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Recent geophysical campaigns in the Alps produce images with seismic property variations along the slab of sufficiently fine resolution to be interpreted as rock transformations. Since the reacting European lower crust is presumed responsible for the variations of velocities at the top of the Alpine slab, we sampled local analogs of the lower crustal lithologies in the field and modeled the evolution of equilibrium seismic properties during burial, along possible pressure‐temperature paths for the crustal portion of the slab. The results are then compared to the range of the S‐wave velocities obtained from the S‐wave velocity tomography model along the CIFALPS transect. The velocity increase from 25 to 45 km within the slab, in the tomographic model is best reproduced by the transformation of specific lithologies in the high‐pressure granulite facies along a collisional gradient (30°C/km). Although the crust is certainly not completely homogeneous, the best candidates for the rocks that make up the top of the Alpine dip crustal panel are a kinzigite from Monte San Petrone, a gneiss from the Insubric line, and blueschist mylonite from Canavese. While they may not represent the entirety of the crust, they are sufficient to explain the tomographic velocity of the Alpine slab. A lateral lithological contrast inherited from the Variscan orogeny is not required. Eclogitization, suggested as the first‐order transformation in convergence zones, could be a second‐order transformation in collisional wedges. These results also imply a partially re‐equilibrated thermal gradient, consistent with the Alpine thermal state data at depth. Plain Language Summary: Tomography, that is, imaging of deep geological structures based on seismic wave travel to time anomalies, is now so sensitive that it allows us to see changes in the properties of rocks buried at depth beneath mountain belts. In the Alps, the European plate is imaged down to 80 km and shows a sharp velocity increase close to 30 km depth. By calculating the bulk seismic wave velocities on exhumed analogs sampled throughout the Western Alps, the present study proposes to interpret the velocity jump as the consequence of the transformation of the European lower crust from amphibolite to granulite (the high‐temperature metamorphic rocks produced during a collision) rather than the usually admitted transformation to eclogite (the higher pressure metamorphic rocks produced during subduction). This has implications regarding the present‐day thermal structure of the Alps: the Western Alps are not a frozen subduction zone but a collision zone exposing subduction‐related rocks and structures at the surface only. Key Points: The lower crustal top of the European Alpine slab is mostly composed of felsic to intermediate rocksThe transformation of hydrated rocks into HP granulites along a collision gradient reproduces the slab tomographic velocity increase at 30 kmThe often‐supposed eclogitization produces velocities that are significantly higher than the crustal top of the European Alpine slab [ABSTRACT FROM AUTHOR]