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Leuconemacris acuminata, sp. nov. (Figs 1–5, 22–23) Description. Male. Small size. Head. Head 1.0–1.1 times longer than pronotum; frons slightly oblique in lateral view; fastigium triangular, apex acutely angular, dorsum weakly depressed with distinct median and lateral carinae, median carina continuing to half of occiput; vertex between eyes 1.6–1.8 times as wide as frontal costa between antennae; frontal costa broad and shallowly concave at whole length, lateral margins nearly parallel but gradually expanded below median ocellus. Antennae thick, 20 segments, surpassing hind margin of pronotum, basal segments weakly compassed, median segment (9 th –13 th) mean 1.3 times as long as wide. Eyes roundly triangular, longitudinal diameter 1.2–1.3 times horizontal diameter and 0.9–1.0 times the length of subocular furrow. Thorax. Pronotum with anteral margin nearly straight, posteral margin nearly straight with a small median notch; median and lateral carinae distinct, lateral carinae a little recurved medially; prozona 1.4–1.6 times as long as metazona. Mesosternal interspace trapezoidal, minimum width 2.0 times larger than length; metasternal lobes separate. Tegmen abbreviated, 2.5 times as long as wide, dorsally touching each other, apex rounded, not reaching or just reaching 1/3 of hind femur, or surpassing hind margin of 5 th abdominal tergite; anterior margin sinuate, precostal area ending at half of whole length; costal area 1.7–2.0 times as wide as subcostal area, medial area weakly wider than or equal to radial area. Hind wing vestigial, not surpassing half of 1 st abdominal tergite. Hind femur 4.0–4.3 times longer than wide, inner side with 44–49 conical stridulatory pegs. Hind tibia with 11 spines on both dorsal margins, outer apical spine absent. Abdomen. Tympanum developed. 10 th abdominal tergite excised in middle, with furculae. Epiproct rhombic, as long as wide, shallowly furrowed at basal half, apex rounded. Cerci long conical, finely surpassing apex of epiproct, apex blunt. Subgenital plate distally observably slimmed, long conical, apex acute. Epiphallus with bridge semicircular; lateral plate with outer-lower corner pointed; lophi nearly oblong, distal and basal portions densely interspersed with granules; anchorae stout, pointing ahead and ventrad. Coloration. Body greyish-green or greenish-brown. Antennae with basal segments brown, others dark. Eye brown. Tegmen with basal portion brown and apical portion hyaline but costal area white in basal portion. Hind femur with outer side brown, lower side yellow. Hind tibia yellowish-red. Female. Similar to male, but body stout, eyes longitudinal diameter 1.1 times horizontal diameter and 0.8 times the length of subocular furrow, the notch on the posterior margin of pronotum clearer, tegmen surpassing hind margin of 1 st abdominal tergite. Subgenital plate oblong, hind margin acutely protruding in the middle. Upper ovipositor valves with upper-outer margin rough, lower ovipositor valve nearly straight, margin smooth. Measurements (mm). Length of body: ♁12.0–13.0, ♀ 18.0–20.0; length of pronotum: ♁ 2.4–2.5, ♀ 3.3–3.5; length of hind femur: ♁ 7.9–8.0, ♀ 8.7–9.0. Type material. Holotype: ♁, China, Sichuan, Muli (Bowo), 28.69° N, 101.11° E, alt. 3910 m, 18 July 2021, collected by Rong-Long Yang. Paratypes. 9♁, 3♀, same data, deposited in BMDU. Diagnosis. In this genus, the species is characterised by the male subgenital plate, which is long conical, distally observably slimmed, apex acute. Etymology. The specific epithet refers to the apex of male subgenital plate being acute. Distribution. CHINA: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on pages 267-268, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313

Leuconemacris xinlongensis, sp. nov. (Figs 11–15, 30–31) Description. Male. Small size. Head. Head longer, 1.10 times as long as pronotum; frons slightly oblique in lateral view; fastigium triangular, apex acutely angular, dorsum weakly depressed with distinct median and lateral carinae, median carina continuing to half of occiput; vertex between eyes 1.6–1.7 times as wide as frontal costa between antennae; frontal costa broad and flat but finely depressed around median ocellus, lateral margins parallel but weakly contracted near median ocellus. Antennae short and thick, 20 segments, not reaching hind margin of pronotum, median segment (9 th –13 th) mean 0.8 times as long as wide. Eyes roundly triangular, longitudinal diameter 1.1 times horizontal one and 1.0–1.1 times the length of subocular furrow. Thorax. Pronotum with anteral margin straight, posteral margin with a small median notch; median and lateral carinae distinct, lateral carinae a little recurved medially; prozona 1.4 times as long as metazona. Mesosternal interspace trapezoidal, minimum width 2.1 times larger than length; metasternal lobes separate. Tegmen abbreviated, 2.3 times as long as wide, touching each other in mid dorsal line, apex rounded, not reaching or just reaching half of hind femur, or surpassing hind margin of the 7 th abdominal tergite; anterior margin sinuate, precostal area ending at 1/3 of whole length; costal area 1.6–1.7 times as wide as subcostal area, medial area weakly wider than or equal to radial area. Hind wing vestigial, not surpassing half of 1 st abdominal tergite. Hind femur 3.7 times longer than wide, inner side with 48–53 conical stridulatory pegs. Hind tibia with 11 spines on both dorsal margins, outer apical spine absent. Abdomen. Tympanum developed. 10 th abdominal tergite partially excised in middle, furculae faint. Epiproct triangular, as long as wide, shallowly furrowed at basal half, apex rectangular. Cerci long conical, surpassing hind margin of epiproct, apex blunt. Subgenital plate short conical, apex obtuse. Epiphallus with bridge semicircular; lateral plate with outer-lower corner acute angled; lophi nearly nearly oblong in lateral view, distal portion densely interspersed with granules; anchorae stout, pointing ahead and ventrad. Coloration. Body greyish-green or greyish-brown.Antennae with basal segments brown, others dark. Postocular bands black. Tegmen with anal area yellowish brown, others with basal portion dark and apical portion hyaline but costal area white in basal portion. Hind femur brown or greyish-brown, outer side with two blackish spots in most individuals. Hind tibia greyish-brown. Female. 5 th nymph, similar to male, but the notch on the posterior margin of pronotum deeper. Measurements (mm). Length of body: ♁11.5–12.4; length of pronotum: ♁ 2.4–2.5; length of hind femur: ♁ 7.5–7.6. Type material. Holotype: ♁, China, Sichuan, Xinlong (Tongxiao), 30.85° N, 99.94° E, 24 Aug. 2020, collected by Zhi-Pan Huang. Paratypes. 4♁, 1♀ (5 th nymph), same data, deposited in BMDU. Diagnosis. This new species is similar to L. muliensis, sp. nov., but differs from the latter in: 1) antennae not surpassing hind margin of pronotum, median segment mean 0.8 times as long as wide, in the latter antennae finely surpassing hind margin of pronotum, median segment mean 1.3 times as long as wide, 2) male tegmen with precostal area ending at 1/3 of whole length against 1/ 2 in the latter, 3) male 10 th abdominal tergite with very faint furculae instead of clear furculae in the latter. Etymology. The specific epithet refers to the type locality, Xinlong County, Sichuan Province, China. Distribution. CHINA: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on pages 271-272, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313

Leuconemacris muliensis, sp. nov. (Figs 6–10, 26–27) Description. Male. Small size. Head. Head 1.1 times longer than pronotum; frons slightly oblique in lateral view; fastigium triangular, apex acutely angular, dorsum weakly depressed with distinct median and lateral carinae, median carina continuing to half of occiput; vertex between eyes 1.8 times as wide as frontal costa between antennae; frontal costa broad and shallowly concave at whole length, lateral margins nearly parallel but gradually expanded below median ocellus. Antennae thick, 20 segments, slight surpassing hind margin of pronotum, basal segments weakly compassed, median segment (9 th –13 th) mean 1.3 times as long as wide. Eyes roundly triangular, longitudinal diameter 1.2–1.3 times horizontal diameter and 0.9–1.0 times the length of subocular furrow. Thorax. Pronotum with anteral margin straight, posterior margin with a small median notch; median and lateral carinae distinct, lateral carinae a little recurved medially; prozona 1.4–1.6 times as long as metazona. Mesosternal interspace trapezoidal, minimum width 1.9 times larger than length; metasternal lobes separate. Tegmen abbreviated, 2.2 times as long as wide, dorsally touching each other, apex rounded, reaching 1/3–2/5 of hind femur, or reaching hind margin of 5 th abdominal tergite; anterior margin sinuate, precostal area ending at half of whole length; costal area 1.6–2.0 times as wide as subcostal area, medial area wider than or equal to radial area. Hind wing vestigial, not surpassing half of 1 st abdominal tergite. Hind femur 4.0–4.3 times longer than wide, inner side with 58 conical stridulatory pegs. Hind tibia with 11 spines on both dorsal margins, outer apical spine absent. Abdomen. Tympanum developed. 10 th abdominal tergite excised in middle, with furculae. Epiproct pentagonal, 1.0–1.1 times as long as wide, shallowly furrowed at basal half, apex rounded. Cerci long conical, finely surpassing apex of epiproct, apex blunt. Subgenital plate strangely tapered, long conical, apex bluntly acute. Epiphallus with bridge semicircular; lateral plate with outer-lower corner ectad protruding; lophi with distal and basal portions densely interspersed with granules; anchorae stout, pointing ahead and ventrad. Coloration. Body greyish-green or greenish-brown. Antennae with basal segments brown, others dark. Eye brown. Tegmen with basal portion brown and apical portion hyaline but costal area white in basal portion. Hind femur with outer side brown, lower side yellow. Hind tibia yellowish-red. Female. Similar to male, but body stout, eyes longitudinal diameter 1.2 times horizontal diameter and 0.8 times the length of subocular furrow, the notch on the posterior margin of pronotum clearer, tegmen surpassing hind margin of 1 st abdominal tergite. Subgenital plate oblong, acutely protruding in the middle. Upper ovipositor valves with upper-outer margin rough, lower ovipositor valve nearly straight, edge rough. Measurements (mm). Length of body: ♁12.0–13.0, ♀ 18.5–20.0; length of pronotum: ♁2.4–2.5, ♀ 3.3–3.5; length of hind femur: ♁7.9–8.0, ♀ 8.7–9.0. Type material. Holotype: ♁, China, Sichuan, Muli (Bowo), 28.97° N, 101.11° E, alt. 4028 m, 18 July 2021, collected by Rong-Long Yang and Zhi-Pan Huang. Paratypes. 20♁, 20♀, same data, deposited in BMDU. Diagnosis. This species is closely related to L. xiangchengensis, in general features, especially in the length of male tegmen, in the length of precostal area being 1/2 of whole length, in the same ratio of costal area comparing subcostal area in width, and the undulate upper-outer margin of upper ovipositor valves. But it differs from the latter in: 1) head 1.1 times as long as pronotum (length of head less than that of pronotum in the latter); 2) longitudinal diameter of eye larger than transverse diameter 1.2–1.3 times (the longitudinal diameter a little larger than or equal to transverse diameter in the latter); 3) antennae with median segment (9 th –13 th) mean 1.3 times as long as wide (the width of median segment larger than the length in the latter); 4) male tegmen with anterior margin sinuate (the margin nearly straight in the latter). Etymology. The specific epithet refers to the type locality, Muli County, Sichuan Province, China. Distribution. CHINA: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on pages 269-271, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313

Key to the species of Leuconemacris Zheng, 1988 1. Tegmen longer, reaching the base of epiproct............................................................... 2 –. Tegmen shorter, just reaching the 4 th 7 th abdominal tergite..................................................... 5 2. Male tegmen with costal area broad, maximum width 2.0 times as wide as subcostal area, maximum width of medial area 2.0 times or more than that of radial area...................................................................... 3 –. Male tegmen with costal area narrower, maximum width 2.0 times less than that of subcostal area, maximum width of medial area 2.0 times less than that of radial area.................................................................. 4 3. Tegmen longer, mean 6.9 mm in male or 3.5 mm in female; inner side of hind femur with 51 conical stridulatory pegs in male......................................................................................... L. litangensis –. Tegmen shorter, mean 5.3 mm in male or 2.3 mm in female; inner side of hind femur with 40 short thick conical stridulatory pegs in male............................................................................... L. xizangensis 4. Female tegmen longer, reaching half of 2 nd abdominal tergite; inner side of male hind femur with 54 conical stridulatory pegs which distally suddenly thinned, apex acute................................................... L. daochengensis –. Female tegmen shorter, not reaching hind margin of 1 st abdominal tergite; inner side of male hind femur with 50 long conical stridulatory pegs which distally gradually thinned.................................................. L. longipennis 5. Male tegmen with anterior margin nearly straight; length of head shouter than that of pronotum....................... 6 –. Male tegmen with anterior margin sinuate; length of head longer than that of pronotum.............................. 7 6. Male tegmen with precostal area broad, ending at 2/3 of whole length; costal area slightly wider than that of subcostal area; upper ovipositor valves with upper-outer margin smooth............................................. L. breviptera –. Male tegmen with precostal area narrow, ending at 1/2 of whole length; costal area 1.5–2.0 times as wide as subcostal area; upper ovipositor valves with upper-outer margin undulate....................................... L. xiangchengensis 7. Male epiproct rhombic, 1.3 times as long as wide; subgenital plate distally observably slimmed, long conical, apex acute......................................................................................... L. acuminata sp. n. –. Male epiproct triangular, as long as wide; subgenital plate short conical, apex blunt................................. 8 8. Male antennae longer, surpassing hind margin of pronotum, median segment 1.3 times as long as wide; tegmen with precostal area ending at 1/2 of whole length; furcula on hind margin of 10 th abdominal tergite distinct............ L. muliensis sp. n. –. Male antennae shorter, not surpassing hind margin of pronotum, median segment 0.8 times as long as wide; tegmen with precostal area ending at 1/3 of whole length; furcula on hind margin of 10 th abdominal tergite indistinct. L. xinlongensis sp. n., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on page 267, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313, {"references":["Zheng, Z. M. (1988) A study on the Leuconemacris gen. nov. of grasshoppers from Hengduan Shan range of China. Etomotaxonomia, 10 (3 - 4), 283 - 292."]}

Leuconemacris breviptera (Yin, 1983) http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105282 Podismopsis breviptera Yin, 1983: 36–38, 44–45, figs 5–8, pl. I: 3–4 (holotype – male, China: Sichuan, Litang, 4300 m; in NPIB). Leuconemacris breviptera (Yin, 1983); Zheng, 1988: 285. Leuconemacris brevipennis (Yin, 1983); Zheng & Xia, 1998: 274–275, fig. 143 (lapsus calami). Distribution. China: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on page 269, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313, {"references":["Yin, X. C. (1983) Four new genera and seven new species of grasshoppers from Hengduan Shan range of China. Zoological Research, 4 (1), 35 - 46.","Zheng, Z. M. (1988) A study on the Leuconemacris gen. nov. of grasshoppers from Hengduan Shan range of China. Etomotaxonomia, 10 (3 - 4), 283 - 292.","Zheng, Z. M. & Xia, K. L. (1998) Fauna Sinica, Insecta, Vol. 10, Orthoptera, Acridoidea, Oedipodidae and Arcypteridae. Science Press, Beijing, 616 pp."]}

Leuconemacris daochengensis Zheng, 1988 (Figs 24–25) http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105283 Leuconemacris daochengensis Zheng, 1988: 285–286, 291–292, figs 3, 13–14, 25–30 (holotype – male, China: Sichuan, Daocheng; in IZCAS); Zheng & Xia, 1998: 269–271, fig. 140. Material examined. 27♁, 22♀, China: Sichuan, Daocheng (Yading), 28.56° N, 100.19° E, alt. 4151–4400 m, 21 Aug. 2020, collected by Rong-Long Yang and Zhi-Pan Huang, deposited in BMDU. Distribution. China: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on page 269, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313, {"references":["Zheng, Z. M. (1988) A study on the Leuconemacris gen. nov. of grasshoppers from Hengduan Shan range of China. Etomotaxonomia, 10 (3 - 4), 283 - 292.","Zheng, Z. M. & Xia, K. L. (1998) Fauna Sinica, Insecta, Vol. 10, Orthoptera, Acridoidea, Oedipodidae and Arcypteridae. Science Press, Beijing, 616 pp."]}

Leuconemacris litangensis (Yin, 1983) http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105284 Podismopsis litangensis Yin, 1983: 35–36, 44, figs 1–4, pl. I: 1–2 (holotype – male, China: Sichuan, Litang, 3800–4200 m; in NPIB). Leuconemacris litangensis (Yin, 1983); Zheng, 1988: 284; Zheng & Xia, 1998: 267–268, fig. 138. Distribution. China: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on page 269, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313, {"references":["Yin, X. C. (1983) Four new genera and seven new species of grasshoppers from Hengduan Shan range of China. Zoological Research, 4 (1), 35 - 46.","Zheng, Z. M. (1988) A study on the Leuconemacris gen. nov. of grasshoppers from Hengduan Shan range of China. Etomotaxonomia, 10 (3 - 4), 283 - 292.","Zheng, Z. M. & Xia, K. L. (1998) Fauna Sinica, Insecta, Vol. 10, Orthoptera, Acridoidea, Oedipodidae and Arcypteridae. Science Press, Beijing, 616 pp."]}

Leuconemacris xiangchengensis Zheng, 1988 (Figs 28–29) http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105287 Leuconemacris xiangchengensis Zheng, 1988: 289–290, 292, figs 7, 21–22, 44–50 (holotype – male, China: Sichuan Province, Xiangcheng, 4550 m; in IZCAS); Zheng & Xia, 1998: 275–276, fig. 144. Material examined. 2♁, 2♀, China: Sichuan, Xiangcheng (Niding), 29.12° N, 99.76° E, alt. 4000 m, 3 Sep. 2020, collected by Zhi-Pan Huang; 5 ♁, 7♀, China: Sichuan, Xiangcheng (Fozhuxia), 29.13° N, 100.05° E, alt. 4800 m, 4 Sep. 2020, collected by Zhi-Pan Huang, deposited in BMDU. Distribution. China: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on page 271, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313, {"references":["Zheng, Z. M. (1988) A study on the Leuconemacris gen. nov. of grasshoppers from Hengduan Shan range of China. Etomotaxonomia, 10 (3 - 4), 283 - 292.","Zheng, Z. M. & Xia, K. L. (1998) Fauna Sinica, Insecta, Vol. 10, Orthoptera, Acridoidea, Oedipodidae and Arcypteridae. Science Press, Beijing, 616 pp."]}

Leuconemacris longipennis Zheng, 1988 http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105285 Leuconemacris longipennis Zheng, 1988: 288–289, 292, figs 4, 15–16, 38–43 (holotype – male, China: Sichuan, Batang, 4450– 4680 m; in IZCAS); Zheng & Xia, 1998: 271–272, fig. 141. Distribution. China: Sichuan., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on page 269, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313, {"references":["Zheng, Z. M. (1988) A study on the Leuconemacris gen. nov. of grasshoppers from Hengduan Shan range of China. Etomotaxonomia, 10 (3 - 4), 283 - 292.","Zheng, Z. M. & Xia, K. L. (1998) Fauna Sinica, Insecta, Vol. 10, Orthoptera, Acridoidea, Oedipodidae and Arcypteridae. Science Press, Beijing, 616 pp."]}

Leuconemacris xizangensis (Yin, 1984) (Figs 32–33) http://orthoptera.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1105288 Podismopsis xizangensis Yin, 1984: 190–191, 276, figs 406–408, pl. XXVII: 206–207 (holotype – male, China: Xizang, Mangkang, 4200 m; in NPIB). Leuconemacris xizangensis (Yin, 1984); Zheng, 1988: 285; Zheng & Xia, 1998: 268–269, fig. 140. Material examined. 8♁, 6♀, China: Xizang, Mangkang (Gato), 29.51° N, 98.53° E, alt. 4000–4200 m, 15 Aug. 2019, collected by Zhi-Pan Huang, deposited in BMDU. Distribution. China: Xizang., Published as part of Mao, Ben-Yong & Huang, Zhi-Pang, 2023, Taxonomy on three allied genera within Arcypterini (Orthoptera: Acrididae) from Qinghai-Xizang Plateau, China, pp. 265-279 in Zootaxa 5239 (2) on page 273, DOI: 10.11646/zootaxa.5239.2.5, http://zenodo.org/record/7624313, {"references":["Yin, X. C. (1984) Grasshoppers and Locusts from Qinghai-Xizang Plateau of China. Science Press, Beijing, 287 pp.","Zheng, Z. M. (1988) A study on the Leuconemacris gen. nov. of grasshoppers from Hengduan Shan range of China. Etomotaxonomia, 10 (3 - 4), 283 - 292.","Zheng, Z. M. & Xia, K. L. (1998) Fauna Sinica, Insecta, Vol. 10, Orthoptera, Acridoidea, Oedipodidae and Arcypteridae. Science Press, Beijing, 616 pp."]}