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This is a data set of monthly averaged variables simulated by the hydrostatic regional atmospheric climate model RACMO2.3p2 over the Antarctic Peninsula. At the lateral and ocean boundaries the model is forced by ERA-Interim reanalysis data every 6 hours from January 1979 to August 2019. After August 2019, the model is forced by ERA5 reanalyses every 3 hours, with an overlap from 2016, to December 2022. The model is run at a horizontal resolution of 5.5 km and 40 vertical levels for the Antarctic Peninsula domain, which, after 2018, constitutes an update of the simulation forced from 1979-2018 by ERA-Interim reported in van Wessem et al., 2018. Upper air relaxation of wind, humidity and temperature is not active for this domain. This version of the model is specifically applied to the polar regions by interactive coupling to a multilayer snow model that calculates melt, refreezing, percolation and runoff of meltwater (Ettema et al., 2010). In addition, snow albedo is calculated through a prognostic scheme for snow grain size (Kuipers Munneke et al., 2011) while a drifting snow scheme simulates the interaction of the near-surface air with drifting snow (Lenaerts et al., 2010). This dataset is provided on a rotated polar coordinate grid. In such a rotated pole projection the grid is defined over the equator and then rotated to the area of interest. One of the advantages is that the grid distance can be defined in fraction of degrees, which results in near equidistant grid cells as long as the domain is small enough, and provides the most accurate model calculations. However, re-projecting these data on other grids is often troublesome, as after rotation the grid is non-equidistant and most software packages cannot directly handle this. Stef Lhermitte provided a nice solution for reprojecting the RACMO data on his gitlab-page: https://gitlab.tudelft.nl/slhermitte/manuals/blob/master/RACMO_reproject.md. The dataset includes the following surface- and atmospheric variables. Additional variables and higher temporal resolutuon up to 3 hourly are available on request: Surface mass balance (SMB) variables (in kg m-2 mo-1 or mm water equivalent mo-1) smb : (Specific) Surface mass balance defined as SMB = Total precipitation + sublimation - runoff snowmelt : Surface snowmelt production refreeze : Refreezing of meltwater snowfall : Solid precipitation precip : Total precipitation (snowfall + rainfall); to calculate rainfall use rainfall = precip - snowfall runoff : Surface meltwater runoff subl : Snow sublimation (including sublimation of drifting snow). Negative values are sublimation, positive values are snow deposition. erds : erosion of drifting snow Atmospheric variables t2m : 2-m Temperature q2m : 2-m Specific humidity rh2m : 2-m Relative humidity (RH) tskin : Surface/skin temperature. Calculated from closing the surface energy budget. psurf : Surface pressure u10m : Zonal wind speed at 10 m v10m : Meridional wind speed at 10 m ff10m : Wind speed at 10 m u0500 : Zonal wind speed at 500 hPa v0500 : Meridional wind speed at 500 hPa z0500 : Geopotential height at 500 hPa Surface Energy Budget (SEB) variables (in J m-2); SEB = LWnet+SWnet+SHF+LHF+GHF Values are monthly cumulative: to convert to W m-2 divide by amount of seconds in a month: 30*24*3600. lwsn : Net longwave radiation (LWnet=LWdown-LWup) swsn : Net shortwave radiation (SWnet=SWdown-SWup) lwsd : Downwelling longwave radiation at the surface swsd : Downwelling shortwave radiation at the surface swsu : Upwelling shortwave radiation at the surface senf : Upward Sensible Heat Flux (SHF) at the surface latf : Upward Latent Heat Flux (LHF) at the surface (our simulated LHF doesn't explicitly close the SEB, as it also includes in-air sublimation, but the effect should be rougly neglible) gbot : Soil/Ground Heat Flux (GHF) Snow variables totpore : Vertically integrated pore space (m) totwat : Total liquid water content of the snowpack (kg m-2) zsnow : Total snowpack thickness (m) Grid, elevation, coordinates and masks in Height_latlon_ANT27.nc (240 by 262 grid boxes) mask2d : Full ice mask fraction (grounded ice + floating ice shelves) [0..1] maskgrounded2d : Grounded ice sheet mask fraction [0..1] height : Surface elevation (m) slope : Surface slope (m m-1) aspect : Direction of surface slope (degrees) lat : Latitude (polar) lon : Longitude (polar) Ice shelf and ice sheet drainage basins mask in TotIS_RACMO_XPEN055_IMBIE2.nc This file contains masks on the RACMO grid for the drainage basins as defined in http://imbie.org/imbie-3/drainage-basins/ (Rignot et al., 2013, IMBIE2, IMBIE3), including masks seperately for the ice shelves they drain into, numbered counterclockwise from 0 to 18. mask2dF : Full ice mask including ice shelves IceShelves : Ice shelf masks GroundedIce : Grounded ice sheet drainage basins, {"references":["Van Wessem, J. M., van de Berg, W. J., Noël, B. P. Y., van Meijgaard, E., Birnbaum, G., Jakobs, C. L., et al. (2018). Modelling the climate and surface mass balance of polar ice sheets using RACMO2, part 2: Antarctica (1979–2016). The Cryosphere, 2018, 1–35. doi:10.5194/tc-2017-202. issn:1994-0440.","Ettema, J., Van den Broeke, M. R., Van Meijgaard, E., Van de Berg, W. J., Box, J. E., and Steffen, K. (2010). Climate of the Greenland ice sheet using a high-resolution climate model – Part 1: Evaluation. The Cryosphere, 4(4), 529–544. doi:10.5194/tc-4-511-2010. issn:1994-0424.","Lenaerts, J. T. M., van den Broeke, M. R., Déry, S. J., König-Langlo, G., Ettema, J., Munneke, P. K., and Kuipers Munneke, P. (2010). Modelling snowdrift sublimation on an Antarctic ice shelf. The Cryosphere, 4(2), 179–190. doi:10.5194/tc-4-179-2010. issn:1994-0424.","Rignot, E., Jacobs, S., Mouginot, J., and Scheuchl, B. (2013). Ice-Shelf Melting Around Antarctica. Science, 341(6143), 266–270. doi:10.1126/science.1235798. issn:0036-8075."]}

Aerial frames and satellite imagery are widely recognized data sources from which to produce maps. For volcanoes, maps enable the quantification of erupted ash and the destruction caused. The last eruptive sequence on Deception Island was endured from 1967 to 1970. Analogue maps were produced via classical photogrammetric methods with a high degree of human intervention mainly to analyse the volcanic-centres areas only. However, historical aerial frames cover the whole of Deception Island. Structure from motion photogrammetry, a near-automated compilation of digital image processing strategies, minimizes the degree of human intervention to produce orthographic mosaics and digital elevation models from digital aerial frames. Orthographic mosaics were produced from historical aerial frames of 1956 and 1968, and a Kompsat-3 image of 2020. Their shared rootmean-square deviation was 1.8 m and 1.7 m in easting and northing, respectively, at ground control points measured with phase-differential global navigation satellite systems. The digital elevation models were processed with a root-mean-square deviation of 2.3 m and 3.6 m from 1956 and 1968 aerial frames, respectively. As the first application, erupted ashfall and the subsequent destruction, mainly at the former Chilean and British bases, were identified, and the volume of erupted ash was assessed to be over 0.16 km3 within the area mapped by these new digital cartographic products. info:eu-repo/semantics/publishedVersion

Subduction zones and their associated slabs are the main drivers of plate tectonics and mantle flow, but how these zones initiate remains enigmatic. In the Scotia Sea region, subduction started in the Late Cretaceous/Early Cenozoic in a pristine ocean basin setting devoid of other subduction/collision zones. How this subduction zone initiated remains intensely debated, as exemplified by the variability of published plate tectonic reconstructions. Despite such variability, several works argue for a subduction initiation mechanism, in which a South America-Antarctica relative plate motion change, in combination with a particular plate boundary geometry in the western Weddell Sea, caused convergence across a transform plate boundary segment that subsequently evolved into a subduction zone. Here we discuss this kinematic model of subduction initiation, and, following geometric and kinematic arguments, highlight several unsolved issues that call for alternative explanations. Furthermore, we present new tectonic reconstructions of the Scotia region involving a simpler middle-Late Cretaceous plate boundary configuration, which avoid the geometric and kinematic problems of earlier reconstructions and that call for a new mechanism of subduction initiation. We refer to this mechanism as Subduction Invasion Polarity Switch (SIPS), which involves a long-lived and wide subduction zone (South American-Antarctic subduction zone) with lower mantle slab penetration, which imposes major horizontal trench-normal compressive deviatoric stresses on the overriding plate. This plate consists of a narrow continental lithospheric (land) bridge at the trench (Cretaceous-Early Cenozoic Antarctica-South America land bridge) with oceanic lithosphere behind it (Weddell Sea-Atlantic Ocean). The stresses cause shortening and thrusting at the continent-ocean boundary in the backarc region of the overriding plate, forcing oceanic lithosphere under continental lithosphere, starting the subduction initiation process, and eventually leading to a new, self-sustaining, subduction zone (Scotia subduction zone) with an opposite polarity compared to the long-lived subduction zone. The model thus involves invasion of a new subduction zone into a pristine ocean basin (Atlantic Ocean), with the primary driver being a long-lived subduction zone in another ocean basin. To test the physical viability of the SIPS model, we have conducted numerical geodynamic simulations of buoyancy-driven subduction. Numerical results demonstrate that the SIPS model is viable, with compressive stresses in the overriding plate resulting from strong trenchward basal drag induced by subduction-driven whole-mantle poloidal return flow and compression at the subduction zone plate boundary. Subduction initiation starts in the overriding plate after ∼100 Myr of long-lived subduction, eventually resulting in the formation of a new, opposite-dipping, subduction zone. This new subduction zone develops at the continent-ocean boundary for models without and with a pre-imposed weak zone. We further propose that the SIPS model might explain subduction initiation elsewhere, including the New Caledonia subduction zone in the Southwest Pacific, the Lesser Antilles-Puerto Rico subduction zone in the Caribbean region, and the subduction zones that consumed the Rocas Verdes and Arperos backarc basins in South America and Central America, respectively. We further postulate that active backarc shortening in the Japan Sea, with eastward under-thrusting of Japan Sea oceanic lithosphere below the Japan arc, represents an early stage of SIPS.

Runoff in polar areas is strongly influenced by rising air temperature because these regions experience the largest air temperature increases in the world. Rising air temperatures affect permafrost, snow cover and glaciers which are the main components of runoff in high latitude regions. Warming is also causing the shift in precipitation phase from solid (snow) to liquid (rain) as well as changes in annual precipitation its distribution. With reducing extent of the snow and ice cover, the energy balance of the Earth's surface is fundamentally affected as the albedo is reduced, which leads to further warming of the Earth's surface. Changes in snow and ice cover lead to significant positive feedback loops in the climate cycle. This study includes analyses of meteorological and hydrological data measured in James Ross Island during the 2018 austral summer. Meteorological data were measured at eight automatic weather stations, and hydrological data were measured in seven catchments. However, in this thesis the analyses are focused on two selected catchments: the partly glaciated Triangular and the glacier-free Bohemian catchments. The influence of different meteorological variables on the catchment runoff was analysed using correlation and regression analyses. The analyses showed the highest...

To study how Odontaster validus can influence the spatial structure of Antarctic benthic communities and how they respond to disturbance, it is necessary to assess potential dietary shifts in different habitats. We investigated the diets of O. validus from Maxwell Bay and South Bay in the West Antarctic Peninsula. A multifaceted approach was applied including in situ observations of cardiac stomach everted contents, isotopic niche, and trophic diversity metrics. Results confirm the flexible foraging strategy of this species under markedly different environmental conditions, suggesting plasticity in resource use. The data also showed evidence of isotopic niche expansion, high δ15N values, and Nacella concinna as a common food item for individuals inhabiting a site with low seasonal sea ice (Ardley Cove), which could have significant ecological implications such as new trophic linkages within the Antarctic benthic community. These results highlight the importance of considering trophic changes of key species to their environment as multiple ecological factors can vary as a function of climatic conditions.

The Antarctic Peninsula (AP) is one of the widely studied polar regions because of its sensitivity to climate change and potential contribution of its glaciers to global sea level rise. Precise digital elevation models (DEMs) at a high spatial resolution are much demanded for investigating the complex glacier system of the AP at fine scales. However, the two most recent circum-Antarctic DEMs, the 12 m TanDEM-X DEM (TDM DEM) from bistatic interferometric synthetic aperture radar (InSAR) data acquired between 2013 and 2014 and the Reference Elevation Model of Antarctica mosaic (REMA mosaic) at an 8 m spatial resolution derived from optical data acquired between 2011 and 2017 have specific individual limitations in this area. The TDM DEM has the advantage of good data consistency and few data voids (approx. 0.85 %), but there exist residual systematic elevation errors such as phase-unwrapping errors in the non-edited DEM version. The REMA mosaic has high absolute vertical accuracy, but on the AP it suffers from extended areas with data voids (approx. 8 %). To generate a consistent, gapless and high-resolution topography product of the AP, we fill the data voids in the TDM DEM with newly processed TDM raw DEM data acquired in austral winters of 2013 and 2014 and detect and correct the residual systematic elevation errors (i.e., elevation biases) in the TDM DEM with the support of the accurately calibrated REMA mosaic. Instead of a pixelwise replacement with REMA mosaic elevations, these provide reference values to correct the TDM elevation biases over entire regions detected through a path propagation algorithm. The procedure is applied iteratively to gradually correct the errors in the TDM DEM from a large to small scale. The proposed method maintains the characteristics of an InSAR-generated DEM and is minimally influenced by temporal or penetration differences between the TDM DEM and REMA mosaic. The performance of the correction is evaluated with laser altimetry data from Operation IceBridge and ICESat-2 missions. The overall root mean square error (RMSE) of the corrected TDM DEM has been reduced from more than 30 m to about 10 m which together with the improved absolute elevation accuracy indicates comparable values to the REMA mosaic. The generated high-resolution DEM depicts the up-to-date topography of the AP in detail and can be widely used for interferometric applications as well as for glaciological studies on individual glaciers or at regional scales.

In this study, we used the stable water isotope record (δ18O) from an ice core drilled in Palmer Land, southern Antarctic Peninsula (AP). Utilizing δ18O we identified two climate regimes during the satellite era. During the 1979–1998 positive interdecadal Pacific oscillation (IPO) phase, a low-pressure system north of the Weddell Sea drove southeasterly winds that are associated with an increase in warm air mass intrusion onto the Larsen shelves, which melted and a decreased sea ice concentration in the Weddell Sea/increase in the Bellingshausen Sea. This climate setting is associated with anomaly low δ18O values (compared with the latter IPO period). There is significantly more melt along the northern AP ice shelf margins and on the Larsen D and southern Larsen C during the 1979–1998 IPO positive phase. The IPO positive climatic setting was coincidental with the Larsen A ice shelf collapse. In contrast, during the IPO negative phase (1999–2011), northerly winds caused a reduction in sea ice in the Bellingshausen Sea/Drake Passage region. Moreover, a Southern Ocean north of the Weddell Sea high-pressure system caused low-latitude warm humid air over the tip and east of the AP, a setting that is associated with increased northern AP snowfall, a high δ18O anomaly, and less prone to Larsen ice shelf melt.

The western Antarctic Peninsula is facing rapid environmental changes and many recent publications stress the need to gain new knowledge regarding ecosystems responses to these changes. In the framework of the Belgica 121 expedition, we tested the use of a nimble vessel with a moderate environmental footprint as an approach to tackle the urgent needs of the Southern Ocean research community in terms of knowledge regarding the levels of marine biodiversity in shallow areas and the potential impacts of retreating glaciers on this biodiversity in combination with increasing tourism pressure. We discuss the strengths and drawbacks of using a 75’ (23 m) sailboat in this research framework, as well as its sampling and environmental efficiency. We propose that the scientific community considers this approach to 1) fill specific knowledge gaps and 2) improve the general coherence of the research objectives of the Antarctic scientific community in terms of biodiversity conservation and the image that such conservation conveys to the general public.

This study presents a 1:25,000 geomorphological map of the northern sector of Ulu Peninsula, James Ross Island, Antarctic Peninsula. The map covers an area of c. 250 km(2), and documents the landforms and surficial sediments of one of the largest ice-free areas in Antarctica, based on remote sensing and field-based mapping. The large-scale landscape features are determined by the underlying Cretaceous sedimentary and Neogene volcanic geology, which has been sculpted by overlying ice masses during glacial periods. Paraglacial and periglacial features are superimposed upon remnant glacial features, reflecting the post-glacial evolution of the landscape. The study area can be broadly separated into three geomorphological sectors, according to the dominant contemporary Earth-surface processes; specifically, a glacierised southern sector, a paraglacial-dominated eastern sector, and a periglacial-dominated central/northern sector. This map provides a basis for further interdisciplinary research, and insight into the potential future landscape evolution of other parts of the Antarctic Peninsula as the climate warms.

Investigating precipitation phase transitions is crucial for improving our understanding of precipitation formation processes and impacts, particularly in Polar Regions. This study uses observational data and numerical modelling to investigate precipitation phase transitions in the western and northern Antarctic Peninsula (AP) during austral summer. The analysis is based on the ERA5 reanalysis product, dynamically downscaled using the Polar-WRF (Polar Weather Research and Forecasting) model, evaluated using regular meteorological observations and additional measurements made during the Year of Polar Prediction special observing period. We analyse three cases of extra-tropical cyclones bringing precipitation with phase transitions, observed at the Chilean station Professor Julio Escudero (King George Island, north of the AP) and the Ukrainian Antarctic Akademik Vernadsky station (western side of the AP) during the first week of December 2018. We use observed and modelled near-surface air temperature and pressure, precipitation amount and type, and vertical temperature profiles. Our results show that precipitation type (snow or rain) is well-represented by ERA5 and Polar-WRF, but both overestimate the total amount of precipitation. The ERA5 daily variability and vertical air temperature profile are close to the observed, while Polar-WRF underestimates temperature in the lower troposphere. However, ERA5 underestimates the temperature inversion, which is present during the atmospheric river event, while Polar-WRF represents that inversion well. The average weekly temperature, simulated with Polar-WRF, is lower compared to ERA5. The Polar-WRF fraction of snow in the total precipitation amount is higher than for ERA5; nevertheless, Polar-WRF represents the precipitation phase transition better than ERA5 during the event, associated with an atmospheric river. These case studies demonstrated a relationship between specific synoptic conditions and precipitation phase transitions at the AP, evaluated the ability of the state-of-the-art reanalysis and regional climate model to represent these events, and demonstrated the added value of combined analysis of observations from the western and northern AP, particularly for characterizing precipitation during synoptic events affecting the entire AP.

February 2020 was anomalously warm in the Antarctic Peninsula region and registered one of the most intense heatwaves ever recorded in Western Antarctica. The event featured unprecedented regional mean temperature anomalies (+4.5¿°C) over the Antarctic Peninsula between 6 and 11 February 2020 and the highest local temperature of the continental Antarctic region. Taking flow analogs of the event from past (1950¿1984) and recent (1985¿2019) periods of the ERA5 reanalysis, here we quantify the role of recent climate change in the magnitude of this 6-day regional heatwave. Results show that 2020-like heatwaves over the Antarctic Peninsula are now at least ~0.4¿°C warmer than in the past period, which represents a ~25% increase in magnitude. Given the observed atmospheric circulation conditions, the probability of experiencing 6-day regional mean anomalies above ~2¿°C has increased ten times since 1950¿1984. The aggravated severity of the event can be largely ascribed to long-term summer warming of the Antarctic Peninsula rather than recent atmospheric circulation trends., We thank David Bromwich for his useful comments and discussions on this research. We want to acknowledge the contribution of the two anonymous reviewers, which provided constructive comments that helped to substantially improve the final form of this manuscript. The AEMET Antarctic program is supported by the Ministerio para la Transición Ecológica y Reto Demográfico and Ministerio de Ciencia e Innovación of the Spanish Goverment. Research activities of S.G. and M.O. are partly funded by ANTALP Research Group, Generalitat de Catalunya. S.G. is supported by the Ministerio de Ciencia e Innovación project – PID2020-116520RB-I00 (MICROAIRPOLAR2) and RTI2018- 098693-B-C32 (WISE-PreP). M.O. is supported by the Ramón y Cajal Program (RYC2015-17597) and the NUNANTAR (PTDC/CTA-GFI/32002/2017) from the Fundação para a Ciência e a Tecnologia of Portugal. D.B. acknowledges support from the H2020 EU project CLINT (Grant Agreement No. 101003876). R.M.T. was supported of the Portuguese Foundation for Science and Technology through the FCT – project JPIOCEANS/0001/2019 (ROADMAP). This research is part of the CSIC Interdisciplinary Thematic Platform (PTI) Clima y Servicios Climáticos (PTI-CLIMA) and POLARCSIC (PTI-POLAR) activities.

Antarctica and the Southern Ocean are unique natural laboratories where organisms adapted to extreme environmental conditions have evolved in isolation for millions of years. These unique biotic communities on Earth are facing complex climatic and environmental changes. Terrestrial ecosystems in the Antarctic Peninsula Region (APR) have experienced the highest rate of climate warming and, being the most impacted by human activities, are facing the greatest risk of detrimental changes. This review provides an overview of the most recent findings on how biotic communities in terrestrial ecosystems of the Antarctic Peninsula Region (APR) are responding and will likely respond to further environmental changes and direct anthropogenic impacts. Knowledge gained from studies on relatively simple terrestrial ecosystems could be very useful in predicting what may happen in much more complex ecosystems in regions with less extreme temperature changes. The rapid warming of the APR has led to the retreat of glaciers, the loss of snow and permafrost and the increase of ice-free areas, with a consequent enhancement of soil-forming processes, biotic communities, and food web complexity. However, most human activity is concentrated in APR coastal ice-free areas and poses many threats to terrestrial ecosystems such as environmental pollution or disturbances to soilcommunities and wildlife. People who work or visit APR may inadvertently introduce alien organisms and/or spread native species to spatially isolated ice-free areas. The number of introduced non-indigenous species and xenobiotic compounds in the APR is likely to be greater than currently documented, and several biosecurity and monitoring activities are therefore suggested to Antarctic national scientific programs and tourism operators to minimize the risk of irreversible loss of integrity by the unique terrestrial ecosystems of APR.

Antarctic Peninsula region is experiencing one of the fastest rates of climate change on Earth. Its waters are known as important feeding grounds for the Antarctic minke whales (Balaenoptera bonaerensis). The purpose of the present study was to reveal the summer and early autumn presence of the Antarctic minke whales in the area adjacent to the Kiev Peninsula of West Antarctica and to estimate the encounter rates of the species in the area. The boat-based photo-identification cetacean studies were initiated as part of the long-term monitoring program based at the Akademik Vernadsky station near the Kiev Peninsula of West Antarctica. From 22 January to 7 April 2019, 35 boat and yacht cruises of the 821 nautical miles of total length were conducted. There were encountered 13 Antarctic minke whales in 7 sightings. The encounter rate was 0.015 whales per nautical mile. Minke whales were encountered only in 5% of the total sightings. Three more whales were opportunistically seen from the top of Galindez Island. There were single whales sighted and small groups of up to 3 specimens (Med = 2). At least 2 individuals were identified as juveniles. Primary behavior for whales in 7 sightings was foraging, and 2 groups were observed while travelling. A total 9 individuals of the Antarctic minke whales were photo-identified during the survey, and no matches were found between the different encounters. Our pilot study indicates summer and early autumn presence of the Antarctic minke whales in the area adjacent to the Kiev Peninsula. But encounter rates seem to be low in comparison with results of some previous surveys. Our results show the possibility to monitor minke whales in the area, and further long-term complex monitoring is essential for understanding the ecology and population dynamics of the Antarctic minke whales in rapidly changing marine environment of the Antarctic Peninsula.

The Antarctic Peninsula region is of growing interest due to the regional climate change features and related atmospheric circulation patterns. The regional mesoscale atmospheric model Polar Weather Research and Forecasting (WRF) v4.1.1 was used in this research to study a heavy precipitation event over the Ukrainian Antarctic Akademik Vernadsky station region (Antarctic Peninsula). The passage of the cyclone over the Antarctic Peninsula as a typical synoptic process as well as a case of the daily precipitation maximum amount of 2018 were chosen for investigation in this research. The estimation of the modelling quality and downscaling was done by comparing the obtained results with in-situ observation at the Akademik Vernadsky station and cross-domain tracking of average meteorological values and their deviation. The concept of the nested domains allowed to increase the horizontal resolution of the simulated atmosphere up to 1 km and to reproduce the wind regime of this region with high quality. Comparison with measured data showed a significant improvement in wind simulation with increasing of resolution, but worse representation of surface temperature and humidity. The Polar WRF made a general cooling of near surface temperature of 2 °C during the period of simulation and increased precipitation amount by 4.6–8.4 mm (12–21%) on average over the territory relative to the initial data from Global Data Assimilation System. This can be explained by the contribution of noise and imperfection of the model (including static input data of the terrain description). Based on the modelled results, the interaction of wind flow with the mountainous terrain of the Antarctic Peninsula creates a range of complex dynamic effects in the atmosphere. These effects cause local precipitation maxima both over the Peninsula and over the adjacent ocean. These are, respectively, bay-valley areas of increased precipitation and increased precipitation on the crests of shock waves from orographic obstacles. Under certain background wind conditions, the influence of the latter effect can reach the Akademik Vernadsky station and cause the formation of heavy precipitation here.

Background Advances in biologging technology allow researchers access to previously unobservable behavioral states and movement patterns of marine animals. To relate behaviors with environmental variables, features must be evaluated at scales relevant to the animal or behavior. Remotely sensed environmental data, collected via satellites, often suffers from the effects of cloud cover and lacks the spatial or temporal resolution to adequately link with individual animal behaviors or behavioral bouts. This study establishes a new method for remotely and continuously quantifying surface ice concentration (SIC) at a scale relevant to individual whales using on-animal tag video data. Results Motion-sensing and video-recording suction cup tags were deployed on 7 Antarctic minke whales (Balaenoptera bonaerensis) around the Antarctic Peninsula in February and March of 2018. To compare the scale of camera-tag observations with satellite imagery, the area of view was simulated using camera-tag parameters. For expected conditions, we found the visible area maximum to be ~ 100m2 which indicates that observations occur at an equivalent or finer scale than a single pixel of high-resolution visible spectrum satellite imagery. SIC was classified into one of six bins (0%, 1–20%, 21–40%, 41–60%, 61–80%, 81–100%) by two independent observers for the initial and final surfacing between dives. In the event of a disagreement, a third independent observer was introduced, and the median of the three observer’s values was used. Initial results (n = 6) show that Antarctic minke whales in the coastal bays of the Antarctic Peninsula spend 52% of their time in open water, and only 15% of their time in water with SIC greater than 20%. Over time, we find significant variation in observed SIC, indicating that Antarctic minke occupy an extremely dynamic environment. Sentinel-2 satellite-based approaches of sea ice assessment were not possible because of persistent cloud cover during the study period. Conclusion Tag-video offers a means to evaluate ice concentration at spatial and temporal scales relevant to the individual. Combined with information on underwater behavior, our ability to quantify SIC continuously at the scale of the animal will improve upon current remote sensing methods to understand the link between animal behavior and these dynamic environmental variables.

The purpose of the article is to summarize the results of field studies carried out in the framework of the State Antarctic Research Program for 2011–2020 and undertaken to identify areas of special interest for small-scale marine spatial protection in the Argentine Islands water area. The scientific results reported in the publications and relevant meetings’ documents officially presented by the Ukrainian delegation in 2012–2019 were summarized and interpreted in view of the Antarctic Treaty System (ATS) provisions on marine spatial protection. GIS analysis was applied to field data. It is shown that the development of protection regime for small-scale underwater landscape is based on systematic biogeographical data collected during seasonal and year-round Ukrainian Antarctic expeditions in 2011–2019. In order to harmonize the application of all available tools of spatial ecosystem-based protection across the ATS, it is proposed to designate the long-term environmental monitoring sites around Ukrainian Antarctic Akademik Vernadsky station as a new Antarctic Specially Protected Area (ASPA) consisting of terrestrial and marine components. It is shown that the marine component of the proposed ASPA could serve as one of the scientific reference areas of the broad-scale Marine Protected Area in Domain 1 within the Antarctic Peninsula region for assessing the impact of climate change on benthic communities, as well as breeding success and distribution of penguin colonies.

В результате исследований магнитных и плотностных свойств более чем 500 образцов мезозойских и па­лео­ценовых интрузивных и вулканогенных пород, отобранных в западной части Антарктического полу­ост­рова (район станции Академик Вернадский), получена новая информация о петрофизических харак­те­ристиках разных породообразующих комплексов. Породы характеризуются широким диапазоном значений как магнитных (магнитная восприимчивость – от 0.0016 до 0.26 ед. СИ, естественная остаточная намагниченность – от 0.0045 до 8.37 А/м), так и плотностных (от 2.57 до 3.3 гм3) параметров. Различия в плотностных и магнитных характеристиках пород разных петрографических групп укладываются в общие закономерности их изменений в породах кислого, среднего и основного составов. Соответственно средние значения плотности составляют 2.65, 2.77 и 2.9 г/см3, магнитной восприимчивости – 0.02, 0.036 и 0.068 ед. СИ, естественной остаточной намагниченности – 0.16, 0.3 и 2.08 А/м. Новый фактический материал является базовым для параметризации гравитационных и магнитных моделей, рассчитываемых по аномальным потенциальным полям.

Rock samples from the Early to Late Cretaceous marine and non-marine strata of Livingston Island, South Shetland Islands, Antarctica, were studied. Organic matter (OM) originates predominantly from terrestriallyderived organic particles. Mostly highly fragmented plant remains, and locally vitrinite/semifusinite as part of plant compressions/imprints or thin vitrain/fusain lenses, are present. Bulk geochemical results indicate Type III kerogen with poor hydrocarbon generation potential. Tmax and molecular proxies (i.e hopane and sterane isomerization ratios, MPI-1, etc.), together with the local presence of oily droplets, argue for an early oil window maturity, probably caused by regional volcanic activity. In contrast to the results of petrography and Rock-Eval pyrolysis, molecular composition of extractable OM is dominated by short-chain n-alkanes, saturated fatty acids (FAs) and n-alkanols implying a major contribution of autochthonous OM. This contradiction is most likely caused by decomposition of OM from plant tissues, the predominance of wood remains (i.e., xylites) and the low input of FAs from cuticular waxes. The presence of terrestrial OM is indicated by plant wax-derived lipids and resinous compounds. Prominent even carbon number predominance in the short-chain n-alkane range argue for deposition/decomposition under reducing environmental settings, most probably caused by high sedimentation rates and quick burial. Terpenoid biomarker compositions indicate contribution from conifers, whereas input from phytoplankton/algal organic matter is evidenced by C27 steroids, the presence of 4-methyl steranes and dinosterol. Low concentrations of hopanoids and C15 and C17 branched n-FAs argue for limited bacterial input. Only in one sample from Hannah Point, a difference between the δ13C values of the short-chain versus long-chain n-alkanes was obtained, arguing for mixed OM sources (i.e., autochthonous input, land plants). The presence of charred organic matter in most of the studied samples, together with the considerable concentrations of combustion-derived PAHs, argue for the common occurrence of wildfires within the terrestrial environments. Despite the active volcanic arc settings, the compound specific carbon and hydrogen isotopes do not record significant changes in climatic/hydrological conditions, although some short-term events cannot be ruled out. Instead, the δ13C and δ2H isotopic fractionation is most likely caused by the specific biosynthetic mechanisms of the plants and the global carbon and hydrogen cycles.

Benthic samples were collected during two expeditions near the Antarctic Peninsula and in the South-Eastern Weddell Sea. During these studies, a new species of Ampharetidae Malmgren, 1867, Anobothrus konstantini Säring & Bick sp. nov., was found. Here we present a detailed description of this species. We used the traditional light microscope and scanning electron microscope (SEM) to identify and describe the diagnostic characters: a circular glandular band on segment 6; an elongate ridge between the notopodia on segment 12 and modified notochaetae on this segment; 16 thoracic, two intermediate and ten abdominal segments. For the first time, micro-computed tomography (micro-CT) was used for a species description of Anobothrus. Micro-CT provided information on the shape of the prostomium (Ampharete-type) and the arrangement of branchiae (four pairs in two rows, without a gap). In addition, we provide quantitative information on the environmental niche based on sediment parameters (chlorophyll a content, organic matter content, chloroplast equivalent, grain size) for the new Anobothrus species, relevant for, e.g., species distribution modelling. Finally, an identification key for all Anobothrus species is provided.

9 pags, 4 figs. -- The datasets for atmospheric DMS, its oxidative products, and aerosol size distribution are available at the Korea Polar Data Center (https://dx. doi.org/doi:10.22663/KOPRI-KPDC-00001657.4). The sea ice data can be downloaded from the NSIDC website (https://nsidc.org/data/G02135/versions/3). The MODIS-Aqua products can be downloaded from the NASA Ocean Color website (https://oceandata.sci.gsfc.nasa. gov/MODIS-Aqua/). MIMOC is available at https:// www.pmel.noaa. gov/mimoc/. The PHYSAT products can be accessed from the PHYSAT website (https://log.cnrs.fr/Physat-332). The code for the CAM-Chem model is available at https://www2.acom.ucar.edu/gcm/cam-chem. -- Supplementary data to this article can be found online at https://doi.org/10.1016/j.scitotenv.2021.150002., Dimethyl sulfide (DMS) produced by marine algae represents the largest natural emission of sulfur to the atmosphere. The oxidation of DMS is a key process affecting new particle formation that contributes to the radiative forcing of the Earth. In this study, atmospheric DMS and its major oxidation products (methanesulfonic acid, MSA; non-sea-salt sulfate, nss-SO42-) and particle size distributions were measured at King Sejong station located in the Antarctic Peninsula during the austral spring-summer period in 2018-2020. The observatory was surrounded by open ocean and first-year and multi-year sea ice. Importantly, oceanic emissions and atmospheric oxidation of DMS showed distinct differences depending on source regions. A high mixing ratio of atmospheric DMS was observed when air masses were influenced by the open ocean and first-year sea ice due to the abundance of DMS producers such as pelagic phaeocystis and ice algae. However, the concentrations of MSA and nss-SO42- were distinctively increased for air masses originating from first-year sea ice as compared to those originating from the open ocean and multi-year sea ice, suggesting additional influences from the source regions of atmospheric oxidants. Heterogeneous chemical processes that actively occur over first-year sea ice tend to accelerate the release of bromine monoxide (BrO), which is the most efficient DMS oxidant in Antarctica. Model-estimates for surface BrO confirmed that high BrO mixing ratios were closely associated with first-year sea ice, thus enhancing DMS oxidation. Consequently, the concentration of newly formed particles originated from first-year sea ice, which was a strong source area for both DMS and BrO was greater than from open ocean (high DMS but low BrO). These results indicate that first-year sea ice plays an important yet overlooked role in DMS-induced new particle formation in polar environments, where warming-induced sea ice changes are pronounced., We thank the overwintering staff for assisting us in maintaining the aerosol equipment at the King Sejong station. This study was supported by the KOPRI project (PE21030 and PE21120) . KL was supported by the National Research Foundation of Korea (NRF-2021R1A2C3008748) .

The Snow Hill Island Formation (SHIF; late Campanian – early Maastrichtian) crops out in the northeast of the Antarctic Peninsula and constitutes the basal part of the late Campanian-early Maastrichtian sedimentary succession of the James Ross Basin (NG Sequence). Its major exposures occur at the James Ross and Vega islands. Several fossil-bearing localities have been identified in the SHIF providing a valuable fauna of invertebrates and vertebrates, and flora. Our study focuses on the vertebrate fauna recovered at Gamma and Cape Lamb members of the SHIF. The marine vertebrate assemblages include chondrichthyans, actinopterygians, and marine reptiles (elasmosaurid plesiosaurs and mosasaurs). A diverse terrestrial vertebrate assemblage has been reported being characterized by dinosaurs (sauropod, elasmarian ornithopods, nodosaurid ankylosaur, and a paravian theropod), pterosaurs and birds. Most SHIF dinosaurs share close affinities with penecontemporaneous taxa from southern South America, indicating that at least some continental vertebrates could disperse between southern South America and Antarctica during the Late Cretaceous. The Snow Hill Island Formation provides the most diverse Late Cretaceous marine and continental faunas from Antarctica. The present study summarizes previous and new vertebrate findings with the best actualized stratigraphical framework, providing a more complete fauna association and analyzing further perspectives.

The disintegration of the eastern Antarctic Peninsula’s Larsen A and B ice shelves has been attributed to atmosphere and ocean warming, and increased mass-losses from the glaciers once restrained by these ice shelves have increased Antarctica’s total contribution to sea-level rise. Abrupt recessions in ice-shelf frontal position presaged the break-up of Larsen A and B, yet, in the ~20 years since these events, documented knowledge of frontal change along the entire ~1,400 km-long eastern Antarctic Peninsula is limited. Here, we show that 85% of the seaward ice-shelf perimeter fringing this coastline underwent uninterrupted advance between the early 2000s and 2019, in contrast to the two previous decades. We attribute this advance to enhanced ocean-wave dampening, ice-shelf buttressing and the absence of sea-surface slope-induced gravitational ice-shelf flow. These phenomena were, in turn, enabled by increased near-shore sea ice driven by a Weddell Sea-wide intensification of cyclonic surface winds around 2002. Collectively, our observations demonstrate that sea-ice change can either safeguard from, or set in motion, the final rifting and calving of even large Antarctic ice shelves.

The classification of Synthetic Aperture Radar (SAR) images by knowledge-based algorithms with elevation and backscatter thresholds were used in several studies to detect the Wet Snow Radar Zone (WSZ) in the Antarctic Peninsula. To identify it more accurately based on its seasonal variations, this study proposed the additional use of a threshold in synthetic images, created by rationing summer and winter sigma linear images. In our algorithm we used the following thresholds to detect the WSZ in Envisat ASAR imageries, using the Radarsat Antarctic Map Digital Elevation Model as ancillary data: i) -25 dB < s0 < -14 dB; ii) slinear summer / slinear winter < 0.4; iii) elevation H < 1,200 m for northern tip and H < 800 m for southern tip of the Antarctic Peninsula. The classified images were post-processed by a focal majority 5 x 5 filter and superimposed by an image of rock outcrops derived from the Antarctic Digital Database. The ratio image threshold allowed discriminating the WSZ from the Dry Snow Radar Zone and radar shadows, as well as transitional areas between this glacier zone and the Frozen Percolation Radar Zone, which would be classified incorrectly if we used only elevation and backscatter thresholds.

We applied multi–temporal 1D magnetotelluric (MT) surveys to identify space–time anomalies of apparent resistivity (ρa) in the upper lithosphere in the Antarctic Peninsula (the border between the Antarctic and the Shetland plates). We used time series over several weeks of the natural Earth’s electric and magnetic fields registered at one MT station of the Universidad Nacional de Colombia (RSUNAL) located at Seymour–Marambio Island, Antarctica. We associated resistivity anomalies with contrasting earthquake activity. Anomalies of ρa were detected almost simultaneously with the beginning of a seismic crisis in the Bransfield Strait, south of King George Island (approximately 85.000 events were reported close to the Orca submarine volcano, with focal depths < 20 km and MWW < 6.9). We explained the origin of these anomalies in response to fluid migration near the place of the fractures linked with the seismic swarm, which could promote disturbances of the pore pressure field that reached some hundreds of km away.

Ice scouring is one of the strongest agents of disturbance in nearshore environments at high latitudes. In depths, less than 20 m, grounding icebergs reshape the soft-sediment seabed by gouging furrows called ice pits. Large amounts of drift algae (up to 5.6 kg/m2) that would otherwise be transported to deeper water accumulate inside these features, representing an underestimated subsidy. Our work documents the distribution and dimensions of ice pits in Fildes Bay, Antarctica, and evaluates their relationship to the biomass and species composition of algae found within them. It also assesses the rates of deposition and advective loss of algae in the pits. The 17 ice pits found in the study area covered only 4.2% of the seabed but contained 98% of drift algal biomass, i.e., 60 times the density (kg/m2) of the surrounding seabed. Larger ice pits had larger and denser algal accumulations than small pits and had different species compositions. The accumulations were stable over time: experimentally cleared pits regained initial biomass levels after one year, and advective loss was less than 15% annually. Further research is needed to understand the impacts of ice scouring and subsequent algal retention on ecosystem functioning in this rapidly changing polar environment.

Volcanism can play a key role in modulating climate; however, a lack of historical records has limited our comprehension of Antarctic volcanism and its role on the cryosphere. Remote sensing can provide insight into active volcanism in Antarctica during the satellite era, although the evidence is often inconclusive. Here, we use independent evidence from ice cores to validate one such potential volcanic eruption from the sub‐Antarctic Balleny Islands in 2001 CE. Multiple ice cores from downwind of the eruption site, record elevated input of sulfate, microparticles, and the presence of tephra, coincident with the eruption. In‐phase deposition of volcanic products confirmed a rapid tropospheric transport of volcanic emissions from a small‐to‐moderate, local eruption during 2001. Air mass trajectories demonstrated some air parcels were transported over the West Antarctic ice sheet from the Balleny Islands to ice core sites at the time of the potential eruption, establishing a route for transport and deposition of volcanic products over the ice sheet. The data presented here validate previous remote sensing observations and confirms a volcanic event in the Balleny Islands during 2001 CE. This newly identified eruption provides a case study of recent Antarctic volcanism.

The Antarctic krill (Euphausia superba) fishery is the largest fishery in the Southern Ocean, and it has been operating for over 45 years. In the past decade, the spatial distribution of the krill fishery has focused on the Bransfield Strait off the Antarctic Peninsula (subarea 48.1). However, the high fishing effort and climate change have placed great pressure on krill resources in this region, and conservation concerns have been raised. Because aging krill is difficult and uncertain, we developed an integrated size-structured model to estimate the fishing mortality, recruitment and spawning biomass of krill. The results indicated that in 1992–2011, the average spawning biomass of krill ranged from 1.14 × 106 to 1.45 × 106 tons, the estimated biomass of the maximum sustainable yield (BMSY) ranged from 3.96 × 105 to 4.90 × 105 tons and the estimated average recruitment ranged from 2.3 × 1012 to 5.03 × 1012 individuals in the research area. We explored the effect of different data weighting schemes for the length data on the assessment output. However, our estimates also have uncertainties. In addition, an attempt was made to analyze the correlation between model-estimated recruitment and mature biomass with climate change. Future stock assessments of krill resources should be comprehensive and multimethod, and the management of krill resources should be based on science that considers the demand of fishing communities and ecosystem protection.

High-accumulation sites are crucial for understanding the patterns and mechanisms of climate and environmental change in Antarctica since they allow gaining high-resolution proxy records from firn and ice. Here, we present new glacio- and isotope-geochemical data at sub-annual resolution from a firn core retrieved from an ice cap on Plateau Laclavere (LCL), northern Antarctic Peninsula, covering the period 2012–2015. The signals of two volcanic eruptions and two forest fire events in South America could be identified in the non-sea-salt sulphur and black carbon records, respectively. Mean annual snow accumulation on LCL amounts to 2500 kg m−2 a−1 and exhibits low inter-annual variability. Time series of δ18O, δD and d excess show no seasonal cyclicity, which may result from (1) a reduced annual temperature amplitude due to the maritime climate and (2) post-depositional processes. The firn core stratigraphy indicates strong surface melt on LCL during austral summers 2013 and 2015, likely related to large-scale warm-air advection from lower latitudes and temporal variations in sea ice extent in the Bellingshausen-Amundsen Sea. The LCL ice cap is a highly valuable natural archive since it captures regional meteorological and environmental signals as well as their connection to the South American continent.

Southern Ocean ecosystems offer numerous benefits to human society and the global environment, and maintaining them requires well-informed and effective ecosystem-based management. Up to date and accurate information is needed on the status of species, communities, habitats and ecosystems and the impacts of fisheries, tourism and climate change. This information can be used to generate indicators and undertake assessments to advise decision-makers. Currently, most marine assessments are derivative: reliant on the review of published peer-reviewed literature. More timely and accurate information for decision making requires an integrated Marine Biological Observing and Informatics System that combines and distributes data. For such a system to work, data needs to be shared according to the FAIR principles (Findable, Accessible, Interoperable, and Reusable), use transparent and reproducible science, adhere to the principle of action ecology and complement global initiatives. Here we aim to provide an overview of the components of such a system currently in place for the Southern Ocean, the existing gaps and a framework for a way forward.

Fine-scale knowledge of spatiotemporal dynamics in cetacean distribution and abundance throughout the Western Antarctic Peninsula (WAP) is sparse yet essential for effective ecosystem-based management (EBM). Cruise vessels were used as platforms of opportunity to collect data on the distribution and abundance of humpback whales (Megaptera novaeangliae) during the austral summer of 2019/2020 in a region that is also important for the Antarctic krill (Euphausia superba) fishery, to assess potential spatiotemporal interactions for future use in EBM. Data were analyzed using traditional design-based line transect methodology and spatial density surface hurdle models fitted using a set of physical environmental covariates to estimate the abundance and distribution of whales in the area, and to describe their temporal dynamics. Our results indicate a rapid increase in humpback whale abundance in the Bransfield and Gerlache Straits through December, reaching a stable abundance by mid-January. The distribution of humpback whales appeared to change from a patchier distribution in the northern Gerlache Strait to a significantly concentrated presence in the central Gerlache and southern Bransfield Straits, followed by a subsequent dispersion throughout the area. Abundance estimates agreed well with previous literature, increasing from approximately 7000 individuals in 2000 to a peak of 19,107 in 2020. Based on these estimates, we project a total krill consumption of between 1.4 and 3.7 million tons based on traditional and contemporary literature on per capita krill consumption of whales, respectively. When taken in the context of krill fishery catch data in the study area, we conclude that there is minimal spatiotemporal overlap between humpback whales and fishery activity during our study period of November-January. However, there is potential for significant interaction between the two later in the feeding season, but cetacean survey efforts need to be extended into late season in order to fully characterize this potential overlap. Publisher PDF

Polar regions, symbols of wilderness, have been identified as potential sinks of mercury coming from natural and anthropogenic sources at lower latitudes. Changes in ice coverage currently occurring in some areas such as the Antarctic Peninsula could enhance these phenomena and their impacts on local biota. As long-lived species at the top of food chains, seabirds are particularly sensitive to this highly toxic metal with the capacity to be biomagnified. Specifically, their feathers can be useful for Hg monitoring since they mainly accumulate its most toxic and persistent form, methyl-Hg. To that end, feathers of gentoo (Pygoscelis papua), chinstrap (P. antarcticus), and Adélie penguins (P. adeliae) (n = 108) were collected by passive sampling in seven different locations throughout the Antarctic Peninsula area and analyzed by ICP-MS after microwave-digestion. More than 93% of the samples showed detectable Hg levels (range: 6.3¿12,529.8 ng g¿1 dry weight), and the highest ones were found in the feathers of chinstrap penguins from King George Island. Hg bioconcentration and biomagnification seem to be occurring in the Antarctic food web, giving rise to high but non-toxic Hg levels in penguins, similar to those previously found in Arctic seabirds., This research has been funded by the Spanish Ministry of Science and Innovation CGL2004-01348, POL2006-05175, and CGL2007-60369.

Although the glaciers in the Antarctic periphery currently modestly contribute to sea level rise, their contribution is projected to increase substantially until the end of the 21st century. The South Shetland Islands (SSI), located to the north of the Antarctic Peninsula, are lacking a geodetic mass balance calculation for the entire archipelago. We estimated its geodetic mass balance over a 3–4-year period within 2013–2017. Our estimation is based on remotely sensed multispectral and interferometric SAR data covering 96% of the glacierized areas of the islands considered in our study and 73% of the total glacierized area of the SSI archipelago (Elephant, Clarence, and Smith Islands were excluded due to data limitations). Our results show a close to balance, slightly negative average specific mass balance for the whole area of −0.106 ± 0.007 m w.e. a−1, representing a mass change of −238 ± 12 Mt a−1. These results are consistent with a wider scale geodetic mass balance estimation and with glaciological mass balance measurements at SSI locations for the same study period. They are also consistent with the cooling trend observed in the region between 1998 and the mid-2010s.

The aim of the study is to analyze seasonal and interannual changes in sea level on the western coast of the Antarctic Peninsula. Objects of study are seasonal and interannual variability of sea level, air temperature, pressure at sea level, precipitation in the period 1960–2018 at the Faraday/Akademik Vernadsky station, which was considered as representative site for the Antarctic Peninsula. Statistical methods of study were used, including estimates of linear trends of time series using the nonparametric Sen’s estimator of slope. The Mann-Kendall test was used to assess the significance of the slope of the trend. Time variability analysis was performed using wavelet analysis. Using the MATrix LABoratory (MATLAB) software package, squared of wavelet coefficients were calculated depending on the scale and shift or scalograms that characterize the local energy spectrum, and scalegrams calculated by averaging the scalograms by time shifts. The Morlet wavelet transformations were used. The results of the calculations showed that the trend of sea level at the Faraday/ Akademik Vernadsky station in the period 1960—2018, according to observations and correction on glacial isostatic adjustment of the crust is in the range from 3.05 to 3.45 mm/year, which is significantly higher than the global trend of 2.1 mm/year. Sea level scalograms allow estimating time-averaged periods and amplitudes of the coefficients for each season. In the austral winter and spring characteristic periods were 4—6 years, whereas the summer and autumn periods are characterized by 6–8 years, as well as by the highest amplitudes of the coefficients. All seasons are characterized by the appearance of a weakly pronounced period of about 4 years. The presence of peaks in the scalegrams at 6–8 and 3–4 years confirms the relationship of atmospheric and oceanic processes in West Antarctica to the natural variations of the ocean-atmosphere system, such as the Southern Annular Mode and El Niño-Southern Oscillation, which varies with typical 3–4 and 6–8 year periods. It was concluded that, unlike air temperature, the sea level trend is relatively weakly depends on the season, with the exception of autumn, when sea level trend is three times smaller than the average value. At the same time, the air temperature trend is the largest in the austral winter and the lowest in the summer.

Here, a geomorphological map of Horseshoe Island, which is one of the most ice-free islands in Marguerite Bay of the Antarctic Peninsula, is provided. The landforms on the island were mapped by using Google Earth images. Field reconnaissance of the landforms was carried out in March 2018. The island is subdivided into three major geomorphologically different sectors. The northern sector is mostly covered by a remnant of a non-erosive ice cap and has limited glacial landforms and deposits. The central sector is rich in terms of glacial and periglacial landforms and deposits. Glaciers are still sculpting the southern sector and it has extensive features of glacial erosion and deposition. The most common landforms on the island are talus cones, moraines, patterned ground, and raised beaches. The geomorphological map of the island will be a useful base for further geomorphic and/or glaciologic research in this climate-sensitive region.

Oblique zooplankton tows were carried out in the Gerlache Strait from 17 to 22 January 2015 as part of the first Colombian oceanographic expedition to the Antarctic continent, to document the composition and structure of the epipelagic copepod assemblage, taking into account variations at a small spatial scale. The epipelagic environment of the Gerlache Strait was divided into two oceanographic regions during sampling: a stratified northern region and a homogeneous southern region. The epipelagic copepod assemblage comprised 17 species belonging to four orders and 13 families. Adult copepods as well as developmental stages ranging from copepodite II (C2) to copepodite V (C5) from eight different species were recorded in the study area. The dominant species by number was Oithona similis, followed by O. frigida, Ctenocalanus citer, Drescheriella glacialis, and Calanoides acutus. There were significant differences between the epipelagic copepod assemblages present in the northern and southern regions of the Gerlache Strait (Anosim, p = 0.01), with a dissimilarity percentage of 52%. Moreover, Aetideus armatus is reported for the first time in the western sector of the Antarctic Peninsula. The oceanographic conditions and the geomorphological characteristics of the study area modulated the surface circulation pattern, as well as the structure and composition of the primary producers during the study period, determining the spatial variation of the abundance and composition of the epipelagic copepod assemblage in the Gerlache Strait during the 2015 Austral summer.

Meteorological measurements were performed on the non-glaciated coastal area of the Ferraz Station, the Brazilian Antarctic Station, at King George Island. Near the Station there are different topographic attributes, such as sea, glacier and hill, in addition to the intermittent presence of snow. The local atmospheric characteristics depend on wind direction, which is related in small scale to the local features and in large scale to the synoptic systems acting over the region. Westerly winds were the most frequent, with relatively lower speed and higher temperature and humidity values compared to the also frequent easterly winds, related to the coolest and driest winds. The average monthly barometric pressure and wind speed were higher during winter, associated with the presence of high-pressure systems. The radiation balance, measured for the first time in Ferraz Station, evidenced the presence of clouds throughout the year, especially during summer months, associated with the frequent synoptic systems of the region. During nighttimes (daytimes) and in winter (summer) months, the surface lost (gained) energy mainly by net longwave (shortwave) radiation. Reflected shortwave radiation was higher during spring months because of the its relatively higher albedo values combined with the already increased magnitude of the incident shortwave radiation.

Isoprene (C5H8) plays an important role in the formation of surface ozone (O3) and the secondary organic aerosol (SOA) which contributed to the climate change. This study aims to determine hourly distribution of tropospheric isoprene over the Western Coast of Antarctic Peninsula (WCAP) during the Malaysian Antarctic Scientific Expedition Cruise 2016 (MASEC′16). In-situ measurements of isoprene were taken using a custom-built gas chromatography with photoionization detector, known as iDirac. Biological parameters such as chlorophyll a (chl-a) and particulate organic carbon (POC) were compared to the in-situ isoprene measurements. Significant positive correlation was observed between isoprene and POC concentrations (r2 = 0.67, p

The James Ross Basin, in the northern Antarctic Peninsula, exposes which is probably the world thickest and most complete Late Cretaceous sedimentary succession of southern high latitudes. Despite its very good exposures and varied and abundant fossil fauna, precise chronological determination of its infill is still lacking. We report results from a magnetostratigraphic study on shelfal sedimentary rocks of the Marambio Group, southeastern James Ross Basin, Antarctica. The succession studied covers a ~1,200 m-thick stratigraphic interval within the Hamilton Point, Sanctuary Cliffs and Karlsen Cliffs Members of the Snow Hill Island Formation, the Haslum Crag Formation, and the lower López de Bertodano Formation. The basic chronological reference framework is given by ammonite assemblages, which indicate a Late Campanian – Early Maastrichtian age for the studied units. Magnetostratigraphic samples were obtained from five partial sections located on James Ross and Snow Hill islands, the results from which agree partially with this previous biostratigraphical framework. Seven geomagnetic polarity reversals are identified in this work, allowing to identify the Chron C32/C33 boundary in Ammonite Assemblage 8-1, confirming the Late Campanian age of the Hamilton Point Member. However, the identification of the Chron C32/C31 boundary in Ammonite Assemblage 8-2 assigns the base of the Sanctuary Cliffs Member to the early Maastrichtian, which differs from the Late Campanian age previously assigned by ammonite biostratigraphy. This magnetostratigraphy spans ~14 Ma of sedimentary succession and together with previous partial magnetostratigraphies on Early-Mid Campanian and Middle Maastrichtian to Danian columns permits a complete and continuous record of the Late Cretaceous distal deposits of the James Ross Basin. This provides the required chronological resolution to solve the intra-basin and global correlation problems of the Late Cretaceous in the Southern Hemisphere in general and in the Weddellian province in particular, given by endemism and diachronic extinctions on invertebrate fossils, including ammonites. The new chronostratigraphic scheme allowed us to calculate sediment accumulation rates for almost the entire Late Cretaceous infill of the distal James Ross Basin (the Marambio Group), showing a monotonous accumulation for more than 8 Myr during the upper Campanian and a dramatic increase during the early Maastrichtian, controlled by tectonic and/or eustatic causes. Fil: Milanese, Florencia Nidia. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Ciudad Universitaria. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires. Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires; Argentina Fil: Olivero, Eduardo Bernardo. Universidad Nacional de Tierra del Fuego; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Austral de Investigaciones Científicas; Argentina Fil: Raffi, María Eugenia. Universidad Nacional de Tierra del Fuego; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Austral de Investigaciones Científicas; Argentina Fil: Franceschinis, Pablo Reinaldo. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Ciudad Universitaria. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires. Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires; Argentina Fil: Gallo, Leandro César. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Ciudad Universitaria. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires. Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires; Argentina Fil: Skinner, Steven M.. California State University; Estados Unidos Fil: Mitchell, Ross N.. California Institute of Technology; Estados Unidos Fil: Kirschvink, Joseph L.. California Institute of Technology; Estados Unidos. Tokyo Institute of Technology; Japón Fil: Rapalini, Augusto Ernesto. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Ciudad Universitaria. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires. Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Instituto de Geociencias Básicas, Aplicadas y Ambientales de Buenos Aires; Argentina

18 pages, 10 figures, 5 tables, supplementary material https://doi.org/10.1016/j.gca.2021.04.021, Burial fluxes of organic carbon and biogenic silica were determined in 17 continental shelf sediment cores collected from the northern Weddell Sea, the Bransfield Strait, and the southern Drake Passage. Coring sites included open-shelf stations as well as slope and glacial trough environments, with water depths varying from 220 to 760 m. Apparent 210Pb accumulation rates from these cores ranged from 0.04 g m−2y−1 to 0.21 g m−2y−1 (1 to 3 mm y−1), with organic carbon burial rates ranging from 3 to 15 g OC m−2y−1 and biogenic silica accumulation rates ranging from 15 to 126 g SiO2 m−2y−1. OC contents below the surface mixed layer ranged from 0.26 to 1.51 wt. % (avg. 0.64 %). Biogenic silica contents at depth ranged from 2.3 to 11.2 wt. % (avg. 7.5%), with an average bSi/OC ratio (wt. %/wt. %) at depth of 12. Annual OC primary production rates and biogenic silica production rates in the euphotic zone were estimated from satellite chlorophyll-a data in the literature and from a seasonal model for biogenic particle export from surface waters. Based on these biogeochemical data, preservation efficiencies (i.e., mass burial rate/water column production rate) were calculated for organic carbon and biogenic silica. These preservation efficiency values ranged from 2 to 18% (avg. 9%) for OC and 8 to 106% (avg. 54%) for bSi. These relatively high preservation efficiencies resulted from extensive lateral sediment focusing (210Pb Psi (Ψ) values [burial flux/water column production rate] ranging from 2 to 33; avg. of 16), cold bottom water temperatures (2 to −2°C), and relatively high biogenic Si and OC production rates in the euphotic zone. The enhanced preservation efficiency for bSi relative to OC (i.e., 54% vs. 9%) in these Antarctic settings is consistent with the change in the phytoplankton bSi/OC (wt. %/wt. %) value of 2 for this area to the burial bSi/OC value of 12. Excess 210Pb activities in surface sediments varied from 4 to 47 dpm g−1. The surface mixed layer in the seabed varied in thickness from 0 to 4 cm. The penetration of excess 210Pb into these Antarctic Peninsula sediments varied from 6 to 28 cm (avg. 18 cm). The inventory of excess 210Pb in the seabed varied from 13 to 230 dpm cm−2 (avg. 110 dpm cm−2). Although 210Pb was the only radionuclide measured in this study, “apparent” 210Pb sediment accumulation rate (SAR) values from these 17 cores (assuming that deep bioturbation is negligible) are believed to be accurate SAR values because of good agreement between 210Pb and 14C chronologies from nearby cores reported in the literature, This work was funded by the Spanish Ministry of Economy and Competitiveness as part of the project ECOWED (CTM2012-39350-C02-01), With funding from the Spanish government through the ‘Severo Ochoa Centre of Excellence’ accreditation (CEX2019-000928-S)

New, well-preserved fossil starfish material is recorded from the Eocene La Meseta Formation exposed in Seymour Island, Antarctica. The use of new technology (i.e., microCT) on several fragments enabled the visualization of new characters and the differentiation of a new species, Zoroaster marambioensis sp. nov., which was previously identified as Zoroaster aff. Z. fulgens. Diagnostic characters of Z. marambioensis sp. nov. are (i) central disc plate enlarged, lobate and flattened, (ii) disc ring with enlarged, tumid radials and polygonal, flattened inter-radials, (iii) primary spines on disc only present on radials, (iv) oral armature with 1–3 primary spines and 1–2 secondary spines for each prominent adambulacral. The depositional setting represents the outer zone of an estuary dominated by marine processes affected by long lived hyperpycnal flows. We argue that zoroasterids colonized a distal part of the estuary under normal marine salinity and were killed by the input of freshwater carried by a hyperpycnal flow, and immediately buried by fine grained sandstone. Sedimentological data suggest that Z. marambioensis sp. nov. lived in shallow-water environments, it seems possible that they were adapted to higher temperatures than other Recent species of the genus, which inhabit cold, deep marine environments., Facultad de Ciencias Naturales y Museo

This work is a core contribution to the first Marine Ecosystem Assessment for the Southern Ocean (MEASO) of IMBeR’s program ICED.-- 30 pages, 8 figures, 2 tables, supplementary material https://www.frontiersin.org/articles/10.3389/fmars.2021.622721/full#supplementary-material, Knowledge of life on the Southern Ocean seafloor has substantially grown since the beginning of this century with increasing ship-based surveys and regular monitoring sites, new technologies and greatly enhanced data sharing. However, seafloor habitats and their communities exhibit high spatial variability and heterogeneity that challenges the way in which we assess the state of the Southern Ocean benthos on larger scales. The Antarctic shelf is rich in diversity compared with deeper water areas, important for storing carbon (“blue carbon”) and provides habitat for commercial fish species. In this paper, we focus on the seafloor habitats of the Antarctic shelf, which are vulnerable to drivers of change including increasing ocean temperatures, iceberg scour, sea ice melt, ocean acidification, fishing pressures, pollution and non-indigenous species. Some of the most vulnerable areas include the West Antarctic Peninsula, which is experiencing rapid regional warming and increased iceberg-scouring, subantarctic islands and tourist destinations where human activities and environmental conditions increase the potential for the establishment of non-indigenous species and active fishing areas around South Georgia, Heard and MacDonald Islands. Vulnerable species include those in areas of regional warming with low thermal tolerance, calcifying species susceptible to increasing ocean acidity as well as slow-growing habitat-forming species that can be damaged by fishing gears e.g., sponges, bryozoan, and coral species. Management regimes can protect seafloor habitats and key species from fishing activities; some areas will need more protection than others, accounting for specific traits that make species vulnerable, slow growing and long-lived species, restricted locations with optimum physiological conditions and available food, and restricted distributions of rare species. Ecosystem-based management practices and long-term, highly protected areas may be the most effective tools in the preservation of vulnerable seafloor habitats. Here, we focus on outlining seafloor responses to drivers of change observed to date and projections for the future. We discuss the need for action to preserve seafloor habitats under climate change, fishing pressures and other anthropogenic impacts, MB was supported by the Antarctic Climate & Ecosystems Cooperative Research Centre and PEW Charitable Trusts. RD was funded by an Australian Government Research Training Program (AGRTP). SM and HG were funded through NERC core funding to the British Antarctic Survey Biodiversity, Evolution and Adaptation Team. BF was supported by a postdoctoral contract Juan de la Cierva-Incorporación (IJCI-2017-31478) of Ministerio de Ciencia, Innovación y Universidades. AV was funded by the Belgian Science Policy Office (BELSPO, contract n° FR/36/AN1/AntaBIS) in the Framework of EU-Lifewatch. [...] BF received institutional support of the ‘Severo Ochoa Centre of Excellence’ accreditation (CEX2019-000928-S)

Expansive study areas, such as those used by highly-mobile species, provide numerous logistical challenges for researchers. Community science initiatives have been proposed as a means of overcoming some of these challenges but often suffer from low uptake or limited long-term participation rates. Nevertheless, there are many places where the public has a much higher visitation rate than do field researchers. Here we demonstrate a passive means of collecting community science data by sourcing ecological image data from the digital public, who act as “eco-social sensors,” via a public photo-sharing platform—Flickr. To achieve this, we use freely-available Python packages and simple applications of convolutional neural networks. Using the Weddell seal (Leptonychotes weddellii) on the Antarctic Peninsula as an example, we use these data with field survey data to demonstrate the viability of photo-identification for this species, supplement traditional field studies to better understand patterns of habitat use, describe spatial and sex-specific signals in molt phenology, and examine behavioral differences between the Antarctic Peninsula’s Weddell seal population and better-studied populations in the species’ more southerly fast-ice habitat. While our analyses are unavoidably limited by the relatively small volume of imagery currently available, this pilot study demonstrates the utility an eco-social sensors approach, the value of ad hoc wildlife photography, the role of geographic metadata for the incorporation of such imagery into ecological analyses, the remaining challenges of computer vision for ecological applications, and the viability of pelage patterns for use in individual recognition for this species.

This research aims to investigate sediment cores from a glaciomarine environment based on their petrophysical parameters to elucidate lithological and sedimentary issues, as well as to identify a geophysical signature based on their parameter’s response. To achieve this objective, six marine sediment cores were collected in the Central Bransfield Basin - Antarctica, the lengths ranging from 1.5 to 5.2 meters in water depths ranging from 304 to 1463 meters. The cores were submitted to different analyses and values from density, magnetic susceptibility, electrical resistivity, p-wave velocity, total gamma radiation, silt, clay and sand contents, and mean grain size for each core location were considered. Four large lithologies were identified according to their geological and geophysical characteristics. The first group is subglacial deformation till (GC16); the second is massive diamicton (GC13 and GC12), the third group is composed of diamictons from the basin (AM10) and lower slope (GC09); the fourth lithological group is composed of siliceous mud from an upper slope location (GC06A). The characteristics recorded across Central Bransfield Basin (from South Shetlands Islands to Antarctic Peninsula) highlighted the relationship between the lithological content and associated depositional processes with the geophysical proprieties as density and magnetic susceptibility.

It is estimated that the explosive Hudson volcano eruption in Southern Chile injected approximately 2.7 km3 of basalt and trachyandesite tephra into the troposphere between August 8-15, 1991. The Hudson signal has been detected in Antarctica at the eastern sector and in South Pole snow. In this work, we track the Hudson volcanic plume using a dispersion model, remote sensing, and a re-analysis of a high-resolution ice core analysis from the Detroit Plateau in the Antarctic Peninsula and sedimentary records from shallow lakes from King George Island (KGI). The Hudson eruption imprint in these records is confirmed by using a weekly resolved aerosol concentration database from KGI demonstrating that the regional impact of Hudson eruption predominates over the Mount Pinatubo/Phillippines volcanic signal, dated from June 1991, in terms of particulate matter depositions. The aerosol elemental composition of Ca, Fe, Ti, Si, Al, Zn, and Pb increases from 2 to 3 orders of magnitude in background level during the days following the eruption of the Hudson volcano.

16 pages, 7 figures, 3 tables, supporting information https://pubs.acs.org/doi/10.1021/acsearthspacechem.0c00351.-- All data are available on the public repository PANGAEA: https://doi.pangaea.de/10.1594/PANGAEA.927565 (for aerosol particles) and https://doi.pangaea.de/10.1594/PANGAEA.927566 (for seawater), We present ship-borne and land-based measurements of carbohydrate concentrations and patterns in (i) bulk seawater, (ii) sea surface microlayer (SML), and (iii) atmospheric size-resolved aerosol particles (0.05–10 μm) collected in the Western Antarctic Peninsula. In seawater, we find higher combined carbohydrates (CCHO) in both the particulate (PCCHO, 13–248 μg L–1) and dissolved (DCCHO, 14–294 μg L–1) phases than dissolved free carbohydrates (DFCHO, 1.0–17 μg L–1). Moderate enrichment factors are found in the SML samples (median EFSML = 1.4 for PCCHO, DCCHO, and DFCHO). In PM10 atmospheric particles, combined carbohydrates (CCHOaer,PM10 0.2–11.3 ng m–3) were preferably found in particles of two size modes (0.05–0.42 and 1.2–10 μm) and strongly correlated with Na+aer,PM10 and wind speed, hence suggesting oceanic emission as their primary source. In contrast to SML samples, very high enrichment factors for CCHOaer relative to the bulk water (EFaer) were estimated for supermicron (20–4000) and submicron (40–167 000) particles. Notably, the relative atmospheric aerosol monosaccharide compositions strongly differed from the ones sampled in seawater. The prevalence of bacterial monosaccharides (muramic acid, glucosamine) in aerosol particles allows us to suggest a selective consumption and release of polysaccharides by bacteria in the atmosphere. Our results highlight the need to evaluate the role of different ecosystems as aerosol sources around Antarctica, The authors gratefully acknowledge the funding by the Deutsche Forschungsgemeinschaft (DFG, German Research Foundation, Projektnummer 268020496−TRR 172) within the Transregional Collaborative Research Center “ArctiC Amplification: Climate Relevant Atmospheric and SurfaCe Processes, and Feedback Mechanisms (AC)3” in subprojects B04. Additional support through the Leibniz Association SAW funding of the project “Marine biological production, organic aerosol particles and marine clouds: a Process Chain (MarParCloud)” (SAW-2016-TROPOS-2) is also gratefully acknowledged, With the funding support of the ‘Severo Ochoa Centre of Excellence’ accreditation (CEX2019-000928-S), of the Spanish Research Agency (AEI)

Among nudibranch molluscs, the family Tritoniidae gathers taxa with an uncertain phylogenetic position, such as some species of the genus Tritonia Cuvier, 1798. Currently, 37 valid species belong to this genus and only three of them are found in the Southern Ocean, namely T. challengeriana Bergh, 1884, T. dantarti Ballesteros & Avila, 2006, and T. vorax (Odhner, 1926). In this study, we shed light on the long-term discussed systematics and taxonomy of Antarctic Tritonia species using morpho-anatomical and molecular techniques. Samples from the Weddell Sea and Bouvet Island were dissected and prepared for scanning electron microscopy. The three molecular markers COI, 16S, and H3 were sequenced and analysed through maximum-likelihood and Bayesian methods. The phylogenetic analyses and species delimitation tests clearly distinguished two species, T. challengeriana widely spread in the Southern Ocean and T. dantarti endemic to Bouvet Island. Colouration seems to be an unreliable character to differentiate among species since molecular data revealed both species can either have orange or white colour morphotypes. This variability could be explained by pigment sequestration from the soft coral species they feed on. Morphological analyses reveal differences between Antarctic and Magellanic specimens of T. challengeriana. However, the relationship between T. challengeriana specimens from these two regions remains still unclear due to the lack of molecular data. Therefore, the validity of the T. antarctica Martens & Pfeffer, 1886, exclusively found in Antarctic waters requires further systematic work.

The insoluble particulate matter deposited on ice sheets provide key information to reconstruct past climate. The low concentration of some insoluble particulate matter, such as terrigenous particles and microfossils, challenges the efficiency of the recovery and the representativeness of the results. Here we present a new optimized method to extract, quantify and classify targeted low concentration insoluble particulate matter. Particle recovery rates and particle distribution were investigated using polystyrene particle standards filtered through Polycarbonate membrane filters and subsequently scanned in a scanning electron microscope. Experimental results in continuous and discrete sampling systems reveal consistent trends in the transport and removal of particulate material inside a filtration system. Statistical simulations are used to optimize the sample analyses required to achieve representative results. The analysis of diatoms in ice cores using this new method uncovered their potential to hold valuable climate records from the Antarctic Peninsula region. The data presented here evidence the presence of a measurable amount of marine diatoms with sub-annual variations, highlighting the potential of this record as a seasonal indicator. The new method presented provides an optimized and statistically representative approach for extracting, recovering and analyzing micrometre-sized, low-concentration insoluble particulate matter in ice.

Here, we report on two tarsometatarsi assignable to relatively small-sized Eocene Antarctic penguins, housed in the palaeozoological collections of Naturhistoriska riksmuseet, Stockholm. The Priabonian fossils were collected by museum staff during two joined Argentinean and Swedish expeditions from the Submeseta Formation on Seymour Island, Antarctic Peninsula. One specimen represents a new early sphenisciform, Marambiornopsis sobrali gen. et sp. nov., the sixth small-sized tarsometatarsus-based penguin species known from the Antarctic Eocene. Micro-CT scanning revealed the presence of quite large and essentially empty metatarsal medullary cavities. The second fossil can unequivocally be assigned to Mesetaornis polaris. The specimen represents only the second record of this species and supposedly a relatively young bird. Micro-CT scanning showed that in M. polaris the metatarsal medullary cavities are less developed than in M. sobrali – the cortical and trabecular bone tissues left rather little room for significant hollow spaces. Both specimens also differ in overall density of their trabecular networks. We thank the Argentinean Antarctic Institute (IAA-DNA), the Argentinean Air Force and the Swedish Polar Research Secretariat (SPFS) for logistical support in Antarctica and Jonas Hagström (NRM) for assistance in the field. We thank Vivi Vajda (NRM) for making the CT scanning possible, Ashley Kruger (NRM) for the CT-scanning. Financial support through SYNTHESYS funding made available by the European Community – Research Infrastructure Action under the FP6 Structuring the European Research Area Programme, project SE-TAF-4399 to PJ; from the Consejo Nacional de lnvestigaciones Cientificas y Tecnicas (CONICET grant PIP 0462) and from the Argentinean National Agency for Promotion of Science and Technology (ANPCyT grant PICTO 0093/2010) to MR; from the Swedish Research Council (VR Grant 2009-4447) and from the Carl Tryggers Foundation (CTS grant 20:300) to TM is gratefully acknowledged.