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The family Inversiulidae, and its only genus Inversiula, are unique among cheilostome bryozoans owing to the reverse orientation of the operculum, which hinges distally instead of proximally. Inversiula currently includes four extant and one fossil species almost exclusively distributed in the Southern Hemisphere. A phylogenetic analysis of the Inversiulidae based on morphological characters was performed, showing the existence of two clades with contrasting development of the frontal shield. As a result of this analysis, Parainversiula gen. nov. is erected to contain inversiulid species with an umbonuloid frontal shield structure and fewer than 30 central spoked pores. Two poorly known fossil species from the early Miocene of Patagonia (Argentina), originally assigned to the genus Microporella, are here redescribed as Parainversiula sp. and P. chubutiana (Canu) comb. nov. The historical, time-calibrated biogeography of Inversiulidae was estimated using an event-based, multi-model quantitative analysis. The analyses estimated that Australasia was the ancestral area of Parainversiula and a dispersion of the genus into South America occurred between the late Eocene and Oligocene (c. 23–42 Ma), an event that could have been helped by the origin of the Antarctic Circumpolar Current. [ABSTRACT FROM AUTHOR]

This study describes two new species of cheilostome bryozoans associated with gastropod shells from the Monte León Formation (Burdigalian, early Miocene), Argentine Patagonia. Burdwoodipora griffini sp. nov. is the second known species and the first fossil representative of the genus, whose geographic distribution is so far restricted to the Magellan region in the southern southwest Atlantic. it differs from the type species, B. paguricola, in lacking suboral adventitious avicularia in most autozooids. Odontoporella miocenica sp. nov. is morphologically very close to the type species, the Recent O. adpressa, from which it differs in having smaller zooids with fewer areolae and relatively wider orifices. The stratigraphic range of Odontoporella is here extended from the Pleistocene to the early Miocene. its geographic distribution includes southern south America, New Zealand, and New caledonia, suggesting an austral origin of the genus in the Paleogene of former Gondwana. The possible association of these two species with paguroid crabs is discussed. [ABSTRACT FROM AUTHOR]

Abstract: The Paleogene to Neogene San Julián and Monte León formations underlie the central and eastern parts of the Austral Basin (Patagonia, Argentina), and represent coastal plain to shallow shelf systems within a retroarc foreland basin east of the Andes. The formations are characterized by prominent occurrences of thin (<1 m) densely compacted fossil concentrations, including the large oyster Crassostrea? hatcheri. Formation ages have been problematic given macrofaunal provincialism effectively precluding precise extrabasinal correlations. A Sr-isotope chronostratigraphy based on fossil shell calcite from oysters, pectinids and brachiopods is presented for a regional NE–SW-oriented transect extending ∼200 km along strike of the Austral Basin. The Paleogene–Neogene boundary (∼23 Ma) may occur within the uppermost San Julián Formation (22.68 Ma, with a range between 22.22 and 23.22 Ma), coincident with a prominent glauconitic interval, or in a meteorically altered erosional paleosurface that caps this formation and may correlate with an eustatic sea level fall coincident with the global boundary. The younger Monte León Formation is entirely of early Neogene (Aquitanian to early Burdigalian) age, with ages of 22.12 Ma (21.68–22.58) at the base to 17.91 Ma (17.51–18.29) at the top. The proposed new chronostratigraphic framework forms a foundation for future differentiation of local, regional, and global sedimentary patterns in the Austral Basin during the Paleogene/Neogene transition; in particular, with contemporaneous basins and tectonic blocks across the southern polar region, such as in Australia, New Zealand, and Antarctica. [Copyright &y& Elsevier]