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A new species of silvanid flat bark beetle, Protoliota paleus sp. nov., is described from mid-Cretaceous Burmese amber. Protoliota paleus nov. sp. differs from a previous member of the genus described in Burmese amber by its small body size, weakly protruding eyes, long mandibles, and narrow pronotum. The new fossil is characterized by extremely elongate antennae and setiferous pads on its fore tarsus. Discussion is presented on the purpose of both of these attributes and how they could play a role in the survival of the species. A list of the superfamily Cucujoidea described from Burmese amber is included.

Seed germination is a crucial stage in the development of all plants. It normally occurs in the soil after the seed has fallen from the mother plant. In some infrequent instances, precocious germin...

Premise of research. This investigation was undertaken to describe a multifloral grass spikelet enclosed in Baltic amber. The fossil, described in the new genus Eograminis, represents the first def...

In this article, a new species of the genus Amberophytum Yu, Slipinski et Pang, 2019 of the family Cerophytidae from mid- Cretaceous Burmese amber is described and illustrated. The new species, A. maculatum s.n. differs from A. birmanicum Yu, Slipinski et Pang, 2019 in the smaller body size, shorter metatarsomere 1, and more convex body. A key to the species of the genus Amberophytum is presented.

Tropidogyne euthystyla, described here, is the fourth species of this fossil genus to have been reported from amber deposits in northern Myanmar. The species are alike in features of the calyx, the shape and venation of the inferior ovary, and the absence of petals. They differ in the number and form of the styles, the lobing of the epigynous disc, and whether the flowers are apparently bisexual or unisexual. In the one species for which several flowers are available for study, T. pentaptera, floral diameters vary from 3.5 to 5.0 mm. The present species, known only from a single flower, is the smallest in the genus, with a floral diameter of only 2 mm. It differs from the 3 previously described species in having 2 stout, erect styles bearing a terminal stigma, whereas the other species have 2 or 3 short or long, arching styles that are decurrently stigmatic along the adaxial surface.

A new genus and species of froghopper, Araeoanasillus leptosomus gen. et sp. nov. (Hemiptera: Cercopoidea, Sinoalidae?), is described from mid-Cretaceous Burmese amber. The new genus possesses the following diagnostic characteristics: slender, medium size body (length, 7.0 mm) with head longer than wide, round eyes; antennae slender with eight antennomeres; pedicel very short, shorter than scape; pronotum with a length/width ratio of 2.4; metatibia with three spines, including one short spine near base and two adjacent, long, thick spines near apex; a single series of 16 thick apical teeth (comb) at metatibial apex; tegmen narrow with a length/width ratio of 3.2; tegmen with coastal area and stigmal cell punctate; CuP meeting base of CuA2; and MP branching at middle of wing. In hind wing, Cu vein forked once. A series of plant trichomes adjacent and attached to the specimen suggests that the froghopper’s host plant was a fern.

A new monotypic genus of nogodinid planthoppers (Hemiptera: Fulgoromorpha: Nogodinidae) is described from Dominican amber. The adult female of Jatoba losbrachi gen. et sp. nov. can be distinguished...

The present paper describes a new species of encyrtid wasp, Hambletonia dominicana sp. nov. (Hymenoptera: Encyrtidae) from Dominican amber. Diagnostic characters include long antennae that are almost twice the length of the head, the absence of long, coarse, flattened setae on the dorsal apical margin of the pedicel, and an elliptical club subequal in length to that of the funicle. This first described fossil species of Hambletonia provides views of new and possible unique morphological features of encyrtid wasps that existed in the Neotropics during the mid-Tertiary.

A new species of spiroplasmid, Spiroplasma burmanica sp. nov. (Mollicutes: Entomoplasmatales: Spiroplasmataceae) is described from the body cavity of a fossil plant louse (Psylloidea: Sternorrhyncha) in Burmese amber. The new species is pleomorphic with body shapes varying from oval to helical. The majority of the helical cells occur in the head, thorax (including leg cavities) and abdomen of the fossil psyllid. The association between S. burmanica and the psyllid is considered to be a case of symbiosis, similar to extant relationships. This discovery of the first fossil spiroplasmid shows that psyllids carried these microorganisms some 100 million years ago.

A bizarre cylindrical bark beetle from mid-Cretaceous Burmese amber is described as Stegastochlidus saraemcheana, a new genus and species in the subfamily Colydiinae of the family Zopheridae. The male beetle is characterized by elongate protuberances covering its entire dorsal surface, a tarsal formula of 4-4-4 and ten-segmented antennae with the terminal segment expanded into a small club. The fossil is considered to have been a possible predator that lived among moss, lichens and fungi either attached to trees trunks or on the forest floor. A close association with fungi is indicated by strands of conidia attached to the cuticle of the beetle.

Although partially damaged prior to its preservation in coniferous resin, the flower of Cyathitepala papillosa retains enough morphological features to justify its description as a new genus and species of fossil angiosperms. The perianth is incomplete, but there are at least 6 large outer tepals and numerous smaller, spirally arranged inner ones. The limbs of some of the larger tepals are cup-shaped and face abaxially, while the smaller inner tepals are flat or have adaxially folded margins. There is an inner series of erect, apically truncate tepals surrounding the stamens. The surfaces and margins of the tepals are densely papillate. There is a whorl of ca. 12 stamens with anthers that dehisce by two dorsally hinged valves. Nothing remains of the receptacle or pedicel of the flower, but the superior portion of the pistil is conic and bears two columnar, erect styles. The fossil is compared with three previously described fossil genera, Setitheca, Zygadelphus, and Strombothelya, which were suggested as having a possible relationship with certain families of order Laurales.

A flower preserved in 100 Ma amber from Myanmar is described here as Chainandra zeugostylus, a new genus and species of fossil angiosperms. The anthers in Chainandra are sagittate at the base and have only a short connective. They dehisce by means of a circumferential stomium, the dorsal and ventral sides of each locule forming flaps that become widely separated. The style of Chainandra is columnar and two-branched above the middle, and the fully inferior ovary bears an epigynous nectar disc but lacks well-defined ribs or veins. In Tropidogyne, Lacknociona, and Strombothelya, similar genera described earlier from the same am-ber deposits, the styles are either unbranched or are 3–5-branched to near the base, and the ovary is half-inferior to inferior, its lower portion often being dis-tinctly veined and ribbed. Morphology of the stamens and gynoecium of the three genera suggests that they are early members of the eudotyledons.

The flower described here as Valviloculus pleristaminis, a new genus and species of fossil angiosperms, was obtained from Myanmar amber deposits dating to the mid-Cretaceous period. The flower is staminate, with an ovoid, hollow floral cup. The perianth consists of 6 tepals, one of which was lost prior to preservation. Numerous helically arranged stamens are borne at the summit of the flower. The anthers are bilocular, with pollen sacs that dehisce by laterally hinged valves. The tip of the connective bears a cup-shaped appendage. The gynoecium is represented only by a cluster of vestigial styles centered among the stamens. The fossil is provisionally assigned to order Laurales, its closest affinities being with the families Monimiaceae and Atherospermataceae.

A new weevil genus, Tanyrhamphis gen. nov. (type species: Tanyrhamphis rhadinoalibus sp. nov.), of the subfamily Conoderinae in the family Curculionidae, is described from Dominican amber. The new ...

Three new species of ensign wasps (Hymenoptera: Evaniidae), Evaniella setifera sp. nov., Evaniella dominicana sp. nov., and Semaeomyia hispaniola sp. nov. are described from Dominican amber and Hyp...

A fossil flower in mid-Cretaceous amber from Myanmar (Burma) is described as the new genus and species Phantophlebia dicycla. The perianth of the flower is composed of two 5-merous whorls of tepals, the outer tepals being smaller and more irregular in shape than the inner. All the tepals are distinctly vascularized, with few to many major veins that are connected above by an irregular network of veinlets. The flower’s 5 stamens are opposite the inner tepals, with bilocular anthers that dehisce by latrorse, oblique slits. No gynoecium can be seen beneath the air bubble covering the center of the flower. The fossil shares some characteristics with myrsinoid members of family Primulaceae.

The fossil flower described here as Thymolepis toxandra is from amber deposits in Myanmar which have been dated as ca. 100 Ma. The peri-anth consists of 12 tepals of varying size and shape, arranged in decussate pairs at the summit of an obconic floral cup. The epidermis of some of the tepals is densely papillate-secretory. The flower is bisexual, the androecium consisting of 2 stamens whose anthers are bisporangiate, hippocrepiform, and densely hirsute. The gynoecium is visible only as a bilobed stigma or as the tips of two postgenitally fused styles, the re-mainder of the pistil(s) being hidden by the perianth and floral cup. It is suggested that the fossil may be an early representative of family Monimiaceae.

A new fossil angiosperm, Dasykothon leptomiscus, is described from mid-Cretaceous amber deposits in the Hukawng Valley of northwestern Myanmar (Burma). The flower is post-anthesis except for one stamen that retains its pollen. The perianth is slightly irregular and composed of 5 ovate or lance-linear, erect or incurved sepals. The ca. 12 stamens have slender, elongate filaments and dorsifixed, bithecal, longitudinally dehiscent anthers. The superior ovary bears 2 long, curved styles. Germinating pollen grains observed on the stigmas are monoporate and have the peculiar feature of a grooved ridge encircling the grain from pole to pole. This structure is hypothesized to be an evolutionary elaboration of the meridionally syncolpate sulcus found in some members of the Atherospermataceae. The generalized floral morphology of the fossil prevents ready taxonomic assignment to a modern family.

Summary A new species of fossil mermithid, Heydenius simulphilus sp. n. (Nematoda: Mermithidae), is described from two parasitic juvenile specimens adjacent to a male black fly (Diptera: Simuliidae) in Baltic amber. It is proposed that the nematodes emerged from their developmental sites in the haemocoel of the black fly host through a wound in the abdomen of the latter, as indicated by the release of a droplet of haemolymph and damaged cuticle. Various internal structures of the nematodes are identified and related to those found on extant developing mermithids. This is the first fossil record of mermithid parasitism of a black fly.

A new species, Anthonomus aphelomerus Legalov and Poinar, sp. nov. from Early Miocene Dominican amber is described. The new species differs from Anthonomus browni Poinar and Legalov, 2015 in the smaller body size, thinner, weakly curved rostrum and weakly clavate femora; it is differs from Anthonomus clarki Legalov, 2019 in the longer rostrum and slightly convex body; it also differs from Anthonomus cruraluma Poinar and Legalov, 2015 in the profemora with one tooth, and the longer and weaker curved rostrum. A key to species of the genus Anthonomus from Dominican amber is given.

The present work investigates the mining and sale of fossiliferous Burmese amber to determine if the profits are being used by the Myanmar military to commit atrocities against minority groups or ethnic armies within the country, as recently implied. Our conclusion, based on the information available to us, is that the great majority of Burmese amber fossils up to 2016 were taken across the border and sold in China. At this time, the amber mines were under the control of the Kachin Independent Army (KIA) or unregulated. After heavy fighting between the KIA and the Myanmar military from June 2017 to June - July 2018, the amber mines in the Hukawng Valley were occupied by the Myanmar government and presently, most of these mines are closed or are operated illegally or worked only intermittingly on a small scale. We have not been able to substantiate claims that millions of dollars from the sale of Burmese amber fossils were used by the Myanmar government to dominate minority groups within the country. At least up to 2017, there were no, or only insignificant funds, received by the Myanmar government from the mining and sale of Burmese amber. Thus we see no reason to halt the purchase of Burmese amber fossils or for editors and publishers to reject scientific papers describing these rare inclusions, which otherwise would be used for carvings or jewelry and become lost to science.

A new genus and species of Empididae (Diptera) in mid-Cretaceous Burmese amber is described as Neoalavesia hadroceria gen. et sp. nov. in the family Atelestidae. The male is characterised by its sm...

Presently, three families of Coleoptera (Meloidae, Ripiphoridae and Cleridae) produce triungulin larvae that parasitize aculeate Hymenoptera, especially various lineages of social and solitary bees, as well as wasps and other insects. The discovery of a fossil bee with associated beetle triungulins in mid-Cretaceous Burmese amber provides the earliest known date for the co-evolution between bees and beetle triungulins, specifically those of the family Cleridae (Coleoptera). The fossil bee has been described in a new family and the beetle triungulins are described in the present work in the form genus and species Anebomorpha cercorhampha gen. et sp. nov. (Coleoptera: Cleridae). The description of Anebomorpha cercorhampha is based on 21 triungulins of the family Cleridae, five of which are in direct contact with the primitive bee in the amber. The remainder are at various distances behind the bee. This is the earliest fossil evidence of co-evolution between beetle triungulins and a member of the aculeate Hymenoptera.

A new family, genus and species of scale insect is described from mid-Cretaceous Burmese amber. The male specimen of Macrodrilus bostrychus gen et sp. nov. in the new family Macrodrilidae (Hemipter...

Crickets (Orthoptera: Grylloidea) are a highly diverse and successful group that due to their chirping are often heard more often than they are seen. Their omnivorous diet allows them to exist in a variety of terrestrial habitats around the world. In some environments, cricket populations can build up and become plagues, resulting in significant damage to seedling crops. A new genus and species of cricket, Pherodactylus micromorphus gen. et sp. nov. (Orthoptera: Gryllidae) is described from mid-Cretaceous Myanmar amber. The new genus is characterized by the following features: head without prominent bristles, pronotum longer than wide, middle of pronotal disk with two distinct large dark “eyespots”, fore leg robust and 3 apical spurs arranged on inner side of fore leg tibia. Shed portions of a lizard skin adjacent to the specimen reveal possible evidence of attempted predation.

Ophiocordyceps dominicanus Poinar & Vega sp. nov. in Dominican amber and Polycephalomyces baltica Poinar & Vega sp. nov. (Hypocreales: Ophiocordycipitaceae) in Baltic amber are described as entomopathogenic fungi of bark lice (Psocoptera). The specimens possess several features unknown in extant synnematous entomopathogenic fungi such as a tubular dark synnema with a straight, pointed tip bearing spores over the entire surface in O. dominicanus, and a globular yellowish synnema developing on the tip of the host’s antenna in P. baltica. These are the only known fossil entomopathogenic fungi of bark lice, making them unique not only for their characters but also in respect to their selection of developmental sites on their bark lice hosts.

Male and female thalli of a mid-Cretaceous fungus attached to a thrip (Thysanoptera) in Burmese amber are described as Philothysanus burmanicus gen. et sp. nov. in the family Spheciophilaceae (Ascomycota). Both spermatia and ascospores are present. Currently, Philothysanus burmanicus gen. et sp. nov., together with the previously described Spheciophilia adercia Poinar, represent the oldest known ectoparasitic fungi of insects and their age can be used as a reference point in future phylogenetic studies on the origin of fungi with this life style.

A tylenchid nematode parasite of a male long-legged fly, Tachytrechus sanus Osten Sacken (Diptera: Dolichopodidae) from Montana, USA is described as Parasitylenchus myiophagus n. sp. (Nematoda: Parasitylenchidae). The new species is characterized by the presence of an extremely long first generation female and numerous short and wide second generation males and females produced in “enclosed” clusters in the host’s body cavity. Both generation female nematodes are ovoviviparous, with short stylets lacking knobs and simple tails lacking spikes, palps or mucrons. The second generation males have paired, separate spicules, short stylets, and a bursa but no visible gubernaculum. The fly host shows evidence of demasculinization, which is attributed to nematode parasitism. The gonads of the second generation adults are infected with a microsporidium (Microsporidia), which is a new host record for tylenchid nematodes. A tylenchid-infected Baltic amber dolichopodid shows that associations between these two organisms extend back at least to the Eocene.

The fossil record of ambrosia beetles is summarized and a new genus and species in the subfamily Mecopelminae, Gongyloceria dominicana gen. et sp. nov. is described. The new genus differs from the extant genus Mecopelmus Blackman, 1944, possessing weakly elongated tarsi, finely faceted eyes, a scape not reaching the posterior margin of the eye, coarsely sculptured pronotum and elytra, and a larger body size. It is the first Miocene record of the Mecopelminae. The genus Xyleborites Wickham, 1913, placem. n. is transferred from the Scolytidae to the Platypodidae. A list of the fossil Platypodidae, including a key to the subfamilies and tribes of ambrosia beetles, is presented. Xyleborites longipennis Wickham, 1913 and Gongyloceria dominicana sp. nov. may have been related to Paullinieae recorded from the late Eocene and the early Miocene of North and Central America. Distribution maps with fossil records for ambrosia beetles are included.

A new fern, Cladarastega burmanica gen. et sp. nov. (Dennstaedtiaceae: Polypodiales) is described from a fertile pinnule in Burmese amber. The new species has dentate margins with acute apices, furcated 1 and 2 forked free veins angled toward pinnule apex, elongate, multicellular and glandular hairs on the epidermis, epidermal peltate scales, sori roundish, apical and subapical on abaxial surface of fronds near vein endings and paraphyses. There are both inner and outer cup-shaped indusia. The inner indusium is initially solid but then disintegrates while still attached to the sorus. The sporangium is stalked with a vertical or slightly oblique annulus. The spores are tetrahedral trilete. Related topics discussed are phylogenetic studies on ages pertaining to the origin of the Dennstaedtiaceae in comparison to the age of the fossil, associations with competing angiosperms and insect herbivores of ferns.

The subterranean termite Reticulitermes flavipes (Kollar, 1837) (Isoptera: Rhinotermitidae) occurs throughout the eastern USA and populations have been found as far west as Oregon. As with other termites, eastern subterranean termites live in colonies composed of workers, soldiers and reproductives. The blind, wingless workers, which are only between 3 and 4 mm long, feed on wood and can cause serious damage to wooden structures throughout their range (Edwards & Mill, 1986). Mermithid parasitism of termites is a rare occurrence and this report represents the first case of mermithid parasitism of termites in North and Central America.

The flower described here as Zygadelphus aetheus gen. et sp. nov. was obtained from amber mines in Myanmar, preserved in marine sedimentary deposits dating to the mid-Cretaceous Period, ca. 99 My in age. The perianth consists of ca. 10 spirally arranged tepals that vary in size and shape. The stigmatic tips of two styles are visible, the remainder of the gynoecium being hidden from view by the crowding of stamens and perianth parts toward the center of the flower. There is a whorl of 4 stamens, the anthers of which have the feature, probably unique among angiosperms, of possessing a small but complete accessory stamen arising as an appendage on their dorsal surface. Anthers of both types of stamens are bilocular and dehisce by dorsally hinged valves. Presence of pollen in the accessory anthers indicates that they functioned in reproduction. Under the assumption that the fossil is not merely a teratological mutant in an otherwise normal-flowered species, it is here given taxonomic status as representing a peculiar type of floral development, now extinct, that existed during an earlier stage of angiosperm evolution.

The fossil flower described here is the third species of Tropidogyne to have been collected from mid-Cretaceous amber deposits in the Hukawng Valley of northwestern Myanmar. The flower of Tropidogyne lobodisca differs from the 2 previously described species, T. pikei and T. pentaptera, in lacking stamens and having a 5-lobed nectar disc covering the apex of the ovary. Its 2 slender, curved, attenuate styles are like those of T. pentaptera in being stigmatic along the adaxial surface. The new species has 5 spreading, reticulately-veined sepals, a generic character of Tropidogyne. An unusual, probably teratological, feature is the presence of 2 sepal-like staminodes on one side of the flower, inserted at the base of the nectar disc where stamens would otherwise be found. The inferior portion of the pistil is obconic, with 5 distinct veins that connect to the mid-nerves of the sepals.