shaanxi). Remarks. The species differs from all other species of the Siobla formosana group in the metanotum and metapleuron largely, the first abdominal tergite entirely shining yellow brown; wings hyaline with distinct yellow tinge, venation largely yellow brown; the hind coxa and hind tibia entirely yellow brown; and the malar space as long as diameter of the lateral ocellus. The female of the species is described for the first time., Published as part of Niu, Gengyun & Wei, Meicai, 2013, Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae), pp. 41-68 in Zootaxa 3746 (1) on pages 52-53, DOI: 10.11646/zootaxa.3746.1.2, http://zenodo.org/record/285307

Siobla carinoclypea Niu &Wei, 2010 (Figs 1 c, 1 d, 4 c, 4 d, 7 a–c, 7 r, 7 o, 9 e, 9 r, 10 o, 10 p) Siobla carinoclypea Niu & Wei, 2010 b: 913. ♀, ♂, type locality: Mt. Huping, Hunan (China). Description. Holotype: ♀. Length, 12 mm (Figs 1 c, 4 c). Body and legs black; narrow posterior margin of pronotum dark brown, 2 nd abdominal tergite with small white spots near spiracles; anterior of fore femur and most of tibia and of tarsus brown, middle tarsus blackish brown, hind trochanter yellowish brown. Body hairs silver, hairs on dorsal side of head pale brown. Wings hyaline, pterostigma and veins deep brown or blackish brown, basal 0.2 of pterostigma pale brown. Punctures on head dense and coarse without shiny interspaces, labrum sparsely punctured; prescutum, scutum, and anterior slope of mesoscutellum densely punctured, with narrow interspaces; lateral parts of mesoscutellum sparsely punctured, interspaces shiny; posterior slope of mesoscutellum densely punctured, without interspaces; posttergite shiny with microsculptured laterally; parapsides with large and sparse punctures and dense microsculpture; metascutellum with anterior slope shiny and posterior slop densely punctured; metapostnotum shiny with margins slightly microsculptured; middle and upper of mesepisternum densely and coarsely punctured, without shiny interspaces, venter with large and shiny interspaces between small and sparse punctures; anepimeron coarsely punctured, upper of katepimeron with large punctures, interspaces obviously longer than diameter of punctures, middle and lower of katepimeron microsculptured; upper of metepisternum with dense and small punctures, interspaces shiny, narrower than diameter of punctures, venter sparsely punctured, shiny; basin of metepimeron microsculptured, dorsum with large punctures and posterior corner shiny. First abdominal tergite microsculptured, shiny, lateral edges slightly punctured; other tergites obviously microsculptured, mat, lateral edges of both 3 rd and 4 th tergites, posterior of 4 th tergite, and most of 5 th to 8 th tergites obviously punctured. Hairs on dorsum of head straight, about as long as diameter of lateral ocelli. Clypeus shallowly emarginate (Fig. 7 r), middle of clypeus with a transverse carina (Fig. 7 o); malar space as long as transverse diameter of median ocellus; shortest distance between eyes below antennal sockets in front view 1.1 × eye height; middle fovea shallow and broad; interocellar furrow deep, postocellar furrow broad and shallow; postocellar area 1.2 × as broad as long, distinctly elevated and about as high as top of ocelli, with an obscure middle ridge; lateral furrows straight, parallel and gradually shallow posteriorly; head behind eyes nearly as long as eye in dorsal view; curved laterally and then convergent posteriorly; genal carina developed, complete (Fig. 7 a). Antenna as long as head, thorax and basal two abdominal tergites together, as long as vein C, 2 nd antennomere 1.5 × as long as broad, relative length of 3 rd to 4 th antennomeres as 1.8, 5 th to 8 th antennomeres weakly compressed, inner and outer sides of 5 th to 8 th antennomeres with weak furrows, 7 th antennomere 2.3 × as long as broad. Middle furrow of prescutum distinct; mesoscutellum distinctly elevated, obviously higher than scutum. Metabasitarsus 4.7 × as long as broad, nearly as long as following 4 tarsomeres combined and 1.4 × length of following 3 tarsomeres combined, hind inner tibial spur 0.45 × length of metabasitarsus. Pulvilli of hind tarsus small. Ovipositor sheath 1.1 × length of middle tibia in lateral view; exposed part of apical sheath about 2 × as long as broad, apical sheath 1.4 × as long as basal sheath, apical sheath narrow in dorsal view, exposed part 4 × length of cerci, lateral setae on both sides of sheath straight, forming an angle about 50 degrees. Lancet broadened before apex, with 18 serrulae (Fig. 9 e), middle serrulae each with 14–15 distal fine teeth (Fig. 9 r). Male. Length, 10.5 mm (Figs 1 d, 4 d). Body black; 2 nd to 4 th abdominal tergites, 2 nd to 5 th abdominal sternites, and most of 6 th and 7 th sternites yellowish brown; legs yellowish brown, each coxa, fore and middle trochanter, dorsum of base of fore femur, stripe on dorsum of middle femur, apical 0.2 and stripe on dorsum of hind femur, and apical 0.3 of hind tibia black; similar to female in structure, but malar space 0.6 × transverse diameter of median ocellus (Fig. 7 c); shortest distance between eyes below antennal sockets in front view 0.9 × eye height; head behind eyes 0.5 ×eyes in dorsal view, curved laterally and then distinctly convergent posteriorly; hairs on temple as long as diameter of lateral ocelli (Fig. 7 b); genitalia as in Figs 10 o, 10 p. Variation. Females 10.5–12 mm, males 10.5–11.5mm. Specimens examined. 7 ♀, 11 ♂. Hunan: 1 ♀, “Jiangping, Hupingshan, 2004 -VI- 9, 1200–1600m, Zhou Hu”; [red] ” Holotype ”; “CSCSHT 00810020” (CSCS); 1 ♂, Jiangping, Hupingshan, 2004 -VI- 9, 1200–1600m, Zhou Hu leg.; [yellow] paratype; 2 ♀ 8 ♂, Jiangping, Hupingshan, 2004 -VI- 1, 1200–1600m, Zhou Hu leg.; [yellow] paratype; 1 ♂, Jiangping, Hupingshan, 2004 -VI- 1, 1200–1600m, Zhou Hu leg.; 1 ♂, Huangsang, Suining, 2005 -IV- 21, 600– 900m, Zhou Hu leg.; Guizhou: 1 ♀, Linchang, Leigongshan, 2005 -VI- 1, 1600m, Xiao Wei Leg.; [yellow] paratype; Shaanxi: 1 ♀, Yuantou, Jialingjiang, E. 106 º 59.026´, N. 34 º 13.177´, 1570m, 2007 -V- 26, Jiang Xiaoyu leg.; [yellow] paratype; Guangxi: 1 ♀, Tingcheping, Cenwanglaoshan, N 27 º 12´16 ´´, E. 120 º 13´41 ´´, 1875m, 2012. IV. 29, Wei Meicai, Niu Gengyun; Hubei: 1 ♀, Guitouwan 2150m, N 31 º 28´, E. 110 º09´, Shennongjia, 28.V. 2011, A. Shinohara leg.(NSMT). Distribution. China (Hunan, Guizhou, Shaanxi, Hubei, Guangxi). Remarks. The new species is similar to S. listoni and S. parallela but differs from them in clypeus with an obtuse transverse ridge and a small apical incision; the posterior margin of pronotum, tegula, the middle trochanter, the hind femur, tibia and tarsus black (the posterior of pronotum at most very narrowly margined with brownish line); the abdominal tergites and sternites black, only second tergite with a small white spot near spiracle; the hind femur in male with black macula but at least basal 4 / 5 yellow brown in outer side. The new species differs from S. listoni also in the forewings hyaline; antenna slender and the antennomeres 5–7 not dilated; and the apex of mesoscutellum round. The new species differs from S. parallela also in the pterostigma black brown; the apex of apical sheath not acute in lateral view; the posttergite with a middle carina; the antennomeres 2–3 and posttergite without pale spot. See the remarks under S. atra for the difference between S. carinoclypea and S. atra.

Siobla sturmii (Klug, 1817) (Figs 3 a���b, 4 j���l, 5 q���r, 6 h, 7 h, 8 h, q, 10g, 12 f���i) Tenthredo sturmii Klug, 1817: 116. Macrophya pacifica Smith, 1874: 378, syn. nov. Siobla pacifica: Nakagawa, 1899: 204; Takeuchi, 1936: 86 (in part);?Abe and Togashi, 1989: 555; Lelej &Taeger, 2007: 958; Taeger et al., 2010: 585; Lelej, 2012: 100. ��� Siobla ruficornis Cameron?���: Takeuchi, 1919 b: 13 (in part). Siobla sturmii var. pacifica: Malaise, 1931 a: 121; Malaise, 1934: 24, 26. Siobla sturmii: Takeuchi, 1931: 31;?Naito, 1982: 574; Takeuchi, 1956: 77 (in part); Lee et al., 2000: 119; Katayama, 2004: 96 (in part); Naito et al., 2004: 57 (in part); Lee et al., 2010: 165; Taeger et al., 2010: 585; Niu & Wei, 2010: 50. Siobla ruficornis: Takeuchi, 1936: 85 (in part);?Togashi, 1965: 246; Naito, 1982: 574; Abe and Togashi, 1989: 555 (in part); Yoshida, 2006: 91;?Togashi, 2008: 490. Not Cameron (1876). Siobla sturmii sturmii: Zhelochovtsev & Zinovjev, 1996: 368. Siobla sturmii subsp. pacifica: Zhelochovtsev & Zinovjev, 1996: 368. [Only East Asian references are given.] Distribution. Japan (Hokkaido, southern Kuriles, Honshu, Kyushu), Korea, Russian Far East (Primorskiy kray; Sakhalin), Siberia, Europe. New records from Kyushu and Sakhalin. Primary type examined. Lectotype of Macrophya pacifica hereby designated (Fig. 4 j���l): ♀, ��� Type H. T.��� ���B. M. TYPE HYM. 1.246 ��� ��� Macrophya pacifica, Type, Smith��� ���Kb.��� ���t. 9 / 17.��� ���BMNH(E) # 651765 ��� (BMNH). Other material examined (see Appendix). 165 specimens (57 ♀ and 108 ♂). Variation. The length varies from 9.5 to 12.5 mm in the female and 9.5 to 12.0 mm in the male. The Japanese specimens examined are quite variable in colour pattern, those from Hokkaido usually being darker than those from Honshu and Kyushu. In the female, the clypeus, the metapleuron, and the hind coxa, trochanter and trochantellus are usually entirely black in the Hokkaido specimens, whereas ventral half of the clypeus and the other parts given above are frequently marked with reddish brown (Fig. 12 h, i) in the Honshu and Kyushu specimens, particularly in the specimens from southwestern Honshu. The abdomen in the Hokkaido specimens has only the tergum 2 and part of the tergum 3 reddish brown, while the abdomen in the Honshu specimens has the terga 2 and 3 mostly reddish brown and sometimes the posterior margin of the tergum 1 and the anterior margin of the tergum 4 are also reddish brown. The abdomen in the Kyushu specimens has the terga 2 to 4 almost entirely reddish brown. The male shows some colour variation almost only in the extent of reddish brown areas on the abdomen. The Hokkaido and northern Honshu specimens tend to be darker but variable; the darkest specimens have only parts of the terga 2 and 3 narrowly reddish brown, whereas the lightest specimens have the terga 2 to 4 almost entirely reddish brown. The males from southwestern Honshu and Kyushu almost constantly have the terga 2 to 5 (and often anterior part of 6) reddish brown. Remarks. This species belongs to the S. sturmii group (Niu & Wei 2010). The Japanese specimens are variable in colouration as discussed above. They are usually similar to normal European specimens, such as Klug���s lectotype (see photographs in Taeger & Blank 2011), but some specimens resemble S. ruficornis (Fig. 12 a ���e). Siobla sturmii can be distinguished from S. ruficornis by the shape of the head in dorsal view (Fig. 12 f) and that of the sawsheath in lateral view (Fig. 12 g). The features of the lancet, harpe and penis valve are also different. Among the Japanese congeners, S. sturmii looks similar to S. jucunda and S. japonica, but S. sturmii is separated from these species by the colour of the antenna and hind femur. In S. sturmii, the female antenna is entirely or almost entirely reddish brown, the male antenna has the scape, pedicel, and base of the flagellomere 1 black and the flagellomeres 2 to 9 entirely reddish brown, and the hind femur is basally reddish brown. In S. jucunda, the basal three antennomeres are mostly black in the female, the basal four antennomeres are mostly black in the male, and the hind femur is entirely black. In S. japonica, the antenna and the hind femur are entirely or almost entirely black. Smith (1874) described the female and male of Macrophya pacifica from North Japan, but Kirby (1882) was not able to locate any male specimens in the London collection and listed one female specimen as the type. Here we designate the female specimen as lectotype. An examination of the lectotype has revealed that it is conspecific with S. sturmii and we hereby propose to synonymize S. pacifica with S. sturmii. As discussed elsewhere, ��� Siobla ruficornis Cameron?��� of Takeuchi (1919 b) was a mixture of S. takeuchii and most probably the male of S. sturmii. Takeuchi (1956) listed four specimens from Maruyama, Kunashiri Island. This series of specimens kept in HU, Sapporo, comprises three specimens of S. sturmii and one specimen of S. jucunda (see Appendix). Mainly in local faunal lists, Togashi (1961, 1971, 1978, 1998b, 2002 b), Okutani (1972, 1974b), Togashi et al. (1980), Otsuka (1984), Nambu (1992, 1998), and Katayama et al. (2007) included ��� S. sturmii ��� and Nambu (1992, 1998) and Naito et al. (2004) referred to ��� S. ruficornis ���. These references may actually include S. japonica, S. jucunda, or S. sturmii. Examination of their original material is necessary to establish its true identity., Published as part of Shinohara, Akihiko, Wei, Meicai & Niu, Gengyun, 2013, Revision of Siobla (Hymenoptera, Tenthredinidae) from Japan, pp. 1-40 in Zootaxa 3746 (1) on pages 21-22, DOI: 10.11646/zootaxa.3746.1.1, http://zenodo.org/record/285294

Shinohara, Akihiko, Wei, Meicai, Niu, Gengyun (2013): Revision of Siobla (Hymenoptera, Tenthredinidae) from Japan. Zootaxa 3746 (1): 1-40, DOI: http://dx.doi.org/10.11646/zootaxa.3746.1.1

Siobla semipicta Malaise, 1945 (Figs 3 e, 3 f, 5 i, 5 j, 7 l, 7m, 9 i, 9 v, 10 s, 10t) Siobla semipicta Malaise, 1945: 124. ♀, type locality: Kambaiti, (Myanmar). Siobla semipicta fasciata Malaise, 1945: 124. ♀, type locality: Kambaiti, (Myanmar). Syn. nov. Siobla atra: Malaise, 1945: 123. ♂, misidentification. Description. Lectotype: ♀. Length, 10 mm (Figs 3 e, 5 i). Black, palpi, posterior margin of pronotum (greatest width 0.4 �� length of lateral ocellus), and obscure spots on lateral edges of 2 nd abdominal tergite dirty brown, 10 th tergite and cerci pale brown; legs orange brown, coxae except apical margin, fore and middle 1 st trochanter, apical half of hind femur and apical 0.3 of hind tibia black; body hairs silver; wings hyaline, pterostigma blackish brown. Head densely punctured, clypeus, inner orbit, and postocellar area with visible interspaces, narrower than diameter of punctures, microsculptured; punctures on labrum minute and sparse; mesonotum densely punctured, with interspaces on posterior of prescutum, and middle of scutum as long as or slightly broader than diameter of punctures, without microsculpture, shiny; interspaces on middle of anterior slope of mesoscutellum 2 �� diameter of punctures, shiny, other area of mesoscutellum densely punctured without interspaces; posttergite shiny; parapsides with large and sparse punctures and dense microsculpture; metascutellum with anterior slope shiny and posterior slope densely punctured; metapostnotum and parapsides microsculptured; middle and upper of mesepisternum densely and coarsely punctured, venter with large and shiny interspaces between small and sparse punctures; anterior corner of venter with a small shiny patch; anepimeron coarsely punctured, katepimeron microsculptured, posterior shiny; metepisternum with upper densely punctured, mat, and venter sparsely punctured, shiny; basin of metepimeron shiny, margin of basin microsculptured, dorsum of metepimeron with large punctures and posterior corner shiny. abdominal tergites microsculptured, middle and lateral edges of 1 st abdominal tergite densely punctured (Fig 7 l), 3 rd to 8 th tergites punctured. Hairs on dorsum of head straight, as long as diameter of lateral ocellus, Hairs on mesopleuron 0.9 �� as long as diameter of lateral ocellus. Clypeus flat, anterior margin truncate; malar space as long as transverse diameter of median ocellus; shortest distance between eyes below antennal sockets in front view 1.1 �� as long as eye height; middle fovea broad and shallow; interocellar furrow narrow and deep, postocellar furrow broad and shallow; postocellar area 1.6 �� as broad as long, obviously lower than top of ocelli, middle weakly elevated; lateral furrows deep, nearly straight, divergent posteriorly; head behind eyes 0.8 �� length of eye in dorsal view; curved laterally and then convergent posteriorly; genal carina sharp, complete. Antenna shorter than head, thorax and 1 st and 2 nd tergites together, as long as vein C, 2 nd antennomere 1.3 �� as long as broad, relative length of 3 rd to 4 th antennomeres as 1.6, 5 th to 8 th antennomeres hardly compressed, not enlarged, inner and outer sides of 6 th to 8 th antennomeres with weak furrows, 7 th antennomere about 2.2 �� as long as broad. Middle furrow of prescutum shallow; mesoscutellum obviously elevated, hardly higher than scutum. Metabasitarsus 6.5 �� as long as broad, about 1.3 �� length of following 3 tarsomeres combined, hind inner tibial spur 0.5 �� length of metabasitarsus. Pulvilli of hind tarsus weakly developed, 1 st pulvillus 0.7 �� basal breadth and 0.5 �� apical breadth of metabasitarsus, distance between basal two pulvilli 2.6 �� length of 2 nd pulvillus. Ovipositor sheath as long as middle tibia, exposed part of apical sheath about 1.3 �� as long as broad; apical sheath 1.4 �� length of basal sheath. Lancet with 16 serrulae (Fig. 9 i), middle serrulae each with 8���10 distal fine teeth (Fig. 9 v). Male. Length, 10.5 mm (Figs 3 f, 5 j). Similar to female in structure and color, but pronotum and legs black, fore and middle femora black with anterior of apical 0.3 to 0.4 orange brown, 2 nd and 3 rd abdominal sternites yellowish white; malar space linear, 0.2 �� transverse diameter of median ocellus (Fig. 7 m); head in front view with lower interocular distance 0.8 �� eye height; head behind eyes 0.5 �� eyes in length in dorsal view, curved in middle and narrowing posteriorly. Genitalia as in Figs 10 s, 10t. Variation. The second abdominal tergite of one female is largely reddish brown. This was described as S. semipicta fasciata Malaise, 1945. Specimens examined. MYANMAR: Lectotype of Siobla semipicta (hereby designated) ♀, ��� N. E. BURMA, Kambaiti, 7000ft, 29 / 5, R. MALAISE���; ��� Typus ���; ��� Siobla semipicta Malaise det. 1936 ���;���NHRS-HYME 000005296���; ��� Lectotype, Siobla semipicta Malaise, 1945, Design Niu G. 2012 ��� (NRHS). Paralectotypes of Siobla semipicta: (Taeger, pers. comm..): In the NRHS were four females under S. semipicta. One of these specimens fits the description of S. semipicta fasciata and is considered to be the holotype of the latter nominal taxon. The remaining 3 syntypes show a distinct variability regarding the abdominal coloration as noted by Malaise. 1 ♀, same data as the lectotype but 10 / 6, NHRS-HEVA 000001553; 1 ♀, same data as the lectotype but 9 / 6, NHRS-HEVA 0 0 0 0 0 1554 (both NHRS); 1 ♀, same data as the lectotype but 29 / 5 (BMNH). Holotype of Siobla semipicta fasciata: ♀, "N.E. BURMA Kambaiti 7000 ft. 15 / 5 R. Malaise"; " Paratypus "; " Siobla semipicta n. sp. Malaise det. 1936 "; "NHRS-HEVA 000001552". Paralectotypes of Siobla atra: 1 ♂, N. E. BURMA, Kambaiti, 7000ft, 24 / 5, R. MALAISE (HYME-NHRS 000005305); 1 ♂ same data but 8 / 5; 1 ♂ same data but 14 / 5; 4 ♂ same data but 15 / 5; 3 ♂ same data but 16 / 5; 4 ♂ same data but 17 / 5; 2 ♂ same data but 29 / 5; 1 ♂ same data but 1 / 6; 1 ♂ same data but 4 / 6; 1 ♂ same data but 5 / 6; 1 ♂ same data but 9 / 6; 1 ♂ same data but 10 / 6 (NHRS-HEVA 000001608- NHRS-HEVA 000001629). Other specimens: 21 #♂m from Kambaiti, leg. Malaise (details not recorded). CHINA: Yunnan: 7 ♀ 53 ♂, Xiaoheishan, Longling, Yunnan, E. 98 �� 45.574 ���, N. 24 �� 41.713 ���, 2009 -VI- 2, Wei Meicai, Niu Gengyun, Li Zejian; 4 ♂, Yaojiaping, Lushui, Yunnan, E. 98.710 ��, N. 25.975 ��, 2550m, 2009 -VI- 6, Xiao Wei, Zhong Yihai; 4 ♂, Gaoligongshan, Longling, Yunnan, E. 98 �� 46.062 ���, N. 24 �� 49.700 ���, 2145m, 2009 -VI- 2, Zhong Yihai; 4 ♂, Houqiaozhen, Tengchong, Yunnan, E. 98 �� 12.665 ���, N. 25 �� 21.990 ���, 2196m, 2009 -VI- 1, Zhong Yihai, Xiao Wei. Distribution. China (Yunnan); Myanmar. Remarks. The female of Siobla semipicta Malaise, 1945 is slightly similar to the female of S. atra Malaise, 1945, and therefore Malaise misidentified the males of semipicta as atra. See the key above for the differences between the two species. S. semipicta is also quite similar to S. punctata (Cameron, 1899) of Siobla grahami group in body color but differs from it in the dense punctures on the dorsal side of mesoscutellum., Published as part of Niu, Gengyun & Wei, Meicai, 2013, Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae), pp. 41-68 in Zootaxa 3746 (1) on pages 53-54, DOI: 10.11646/zootaxa.3746.1.2, http://zenodo.org/record/285307

Siobla carinoclypea Niu &Wei, 2010 (Figs 1 c, 1 d, 4 c, 4 d, 7 a���c, 7 r, 7 o, 9 e, 9 r, 10 o, 10 p) Siobla carinoclypea Niu & Wei, 2010 b: 913. ♀, ♂, type locality: Mt. Huping, Hunan (China). Description. Holotype: ♀. Length, 12 mm (Figs 1 c, 4 c). Body and legs black; narrow posterior margin of pronotum dark brown, 2 nd abdominal tergite with small white spots near spiracles; anterior of fore femur and most of tibia and of tarsus brown, middle tarsus blackish brown, hind trochanter yellowish brown. Body hairs silver, hairs on dorsal side of head pale brown. Wings hyaline, pterostigma and veins deep brown or blackish brown, basal 0.2 of pterostigma pale brown. Punctures on head dense and coarse without shiny interspaces, labrum sparsely punctured; prescutum, scutum, and anterior slope of mesoscutellum densely punctured, with narrow interspaces; lateral parts of mesoscutellum sparsely punctured, interspaces shiny; posterior slope of mesoscutellum densely punctured, without interspaces; posttergite shiny with microsculptured laterally; parapsides with large and sparse punctures and dense microsculpture; metascutellum with anterior slope shiny and posterior slop densely punctured; metapostnotum shiny with margins slightly microsculptured; middle and upper of mesepisternum densely and coarsely punctured, without shiny interspaces, venter with large and shiny interspaces between small and sparse punctures; anepimeron coarsely punctured, upper of katepimeron with large punctures, interspaces obviously longer than diameter of punctures, middle and lower of katepimeron microsculptured; upper of metepisternum with dense and small punctures, interspaces shiny, narrower than diameter of punctures, venter sparsely punctured, shiny; basin of metepimeron microsculptured, dorsum with large punctures and posterior corner shiny. First abdominal tergite microsculptured, shiny, lateral edges slightly punctured; other tergites obviously microsculptured, mat, lateral edges of both 3 rd and 4 th tergites, posterior of 4 th tergite, and most of 5 th to 8 th tergites obviously punctured. Hairs on dorsum of head straight, about as long as diameter of lateral ocelli. Clypeus shallowly emarginate (Fig. 7 r), middle of clypeus with a transverse carina (Fig. 7 o); malar space as long as transverse diameter of median ocellus; shortest distance between eyes below antennal sockets in front view 1.1 �� eye height; middle fovea shallow and broad; interocellar furrow deep, postocellar furrow broad and shallow; postocellar area 1.2 �� as broad as long, distinctly elevated and about as high as top of ocelli, with an obscure middle ridge; lateral furrows straight, parallel and gradually shallow posteriorly; head behind eyes nearly as long as eye in dorsal view; curved laterally and then convergent posteriorly; genal carina developed, complete (Fig. 7 a). Antenna as long as head, thorax and basal two abdominal tergites together, as long as vein C, 2 nd antennomere 1.5 �� as long as broad, relative length of 3 rd to 4 th antennomeres as 1.8, 5 th to 8 th antennomeres weakly compressed, inner and outer sides of 5 th to 8 th antennomeres with weak furrows, 7 th antennomere 2.3 �� as long as broad. Middle furrow of prescutum distinct; mesoscutellum distinctly elevated, obviously higher than scutum. Metabasitarsus 4.7 �� as long as broad, nearly as long as following 4 tarsomeres combined and 1.4 �� length of following 3 tarsomeres combined, hind inner tibial spur 0.45 �� length of metabasitarsus. Pulvilli of hind tarsus small. Ovipositor sheath 1.1 �� length of middle tibia in lateral view; exposed part of apical sheath about 2 �� as long as broad, apical sheath 1.4 �� as long as basal sheath, apical sheath narrow in dorsal view, exposed part 4 �� length of cerci, lateral setae on both sides of sheath straight, forming an angle about 50 degrees. Lancet broadened before apex, with 18 serrulae (Fig. 9 e), middle serrulae each with 14���15 distal fine teeth (Fig. 9 r). Male. Length, 10.5 mm (Figs 1 d, 4 d). Body black; 2 nd to 4 th abdominal tergites, 2 nd to 5 th abdominal sternites, and most of 6 th and 7 th sternites yellowish brown; legs yellowish brown, each coxa, fore and middle trochanter, dorsum of base of fore femur, stripe on dorsum of middle femur, apical 0.2 and stripe on dorsum of hind femur, and apical 0.3 of hind tibia black; similar to female in structure, but malar space 0.6 �� transverse diameter of median ocellus (Fig. 7 c); shortest distance between eyes below antennal sockets in front view 0.9 �� eye height; head behind eyes 0.5 ��eyes in dorsal view, curved laterally and then distinctly convergent posteriorly; hairs on temple as long as diameter of lateral ocelli (Fig. 7 b); genitalia as in Figs 10 o, 10 p. Variation. Females 10.5���12 mm, males 10.5���11.5mm. Specimens examined. 7 ♀, 11 ♂. Hunan: 1 ♀, ���Jiangping, Hupingshan, 2004 -VI- 9, 1200���1600m, Zhou Hu���; [red] ��� Holotype ���; ���CSCSHT 00810020��� (CSCS); 1 ♂, Jiangping, Hupingshan, 2004 -VI- 9, 1200���1600m, Zhou Hu leg.; [yellow] paratype; 2 ♀ 8 ♂, Jiangping, Hupingshan, 2004 -VI- 1, 1200���1600m, Zhou Hu leg.; [yellow] paratype; 1 ♂, Jiangping, Hupingshan, 2004 -VI- 1, 1200���1600m, Zhou Hu leg.; 1 ♂, Huangsang, Suining, 2005 -IV- 21, 600��� 900m, Zhou Hu leg.; Guizhou: 1 ♀, Linchang, Leigongshan, 2005 -VI- 1, 1600m, Xiao Wei Leg.; [yellow] paratype; Shaanxi: 1 ♀, Yuantou, Jialingjiang, E. 106 �� 59.026��, N. 34 �� 13.177��, 1570m, 2007 -V- 26, Jiang Xiaoyu leg.; [yellow] paratype; Guangxi: 1 ♀, Tingcheping, Cenwanglaoshan, N 27 �� 12��16 ����, E. 120 �� 13��41 ����, 1875m, 2012. IV. 29, Wei Meicai, Niu Gengyun; Hubei: 1 ♀, Guitouwan 2150m, N 31 �� 28��, E. 110 ��09��, Shennongjia, 28.V. 2011, A. Shinohara leg.(NSMT). Distribution. China (Hunan, Guizhou, Shaanxi, Hubei, Guangxi). Remarks. The new species is similar to S. listoni and S. parallela but differs from them in clypeus with an obtuse transverse ridge and a small apical incision; the posterior margin of pronotum, tegula, the middle trochanter, the hind femur, tibia and tarsus black (the posterior of pronotum at most very narrowly margined with brownish line); the abdominal tergites and sternites black, only second tergite with a small white spot near spiracle; the hind femur in male with black macula but at least basal 4 / 5 yellow brown in outer side. The new species differs from S. listoni also in the forewings hyaline; antenna slender and the antennomeres 5���7 not dilated; and the apex of mesoscutellum round. The new species differs from S. parallela also in the pterostigma black brown; the apex of apical sheath not acute in lateral view; the posttergite with a middle carina; the antennomeres 2���3 and posttergite without pale spot. See the remarks under S. atra for the difference between S. carinoclypea and S. atra., Published as part of Niu, Gengyun & Wei, Meicai, 2013, Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae), pp. 41-68 in Zootaxa 3746 (1) on pages 47-48, DOI: 10.11646/zootaxa.3746.1.2, http://zenodo.org/record/285307

Siobla formosana Takeuchi, 1927 (Figs 1 e, 1 f, 4 e, 4 f, 8 a���c, 9 k, 9 x, 10g, 10 h) Siobla formosana Takeuchi, 1927: 208. ♀, type locality: Musha, Taiwan (China). Siobla formosana: Niu & Wei, 2011: 162. redescription. Distribution. China (Taiwan). Remarks. This species was redescribed in detail in Niu & Wei (2011). Siobla formosana is very close to Siobla fumipennis Malaise, 1945 but differs from it in the basal part of wings in both sexes hyaline; the pterostigma dark brown; the second abdominal tergite entirely yellow brown and metabasitarsus pale brown in female; the abdominal tergite 4 in male largely yellow brown; and the trochanters usually yellow brown. S. formosana is also similar to S. jiangi and S. zuoae. See the key above for the differences between them., Published as part of Niu, Gengyun & Wei, Meicai, 2013, Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae), pp. 41-68 in Zootaxa 3746 (1) on page 48, DOI: 10.11646/zootaxa.3746.1.2, http://zenodo.org/record/285307

Siobla weiweii Niu & Wei, sp. nov. (Figs 6 a, 6 b, 6 c, 6 d, 9 b, 9 o) Description. Holotype: ♀. Length, 12 mm (Figs 6 a, 6 b). Black, labrum blackish brown, palpi dirty brown, entire 2 nd to 4 th abdominal tergites and middle of 10 th tergite, 2 nd to 5 th sternites, middle longitudinal stripe on 7 th sternites, and cerci pale brown; antenna yellowish brown, most of 1 st and of 2 nd antennomeres and base of 3 rd antennomere black, 8 th and 9 th antennomeres blackish brown; legs yellowish brown, fore and middle coxae, most of hind coxa, basal margin of fore femur, basal 0.2 of middle femur, and apical 5 / 6 of hind femur black, spot on venter of apex of hind tibia dark brown; body hairs silver; wings hyaline, pterostigma brown except extreme base pale brown. Head densely punctured, punctures on labrum fine and sparse; prescutum, scutum and anterior slope of mesoscutellum densely punctured, interspaces narrow; punctures on lateral parts of mesoscutellum obvious and sparse, interspaces shiny; posterior slope of mesoscutellum densely punctured, without interspace; posttergite and metapostnotum shiny with margins slightly microsculptured; mesoparapsides with large and sparse punctures and dense microsculpture; metascutellum with anterior slope shiny and posterior slop densely punctured; metapostnotum shiny with margins slightly microsculptured; middle and upper of mesepisternum densely and coarsely punctured, venter with large and shiny interspaces between small and sparse punctures; anepimeron coarsely punctured, katepimeron microsculptured, posterior shiny; metepisternum with upper densely punctured, interspaces narrow, feebly shiny, and venter sparsely punctured, shiny; basin of metepimeron shiny, dorsum of metepimeron with large punctures, posterior corner shiny. First abdominal tergite shiny with very weak microsculpture, lateral edges hardly punctured; other tergites weakly microsculptured, lateral edges of 4 th tergite and entire of 5 th to 8 th tergites punctured. Hairs on dorsum of head straight, 1.2 × as long as diameter of lateral ocellus. Middle of clypeus with transverse carina, anterior margin truncate; malar space 0.9 × as long as transverse diameter of median ocellus; shortest distance between eyes below antennal sockets in front view as long as eye height; middle fovea broad and shallow; interocellar furrow narrow and deep, postocellar furrow narrow and shallow; postocellar area 1.3 × as broad as long, obviously lower than top of ocelli, middle obviously elevated; lateral furrows deep, nearly straight, divergent posteriorly; head behind eyes 0.8 × length of eye in dorsal view; curved laterally and then convergent posteriorly; genal carina sharp, complete (Fig. 6 c). Antenna as long as head, thorax, and 1 st abdominal tergite together, slightly shorter than vein C, 2 nd antennomere 1.4 × as long as broad, relative length of 3 rd to 4 th antennomeres as 1.8, 5 th to 8 th antennomeres weakly compressed, not enlarged, inner and outer sides of 6 th to 8 th antennomeres with weak furrows, 7 th antennomere about 1.9 × as long as broad. Middle furrow of prescutum deep; mesoscutellum elevated with a peak, higher than scutum (Fig. 6 d). Metabasitarsus 4.7 × as long as broad, about 1.2 × length of following 3 tarsomeres combined, hind inner tibial spur 0.4 × length of metabasitarsus. Pulvilli of hind tarsus weakly developed, 1 st pulvillus 0.7 × basal breadth and 0.4 × apical breadth of metabasitarsus, distance between basal two pulvilli 4 × length of 2 nd pulvillus. Ovipositor sheath 1.2 × length of middle tibia, venter margin curved, exposed part of apical sheath slightly narrower than apex of hind tibia, about 1.6 × as long as broad; apical sheath 1.5 × length of basal sheath. Lancet with 17 serrulae (Fig. 9 b), middle serrulae each with 4 distal fine teeth (Fig. 9 o). Male. Unknown. Etymology: This species is named after the first name of the collector of the holotype. Specimens examined. Chongqing: Holotype: 1 ♀, Dafengbao, Huangshui, Shizhu, Chongqing, 2006 -V- 3, Zhang Weiwei leg.; CSCSHT 0 0 810026 (CSCS). Distribution. China (Chongqing). Remarks. The new species is similar to S. acutitheca Niu & Wei, 2010, but differs from it in tegula and pronotum entirely black; the pterostigma black brown; the base of fore and middle femora with black spot; the dorsal side of the apex of lancet acute; serrulae weakly protruding, the 10 th serrulae with 4 distal teeth; and the first pulvillus of hind tarsus very small.

Siobla takeuchii Shinohara, Wei & Niu, 2013 Siobla takeuchii Shinohara, Wei & Niu: real page��� (1108 Nui). ♀, type locality: Honshu, (Japan)., Published as part of Niu, Gengyun & Wei, Meicai, 2013, Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae), pp. 41-68 in Zootaxa 3746 (1) on page 54, DOI: 10.11646/zootaxa.3746.1.2, http://zenodo.org/record/285307

Siobla takeuchii Shinohara, Wei & Niu, sp. nov. (Figs 3 c���d, 5 s���t, 6 i, 7 i, 8 i, r, 10 c, 11 a���d) urn:lsid:zoobank.org:act: 100 E 981 E- 7 E 8 A- 4 F 4 E- 9 DD0-A 3 D 48853 E0D 8 Description. Female. Holotype (Figs 3 c, 5 s). Length 15.0 mm. Body black, apical 0.65 of antennomere 2, antennomeres 3 and 4 entirely, antennomere 5 except extreme apex, palpi, narrow posterior margin of pronotum, narrow anterior margin of tegula, spots on both lateral sides of abdominal tergum 1, abdominal tergum 9 largely, abdominal tergum 10, apex of sheath, and cerci pale brown; abdominal terga 2 and 3 and sterna yellow brown. Legs black, apical 0.25 of fore femur, anterior stripe and apical 0.2 of middle femur, fore and middle tibiae and tarsi, hind trochanter, basal 0.25 of hind femur, basal 0.65 of hind tibia, and entire hind tarsus yellow brown. Hairs on dorsum of head and of thorax yellow brown, hairs on pleuron silver. Fore wings yellowish hyaline, vein C and stigma pale brown, cell Sc infuscated; hind wings hyaline. Dorsum of head densely punctured, frons coarsely punctured, matt; upper part of inner orbit, temple and postocellar area densely punctured without interspaces, feebly shiny. Most of propleuron sparsely punctured; mesonotum sparsely punctured, with narrow shiny interspaces; mesoscutellum densely punctured without interspaces, feebly shiny, with posterior slope coarsely punctured, matt; mesoscutellar appendage shiny at middle; punctures on middle of mesepisternum large and dense, without interspaces, punctures on anterior and posterior margins of mesepisternum sparse, punctures on ventral part of mesepisternum shallow and sparse, interspaces feebly microsculptured, shiny; part of katepimeron densely microsculptured but anterior and posterior margins shiny; metapostnotum microsculptured; metepisternum with dorsal part densely punctured and ventral part of shallowly and sparsely punctured; most of metepimeron microsculptured, rest shiny except dorsal part densely punctured. Abdominal tergum 1 with a few shallow punctures along anterior margin and on lateral sides, shiny; terga 2 and 3 shiny, terga 4 to 9 faintly microsculptured, with lateral sides shallowly punctured. Hairs on dorsum of head and of thorax slightly longer than diameter of median ocellus, straight; hairs on mesopleuron slightly longer than diameter of median ocellus, hardly curved. Clypeus 2.0 �� as broad as long, anterior margin weakly protruding; malar space 0.6 �� diameter of median ocellus; lower interocular distance 1.1 �� eye height (Fig. 11 c); middle fovea small, about as deep as lateral fovea; interocellar furrow wide and deep; postocellar furrows shallow; postocellar area weakly elevated, as high as ocelli, with fine and low middle carina, about 1.4 �� as broad as long; lateral furrows deep, slightly curved in middle, nearly parallel posteriorly; temple 0.6 �� length of eye in dorsal view, anterior 0.25 curved, rest divergent backwards (Fig. 11 a). Antenna about 0.9 �� as long as vein C of forewing, antennomere 2 longer than broad, antennomere 3 1.7 �� as long as antennomere 4, subapical antennomeres weakly enlarged; antennomere 7 2.0 �� as long as broad. Middle furrow on pronotum narrow; mesoscutellum roundly elevated, slightly higher than top of mesoscutum, without carina or acute apex; mesoscutellar appendage almost flat, without middle carina; mesepisternum obviously elevated, ventral spur absent. Hind inner tibial spur 0.5 �� length of metabasitarsus; metabasitarsus 5 �� as long as broad, 1.1 �� length of following 3 tarsomeres combined; pulvilli small, distance between basal two pulvilli 4.5 �� length of second pulvillus. Ovipositor sheath 0.9 �� length of middle tibia, apical sheath 1.4 �� length of basal sheath; lancet as in Fig. 6 i; middle serrulae as in Fig. 7 i. Male. Paratype from Asahidake-onsen. Length 11.5 mm (Figs 3 d, 5 t). Colour and structure similar to female except as follows: Pale line along posterior margin of pronotum extremely narrow; tegula entirely black; most of trochanter black, with broad apical margin whitish; trochantellus largely black; hind femur black except for wide stripe on anterior surface pale brown; antenna with antennomere 2 entirely black and antennomeres 3 to 5 marked with black ventrally; malar space 0.2 �� diameter of lateral ocellus; shortest distance between eyes 0.8 �� as long as height of eye; temple about 0.6 �� eye length in dorsal view (Fig. 11 b); hairs on dorsum of head, dorsum of mesoscutellum, and pleuron 1.3 �� as long as diameter of median ocellus, hairs on pleuron distinctly curved. Subgenital plate longer than broad, apical margin rounded (Fig. 11 d); valviceps narrow (Fig. 8 r); gonoforcep as Fig. 8 i. Etymology. This species is named in honour of the late K. Takeuchi, who made a pioneering contribution to the sawfly systematics in East Asia. Distribution. Japan (Hokkaido, Honshu, Kyushu). Holotype (Figs 3 c, 5 s): ♀, Ashiu-enshurin, Miyama-cho, Kyoto, Honshu, 11. VI. 1989, K. Mizuno, NSMT-I- HYM 65836, genitalia slide N 20130906 B (NSMT). Paratypes: 165 specimens (57 ♀ and 108 ♂, see Appendix). Variation. The length varies from 11.0 to 15.5 mm in the female and 9.5 to 13.0 mm in the male. The colouration of the antenna shows large variation, but the apical four or five antennomeres are always black and the scape and pedicel are usually partly or entirely black in both sexes. The other antennomeres are usually mostly or entirely brown, though the antenna is almost entirely black in very dark specimens. In the female, the mesoscutellum often has an obscure brownish spot medially, the hind trochanter is sometimes marked with black ventrally, and the abdominal terga 2 and 3 are often marked with black. In the male, the pale line along the posterior margin of the pronotum may be missing, the hind trochanter and trochantellus are always marked with black, the abdominal terga 2 (except for the narrow anterior margin) and 3 are always yellow brown, and the tergum 4 is often yellow brown at base. The ranges of some ratio variations are given in the key. Remarks. Siobla takeuchii is a member of the S. formosana group defined by Niu & Wei (2010), though the males could run to the S. sturmii group or to the S. ferox group in the key (Niu & Wei 2010). The new species closely resembles S. hirasana Takeuchi, 1929, belonging to the S. ferox group. In the female, S. takeuchii has the anterior margin of the clypeus weakly protruding (Fig. 11 c), the labrum dark brown basally, the posterior margin of the pronotum, tegula, and mesoscutellum usually entirely black (mesoscutellum rarely with small obscure spot medially), the metabasitarsus equal in length to the following three tarsomeres combined, hind trochanter sometimes marked with black, and the sawsheath black, except at apex. The female of S. hirasana has the anterior margin of the clypeus nearly truncate (Fig. 11 g), the labrum whitish basally, the broad posterior margin of pronotum, tegula, and mesoscutellum always mostly yellowish brown, the hind trochanter never marked with black, and the sawsheath entirely pale brown. In the male of S. takeuchii, the tegula and mesoscutellum are entirely black, the hind trochanter and trochantellus are usually largely blackish brown or black, and the subgenital plate is narrowly rounded at apex (Fig. 11 d), whereas in S. hirasana the tegula and mesoscutellum are largely yellowish brown, the hind trochanter and trochantellus are usually entirely whitish brown, rarely partly marked with blackish brown, and the subgenital plate is broadly rounded at apex (Fig. 11 h). The two species also differ in the shape of the head in dorsal view (Fig. 11 a���b, e���f). Takeuchi���s (1919 b) description of ��� Siobla ruficornis Cameron?��� was based on an unknown number of specimens from Yoshino, Nara prefecture, and two male specimens from ���Shinano��� (=Nagano prefecture). Two pairs of S. takeuchii collected in Yoshino in 1916 and kept in Takeuchi���s collection in OPU agree with the description of ��� Siobla ruficornis Cameron?��� by Takeuchi (1919 b) and should belong to his original material. Takeuchi (1919 b, p. 14) noted that the two male specimens from ���Shinano��� differed from the Yoshino specimens in that the antenna was yellowish brown except for the basal three antennomeres and in that the yellowish brown parts of the body were more reddish. We were not able to locate those two males in Takeuchi���s collection but they probably belong to S. sturmii because of the colour pattern of the antenna and the hind femur. The male specimen of ��� S. ferox ��� shown by Esaki et al. (1939) in their figure 592 2 in plate 152 has apically black antenna and entirely black dorsum of the thorax. This specimen was not available for the present study but it most probably belongs to S. takeuchii, because S. ferox has the apex of the antenna usually brown, though sometimes darkened, and has the dorsum of the thorax and the abdominal tergum 1 largely marked with pale brown. Togashi���s (1955) figure of the penis valve of ��� S. ferox ��� agrees most closely with that of S. takeuchii. Some specimens cited as S. ferox by Naito (1982) and Naito et al. (2004) and those listed under S. sturmii and Siobla sp. D by Katayama (2004) also belong to the present new species (see Appendix)., Published as part of Shinohara, Akihiko, Wei, Meicai & Niu, Gengyun, 2013, Revision of Siobla (Hymenoptera, Tenthredinidae) from Japan, pp. 1-40 in Zootaxa 3746 (1) on pages 22-26, DOI: 10.11646/zootaxa.3746.1.1, http://zenodo.org/record/285294

Key to the Japanese Species of Siobla 1 Female.............................................................................................. 2 - Male............................................................................................... 10 2 Body metallic blue........................................................... Siobla metallica Takeuchi, 1929 - Body black with brownish marks, not metallic blue........................................................... 3 3 Mesoscutellum and most or entire tegula yellowish brown..................................................... 4 - Mesoscutellum black, rarely with small spot at middle; tegula black, with very narrow outer margin sometimes yellowish brown............................................................................................... 6 4 Clypeus, at least apical 4 antennomeres, mesoscutellar appendage, fore and middle trochanters, fore trochantellus, and often middle trochantellus and bases of fore and middle femora black; metascutellum black, often with obscure yellowish macula medially; metapostnotum distinctly microsculptured, matt (Fig. 10 d)................... Siobla hirasana Takeuchi, 1929 - Clypeus, antenna, mesoscutellar appendage, metascutellum, and fore and mid trochanters, trochantelli and femora yellowish brown; metapostnotum hardly microsculptured, shiny (Fig. 10 e���f)............................................... 5 5 Eyes small, shortest distance between eyes below about 1.2���1.3 �� longest axis of eye (Fig. 10 a); postocellar area with distinct longitudinal median carina; mesoscutellum flat, not elevated above level of mesoscutum; forewing apically distinctly infuscated; hind tibia mostly orange, with only obscure black spot near apex; abdomen with tergum 1 largely black, segments 4 to 7 mostly whitish or pale orange ventrally, and sternum 7 with each lateral part roundly incised; lancet long (Fig. 6 g); penis valve as in Fig. 7 l...................................................... Siobla pulchra Shinohara, Wei & Niu, sp. nov. - Eyes large, shortest distance between eyes below about 1.0��� 1.1 �� longest axis of eye (Fig. 10 b); postocellar area without distinct longitudinal median carina; mesoscutellum strongly roundly elevated, higher than mesoscutum; forewing without distinct infuscation apically; hind tibia with apical third black; abdomen with tergum 1 largely orange, segments 4 to 7 usually mostly blackish ventrally, and sternum 7 with each lateral part nearly truncate; lancet normal (Fig. 6 b); penis valve as in Fig. 7 n.................................................................................. Siobla ferox (Smith, 1874) 6 Antenna black with apical 4 antennomeres creamy white (terminal antennomere usually darkened); pale areas on abdominal tergum 1 and tibiae of all legs creamy white; eyes large, shortest distance between eyes below about as broad as longest axis of eye; ovipositor sheath 1.0��� 1.2 �� middle tibia in length; mesoscutellum strongly elevated, much higher than mesoscutum................................................................................ Siobla apicalis Takeuchi, 1929 - Antenna largely reddish brown or largely black, apical 4 antennomeres not paler than antennomeres 4 and 5; pale areas on abdominal terga and tibiae yellowish brown; shortest distance between eyes below about 1.0��� 1.2 �� longest axis of eye; ovipositor sheath 0.7 ���1.0 �� middle tibia in length; mesoscutellum weakly elevated, about as high as mesoscutum............... 7 7 Apical 4 antennomeres black............................................................................ 8 - Apical 4 antennomeres reddish brown..................................................................... 9 8 Antenna with flagellum entirely or almost entirely black, with one or two basal flagellomeres often marked with brown dorsally; subapical antennomeres strongly dilated (Fig. 9 j), antennomere 7 with length about 1.5 �� breadth; shortest distance between eyes below 1.3 �� longest axis of eye (Fig. 9 h); broad posterior margin of pronotum usually reddish brown; hairs on temple curved at apex, length about 1.6���1.8 �� diameter of middle ocellus; hind leg with coxa, trochanter, trochantellus and femur all black; small species, length 9.5 ���12.0 mm.................... Siobla japonica Shinohara, Wei & Niu, sp. nov. - Antenna with basal three flagellomeres usually reddish brown, rarely almost entirely black; subapical antennomeres weakly dilated, antennomere 7 with length about 1.7���1.9 �� breadth; shortest distance between eyes below 1.1 �� longest axis of eye (Fig. 11 e); pronotum entirely black, or very narrow posterior margin brownish; hairs on temple straight, length about 1.0��� 1.2 �� diameter of ocellus; hind leg with apex of coxa, trochanter, trochantellus and base of femur pale brown; large species, length 11.0��� 15.5 mm................................................. Siobla takeuchii Shinohara, Wei & Niu, sp. nov. 9 Ovipositor sheath about as long as middle tibia; scape, pedicel and basal third of antennomere 3 largely black; apical sheath (=valvula 3) in lateral view as broad as hind tibia at apex in lateral view; abdominal tergum 2 usually coriaceous..................................................................................... Siobla jucunda (Mocs��ry, 1909) - Ovipositor sheath 0.7���0.9 �� middle tibia in length; basal 3 antennomeres entirely reddish brown or nearly so; apical sheath in lateral view clearly narrower than apex of tibia in lateral view; abdominal tergum 2 usually smooth.................................................................................................... Siobla sturmii (Klug, 1817) 10 Body metallic blue........................................................... Siobla metallica Takeuchi, 1929 - Body without metallic tinge............................................................................ 11 11 Mesoscutellar appendage pale yellow..................................................................... 12 - Mesoscutellar appendage black.......................................................................... 13 12 Eyes small, shortest distance between eyes below 1.0��� 1.1 �� longest axis of eye; postocellar area with distinct median longitudinal carina; mesoscutellum weakly raised, about as high as mesoscutum; hind tibia mostly reddish brown, with black spot on inner surface near apex; forewing distinctly infuscated apically; apical part of abdomen black dorsally and reddish brown ventrally........................................................ Siobla pulchra Shinohara, Wei & Niu, sp. nov. - Eyes large, shortest distance between eyes below 0.7���0.8 �� longest axis of eye; postocellar area without distinct median longitudinal carina; mesoscutellum strongly raised, much higher than mesoscutum; hind tibia with apical half black; forewing hyaline, without distinct apical infuscation; apical part of abdomen black on both sides........... Siobla ferox (Smith, 1874) 13 Antenna with at least 4 apical antennomeres black........................................................... 14 - Antenna with at least 4 apical antennomeres pale............................................................ 16 14 Antenna entirely black; abdominal terga 1���3 usually with distinct microsculpture dorsally, not polished; penis valve short and robust (Fig. 8 m); small species, length 7.5 ���11.0 mm................... Siobla japonica Shinohara, Wei & Niu, sp. nov. - Antenna usually with basal flagellomeres reddish brown; abdominal terga 1���3 usually without distinct microsculpture dorsally, polished; penis valve elongate; large species, length 9.5 ���14.0 mm.............................................. 15 15 Tegula and mesoscutellum usually largely yellowish brown; hind trochanter and trochantellus entirely whitish brown, rarely partly marked with blackish brown.............................................. Siobla hirasana Takeuchi, 1929 - Tegula and mesoscutellum entirely black; hind trochanter and trochantellus largely blackish brown or black........................................................................... Siobla takeuchii Shinohara, Wei & Niu, sp. nov. 16 Antenna black, with antennomeres 6���8 creamy white and antennomere 5 and 9 often partly dark brownish to whitish.................................................................................. Siobla apicalis Takeuchi, 1929 - Antenna reddish brown, with 3 or 4 basal antennomeres partly black........................................... 17 17 Malar space with length about 0.3 �� diameter of lateral ocellus (Fig. 10 g); shortest distance between eyes below about 0.9 �� longest axis of eye; at least apical 0.4 of antennomere 3 and more apical antennomeres entirely reddish brown; hairs on head blackish and straight (Fig. 10 g); hind femur marked with reddish brown.................... Siobla sturmii (Klug, 1817) - Malar space with length about 0.2 �� diameter of lateral ocellus (Fig. 10 h); shortest distance between eyes below about 0.6���0.7 �� longest axis of eye; apical 0.15 of antennomere 3 and more apical antennomeres reddish brown, sometimes antennomere 4 marked with black; hairs on head brownish, weakly curved at apex (Fig. 10 h); hind femur entirely black........................................................................................... Siobla jucunda (Mocs��ry, 1909), Published as part of Shinohara, Akihiko, Wei, Meicai & Niu, Gengyun, 2013, Revision of Siobla (Hymenoptera, Tenthredinidae) from Japan, pp. 1-40 in Zootaxa 3746 (1) on pages 3-5, DOI: 10.11646/zootaxa.3746.1.1, http://zenodo.org/record/285294

Siobla pseudoplesia Niu & Wei, 2012 (Figs 2 g, 2 h, 5 c, 5 d, 6 j–n, 9 h, 9 u, 10 w, 10 x) Siobla pseudoplesia Niu et al., 2012: 410. f #, type locality: Mt. Shennongjia, Hubei (China). Distribution. China (Ningxia, Shaanxi, Shanxi, Henan, Hubei, Sichuan). Remarks. The species is similar to S. acutitheca Niu & Wei, 2010 but differs from it in the ovipositor sheath distinctly shorter than middle tibia, the apical sheath very narrow in lateral view and acute at apex, the middle breadth of apical sheath about half breadth of the apex of hind tibia; the serrulae weakly sclerotized and almost flat, the 10 th serrula with 10–12 minute teeth; the postocellar area as broad as long; the interspaces between punctures on mesonotum microsculptured, almost mat; the abdominal tergite 1 weakly microsculptured, tergites 2–9 hardly microsculptured; and the first pulvillus small, distinctly shorter than half apical breadth of metabasitarsus.

Siobla pulchra Shinohara, Wei & Niu, sp. nov. (Figs 2 g���h, 5 o���p, 6g, 7g, 8g, p, 9 a���e, 10 a, e) urn:lsid:zoobank.org:act: 2 A07DDB 0-3 BA 3-4567 -BD 9 B- 5 F 4 D0A 67 CD 54 Siobla ferox: Naito et al., 2004: 56 (in part). Not Smith (1874). Siobla sp. 1: Tanaka & Tanaka, 2007: 109. Description. Female. Holotype (Figs 2 g, 5 o). Length 15.0 mm. Head and thorax black with following parts yellow white: clypeus, labrum, middle of mandible, antenna, posterior corner of pronotum, tegula, mesoscutellum, mesoscutellar appendage, metascutellum, minute stripe on middle of mesepimeron, and upper margin of metepimeron; palpi, narrow stripe on upper half of inner orbit, broad anterior stripe on upper half of hind orbit, and posterior part of postocellar area yellow brown. Abdomen yellow white with tergum 1 largely, small macula on tergum 3, most of terga 4 and 8 dorsally, terga 5���7 dorsally, and basal half of tergum 9 black. Legs yellow brown with base of fore coxa, middle and hind coxae, and apical 0.65 of hind femur black, and ventral side of hind tibia dark brown at apex. Body hairs yellow brown. Wings hyaline with weak yellow brown tinge, apical third distinctly infuscated, vein C and stigma yellow brown, other veins black brown. Apex of hind wing slightly grayish. Dorsum of head coarsely and densely punctured, interspaces between punctures on temple hardly recognizable, without microsculpture; clypeus shallowly and densely punctured, shiny; labrum shallowly and sparsely punctured; punctuation on dorsum of thorax similar to head; mesoscutellum matt, extremely densely punctured; mesoscutellar appendage polished, shiny, lateral corners microsculptured; metascutellum densely and coarsely punctured, anterior slope with linear interspaces between punctures; metapostnotum polished, shiny; mesepisternum feebly shiny, dorsal half coarsely and densely punctured and ventral half with small and shallow, dense, punctures; metepisternum with dorsal part densely punctured and ventral part sparsely punctured, shiny; middle and posterior part of metepimeron shiny, dorsal part of metepimeron coarsely and densely punctured; abdominal terga 1���3 strongly shiny, without microsculpture, other terga weakly microsculptured, terga 4���9 laterally and outer side of hind coxa shallowly and sparsely punctured. Hairs on temple straight, 1 �� diameter of median ocellus. Clypeus 2.2 �� as broad as long, with anterior margin truncate; malar space 1 �� diameter of median ocellus; lower interocular distance 1.2 �� eye height (Fig. 10 a); anterior margin of supraantennal tubercle elevated, small, posteriorly confluent with frontal ridge; middle fovea round and broad, deep; lateral fovea small and deep; interocellar furrow narrow and deep; postocellar furrows broad and shallow; postocellar area weakly elevated, as high as top of ocelli, with sharp middle carina, about 1.4 �� as broad as long; lateral furrows deep, distinctly curved outwards in anterior half and subparallel posteriorly; head behind eyes slightly shorter than eyes in dorsal view, convex at basal 0.4 and distinctly narrowing posteriorly (Fig. 9 a). Antenna shorter than vein C of forewing, about as long as head, thorax and abdominal tergum 1 together; pedicel slightly longer than broad, antennomere 3 1.5 �� as long as antennomere 4, subapical antennomeres slightly enlarged, antennomere 7 2 �� as long as broad (Fig. 9 e). Middle furrow on pronotum deep; mesoscutellum weakly elevated, anterior slope quite flat, much longer than posterior slope, without peak and carina; mesoscutellar appendage almost flat, without middle carina; mesepisternum roundly elevated, ventral spur absent. Apex of hind tibia distinctly enlarged; metabasitarsus 3.8 �� as long as broad, 1.1 �� length of following 3 tarsomeres combined; hind inner tibial spur 0.5 �� length of metabasitarsus; pulvilli small, distance between basal two pulvilli 2.5 �� length of second pulvillus. Hind wing with petiole of anal cell 0.7 �� length of cu-a. Abdominal tergum 1 with distinctly elongated carina just above spiracle; sternum 7 with each lateral part roundly incised; ovipositor sheath 0.93 �� length of middle tibia, apical sheath subequal in length to basal sheath (Fig. 9 d); lancet as in Fig. 6 g, middle serrulae as in Fig. 7 g. Male. Paratype from Nakayama-toge. Length 14.0 mm (Figs 2 h, 5 p). Head and thorax black, labrum, mesoscutellum, mesoscutellar appendage, metascutellum and dorsum of metepimeron yellow white; palpi and tegula yellow brown; antenna yellow brown, scape entirely and pedicel largely black; abdomen yellow brown, basal 0.65 of tergum 1 and dorsum of terga 5���8 black. Legs yellow brown; each coxa, broad and long inner stripe and outer apical 0.35 of hind femur black; dorsal stripe on fore and middle femora and ventral half of hind tibia black brown. Punctuation and structure similar to female except as follows: eyes large, shortest distance between eyes almost as long as height of eye; malar space 0.8 �� diameter of lateral ocellus (Fig. 9 c); lateral side of temples strongly narrowed backward in dorsal view (Fig. 9 b); pulvilli minute; subgenital plate longer than broad, apex round; penis valve as in Fig. 8 p; gonoforceps as in Fig. 8 g, harpe broad, less than 2 �� as long as broad. Etymology. The specific epithet, meaning beautiful, refers to the appearance of the new species. Distribution. Japan (Hokkaido, Honshu). Holotype (Figs 2 g, 5 o): ♀, [JAPAN: Honshu], Minoto, 1850m, Yatsugatake Mts., Nagano Pref. 25���29. VII. 2000, A. Shinohara, NSMT-I-HYM 68233, genitalia slide N 20131105 A (NSMT). Paratypes: 17 specimens (6 ♀ and 9 ♂, see Appendix). Variation. The length varies from 13.5 to 15.0 mm in the female and 12.5 to 14.0 mm in the male. The female specimens examined show little variation in colour pattern. The valvifer 2 (basal sheath) is subequal in length to or slightly longer than the valvifer 3 (apical sheath). Five males from Hyogo Prefecture, western Honshu, have the broad anterior margin of the clypeus pale yellow medially and the hind femur largely black. The ranges of some ratio variations are given in the key. Remarks. This new species belongs to the ferox group defined by Niu & Wei (2010) and is similar to S. fulvomarginata Wei & Nie, 1999, from Henan and Hunan Provinces, China. Siobla pulchra differs from S. fulvomarginata in the following features: head and thorax in both sexes largely black with a few yellow white maculae; eyes small, shortest distance between them longer than height of eye; postocellar area with distinct middle carina; dorsum of mesoscutellum flat, below top of mesoscutum; tergum 1 with distinct carina just lateral of spiracle; dorsum of head and thorax coarsely and densely punctured; apical 0.3 of forewing distinctly infuscate; tergum 1 and dorsum of terga 5���8 largely black and male subgenital plate longer than broad. Siobla pulchra differs from the Japanese congeners in the following characters. Female: clypeus, entire antenna, mesoscutellum, mesoscutellar appendage, metascutellum, and valvifer 2 (basal sheath) yellowish white; upper hind orbit and postocellar area marked with yellowish brown; postocellar area with distinct middle carina; forewing apically distinctly infuscated; mesoscutellum weakly raised; hind tibia mostly orange, with only obscure black spot near apex; tergum 1 largely black; abdominal segments 4 to 7 mostly whitish or pale orange ventrally; sternum 7 with each lateral part roundly incised; valvifer 2 (basal sheath) subequal in length to or slightly longer than valvifer 3 (apical sheath). Male: Clypeus entirely black or with broad anterior margin pale yellow medially; antenna yellowish white, with scape and most of pedicel black, and flagellomere 1 partly darkened; mesoscutellum, mesoscutellar appendage and metascutellum whitish yellow; mesoscutellum weakly swollen; forewing apically distinctly infuscated; hind tibia mostly orange, with only obscure black spot near apex; tergum 1 with basal half black. Tanaka & Tanaka (2007) noted that their Siobla sp. 1 ���resembles S. ferox but the apex of the forewing is dark and there is a difference in hind tibia��� (original in Japanese, p. 109; see also fig. 1 in p. 113). This species is probably S. pulchra., Published as part of Shinohara, Akihiko, Wei, Meicai & Niu, Gengyun, 2013, Revision of Siobla (Hymenoptera, Tenthredinidae) from Japan, pp. 1-40 in Zootaxa 3746 (1) on pages 18-21, DOI: 10.11646/zootaxa.3746.1.1, http://zenodo.org/record/285294

Siobla parallela Niu & Wei, sp. nov. (Figs 2 e, 2 f, 5 a, 5 b, 7 d���f, 9g, 9t, 10 q, 10 r) Description. Holotype: ♀. Length, 10 mm (Figs 2 e, 5 a). Black, labrum yellowish white, palpi, apical half of 2 nd antennomere, base of 3 rd antennomere, posterior margin of pronotum (greatest width 1.0 �� length of lateral ocellus), tegula, apex of mesoscutellum, posterior ridge of mesoparapsides and of metaparapsides, entire 2 nd abdominal tergite, most of 3 rd tergite, 2 spots on 4 th tergite, 2 nd to 4 th sternites, and most of 5 th sternites yellowish brown, middle of 10 th tergites, cerci and posterior margin of sheath pale brown; legs yellowish brown, most of fore coxa, basal half of middle and of hind coxae, and apical half of hind femur black, irregular macula on apical of hind tibia dark brown; body hairs silver; wings hyaline, pterostigma pale brown. Head densely and coarsely punctured without interspaces, labrum sparsely punctured; prescutum, parts of scutum, and anterior slope of mesoscutellum densely punctured, interspaces narrow, lateral parts of mesoscutellum punctured with shiny interspaces; posterior slope densely and coarsely punctured, without interspaces; posttergite and metapostnotum shiny with margins microsculptured; parapsides with large and sparse punctures and dense microsculpture; metascutellum with anterior slope shiny and posterior slop densely punctured; middle and upper of mesepisternum densely and coarsely punctured, venter with large and shiny interspaces between small and sparse punctures; anepimeron coarsely punctured, katepimeron microsculptured, posterior margin shiny; metepisternum with upper densely punctured, mat, and venter sparsely punctured, shiny; basin and posterior corner of metepimeron shiny, dorsum with large punctures. First abdominal tergite hardly microsculptured, shiny; punctures on lateral edges shallow; other tergites obviously microsculptured, parts of lateral edges of 3 rd tergite, lateral edges and posterior margin of 4 th tergite and 5 th to 8 th tergites mostly punctured. Hairs on dorsum of head straight, as long as diameter of lateral ocelli. Clypeus flat, anterior margin truncate; malar space as long as transverse diameter of median ocellus; shortest distance between eyes below antennal sockets in front view 1.1 �� eye height; middle fovea broad and shallow; interocellar furrow deep, postocellar furrow broad and shallow; postocellar area 1.2 �� as broad as long, weakly elevated with an obtuse carina; lateral furrows shallow, straight, parallel posteriorly; head behind eyes 0.9 �� length of eye in dorsal view; curved laterally and then convergent posteriorly; genal carina sharp, complete (Fig. 7 d). Antenna as long as head, thorax and 1 st abdominal tergite together, hardly shorter than vein C, 2 nd antennomere 1.3 �� as long as broad, relative length of 3 rd to 4 th antennomeres as 1.6, 5 th to 8 th antennomeres weakly compressed, enlarged, inner and outer sides of 6 th to 8 th antennomeres with weak furrows, 7 th antennomere about 2 �� as long as broad. Middle furrow of prescutum narrow but shallow; mesoscutellum obviously elevated, slightly higher than scutum. Metabasitarsus 5.5 �� as long as broad, about 1.2 �� length of following 3 tarsomeres combined, hind inner tibial spur 0.4 �� length of metabasitarsus. Ovipositor sheath 1.1 �� middle tibia, venter margin curved, exposed part of apical sheath about 2 �� as long as broad; apical sheath narrow in dorsal view, exposed part 4.5 �� length of cerci, lateral setae on both sides of sheath forming an angle about 40 degree. Lancet broadened before apex, with 14 serrulae (Fig. 9 g), middle serrulae each with 10���12 distal fine teeth (Fig. 9 t). Male. Length, 9 mm (Figs 2 f, 5 b). Similar to female in structure and color, but entire 2 nd and 3 rd abdominal tergites, most of 4 th tergite, and entire 2 nd to 5 th sternites yellowish brown; legs yellowish brown, most of each coxa, apical 0.5 to 0.6 of hind femur, apical 0.25 of hind tibia black, base of fore 1 st trochanter and base of fore femur dark brown; malar space 0.8 �� transverse diameter of median ocellus (Fig. 7 f); head in front view with lower interocular distance 0.9 �� eye height; head behind eyes 0.6 �� eyes in length in dorsal view, curved in middle and narrowing posteriorly (Fig. 7 e); genitalia as in Figs 10 q, 10 r. Variation. The pale spot on the dorsal side of base of flagellum, the pale spot on tergite 4 in female and the black macula on hind femur in male slightly vary in size. Etymology: This species is named after the lateral furrows of postocellar area which are parallel to each other. Specimens examined. 9 ♀, 4 ♂. Hunan: Holotype: 1 ♀, Huangsang, Suining, 600���900m, 2005 -IV- 21, Lin Yang leg.; CSCSHT 0 0 810023 (CSCS); Paratypes: 5 ♀ 4 ♂, Huangsang, Suining, 600���900m, 2005 -IV- 21, Lin Yang, Liu Weixing, Liu Shouzhu, Zhou Hu leg.; 1 ♀, Taoyuandong, Yanling, 1999 -IV- 24, 900��� 1000m, Wei Meicai leg.; 1 ♀, CSCS 13012, Yunfengge, Mt. Yun, Wugang, Shaoyang, E. 110 �� 37.169��, N. 26 �� 38.983��, 1170m, 2013 -IV- 14, Li Zejian leg.; Guizhou: 1 ♀, Linchang, Leigongshan, 2005 -V- 31, 1600m, Liang Minwen leg.. Distribution. China (Hunan, Guizhou). Remarks. The new species is similar to S. listoni Niu & Wei, 2012 but differs from it in the forewing hyaline and pterostigma pale brown; the antenna long and slender, with the subapical antennomeres very weakly enlarged, the 6 th antennomere more than 2 �� as long as broad; the breadth of apical sheath in lateral view distinctly narrower than breadth of metabasitarsus, apical sheath 2 �� as long as basal sheath; the middle serrulae each with 10���12 distal teeth; the dorsal side of mesoscutellum roundly elevated with the posterior margin round; the posttergite and metapostnotum largely polished, posttergite without a middle carina., Published as part of Niu, Gengyun & Wei, Meicai, 2013, Revision of the Siobla formosana group (Hymenoptera: Tenthredinidae), pp. 41-68 in Zootaxa 3746 (1) on pages 50-51, DOI: 10.11646/zootaxa.3746.1.2, http://zenodo.org/record/285307

Siobla ferox (Smith, 1874) (Figs 1 c���d, 3 e���i, 5 c���d, 6 b, 7 b, 8 b, k, 10 b, f) Macrophya ferox Smith, 1874: 379. Siobla ferox: Nakagawa, 1899: 204; Takeuchi, 1919 a: 184; Takeuchi, 1919 b: 12; Yano, 1932: 438 (in part); Malaise, 1934: 24; Esaki et al., 1939: 337 (in part); Takeuchi, 1940: 479; Takeuchi, 1950: 1348; Takeuchi, 1952: 18 (in part); Okutani, 1954: 78; Takeuchi, 1955: 120, plate 54, 810; Kim, 1963: 288; Togashi, 1965: 246 (in part); Okutani, 1965: 162, 471; Kim, 1970: 200, 728; Okutani, 1974 a: 54;?Naito, 1982: 574; Abe and Togashi, 1989: 555 (in part); Kim et al., 1994: 222; Katayama, 2000: 13; Lee et al., 2000: 117; Katayama, 2004: 95; Naito et al., 2004: 56, fig. 105 (in part); Yoshida, 2006: 91 (in part); Togashi, 2008: 490 (in part); Lee et al., 2010: 165; Taeger et al., 2010: 583; Niu & Wei, 2010: 49; Lelej, 2012: 100. Siobla grandis Matsumura, 1912: 50; Takeuchi, 1919 a: 184 (as a synonym of S. ferox); Takeuchi, 1919 b: 12 (as a synonym of S. ferox); Doi, 1938: 33. Siobla ferox var. hirasana: Takeuchi, 1937: 57. Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima), Korea, China. Primary types examined. Lectotype of Macrophya ferox hereby designated (Fig. 3 e���g): ♂, ��� Type H. T.��� ���B. M. TYPE HYM. 1.245 ��� ��� Macrophya ferox, Type, Smith��� ��� 47 22 ��� ���Kb. t. 9 / 20.��� ���BMNH(E) # 651764 ��� (BMNH). Lectotype of Siobla grandis hereby designated (Fig. 3 h���i): ♀, ��� 9 /VI./ 1900, Sapporo���, ��� 26 ���, ��� Siobla grandis Mats., ♀ Type ��� (HU). Other material examined (see Appendix). 484 specimens (137 ♀ and 347 ♂). Variation: The length varies from 12.0 to 16.0 mm in the female and 10.0 to 14.5 mm in the male. In the female, the head capsule is usually largely marked with reddish brown but sometimes it is all black except for the clypeus and malar space. The antenna is always entirely pale brown and the mesoscutum sometimes has large obscure brownish markings. In the male, the anterior margin of the clypeus is usually narrowly creamy white but sometimes the entire clypeus is black. The antenna is pale brown, with blackish areas of variable extent on the scape, pedicel and apical flagellomeres. The palest specimens have an entirely pale brown antenna, whereas the darkest specimens have the scape and four or five apical antennomeres mostly black and the ventral surface of the other antennomeres marked with black. The ranges of some ratio variations are given in the key. Remarks. This is a member of the S. ferox group defined by Niu & Wei (2010). From the other species of Siobla occurring in Japan, S. ferox is distinguished by a combination of the entirely pale brown antenna of the female, the yellow, strongly convex mesoscutellum, the pale yellow mesoscutellar appendage, and the largely reddish brown abdominal tergum 1. Previous studies on Japanese Siobla treated S. ferox as a very variable species. We here recognize four species (S. ferox, S. hirasana, S. pulchra and S. takeuchii), which can be separated by the characters given in the key. In addition to the works cited in the list of synonymy above, Togashi (1961, 1970a, 1983, 1997b, 1998 b), Okutani (1972, 1974b), Otsuka (1984), Nambu (1992, 1998), and Katayama et al. (2007) referred to ��� S. ferox ���. Identity of the species they treated should be confirmed by examining the original material actually used in these works. Takeuchi (1949), Okutani (1954, 1959, 1967) and Nakamura (1962) reported on the host plants of ��� S. ferox ��� and Okutani (1959) gave a description of the larva. However, we were not able to examine any reared adult specimens and thus we cannot confirm which species they actually studied. Smith (1874) did not designate types and did not give the number of specimens he examined when he described Macrophya ferox, whereas Kirby (1882) mentioned that the species was ���Described from one male example (not a female, as stated by Smith)��� and listed one male as ��� Type.��� We designate this male specimen as lectotype of M. ferox Smith, 1874. Matsumura (1912), in the original description of his Siobla grandis, did not designate types and did not give the number of specimens he had. A female specimen (Fig. 4 b) which agrees well with the original description and bears a red label ��� Siobla grandis Mats., ♀ Type ��� (Fig. 4 a). is here designated as lectotype., Published as part of Shinohara, Akihiko, Wei, Meicai & Niu, Gengyun, 2013, Revision of Siobla (Hymenoptera, Tenthredinidae) from Japan, pp. 1-40 in Zootaxa 3746 (1) on pages 5-12, DOI: 10.11646/zootaxa.3746.1.1, http://zenodo.org/record/285294