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In this paper we ask whether we should we re‐examine the future of upland gamebird management and greater federal oversight and partnerships in the twenty‐first century. Management for waterfowl in North America has been successful because of the 1918 Migratory Bird Treaty Act (MBTA) and the subsequent 1986 North American Waterfowl Management Plan (NAWMP). Although the MBTA included most migratory and non‐migratory species, upland gamebirds, including the northern bobwhite (Colinus virginianus; bobwhite), were excluded and retained under state control. Although many waterfowl populations have been increasing, bobwhite populations have declined precipitously during much of the period. Excluding non‐migratory gamebirds from the MBTA meant that the multistate coordinating efforts that made the MBTA successful for increasing the management of waterfowl have not been applied. The National Bobwhite Conservation Initiative (NBCI) has made a strong effort to unite states within the bobwhite range but does not have the federal anchoring and financial support that were given to states by the MBTA and NAWMP and currently integrate adaptive harvest, habitat management, and financial partnerships to acquire and manage wetlands that support waterfowl production. The NBCI Coordinated Implementation Program (CIP) is designed to serve the function of developing and monitoring habitat for bobwhites but is entirely voluntary and dependent entirely on state and non‐governmental organization (NGO) funds, lacking federal grants and Federal Duck Stamp funds. To catch up with the successes of waterfowl, we discuss the implications of increasing coordination, partnerships, and funding mechanisms between the federal government, state governments, and NGOs to provide common landscape‐level population monitoring and modeling, adaptive harvest regulations, habitat management goals, and a national upland gamebird stamp. © 2021 The Wildlife Society. : This paper explores whether upland gamebirds should have increased federal coordination, like migratory gamebirds, to assure better conservation in the twenty‐first century. The authors discuss the implications of increasing coordination, partnerships, and funding mechanisms between the federal government, state governments, and non‐governmental organizations to provide common landscape‐level population monitoring and modeling, adaptive harvest regulations, habitat management goals, and a national upland gamebird stamp. [ABSTRACT FROM AUTHOR]

The purpose for our essay was to help charge discussion among state agency personnel and others about the clear disparity in what has happened to northern bobwhite during the same time that waterfowl management has taken hold. Waterfowl have been supported by a century of such partnerships to build landscape-level organization, habitat conservation, and enhanced population and harvest estimation (Williams et al. 2021). Bobwhites are not waterfowl. [Extracted from the article]

We examined 318 Colinus virginianus (Northern Bobwhite) wings from 27 Kentucky counties for fault bars. Fault bars are areas in flight feathers that lack pigment due to exposure to stressors. We detected fault bars on 38 (11.9%) wings (representing individual Northern Bobwhites) from 14 Kentucky counties; fault bars were most prevalent in juveniles. Fault bars have been documented on Phasianus colchicus (Ring-Necked Pheasant) and many species of passerine and raptorial birds, but this is the first report for the Northern Bobwhite. Examination of wings for fault bars could provide a new method for assessing stress in Northern Bobwhite populations and be useful to evaluate translocations or to select donor populations for translocation. [ABSTRACT FROM AUTHOR]

As a wildlife biologist, I have often wondered about the tendency of land managers toward planting nonnative species for wildlife, purportedly because of superior nutritional characteristics. Therefore, I analyzed seeds of 26 species of plants for 3 key nutritional factors: gross energy, crude protein, and crude fat. Six of the species were nonnative (including 4 common agricultural crops), whereas the other 20 were native North American plants often reported in wildlife food habits studies. Two crop plants (Helianthus annuus L. [Asteraceae] and Glycine max (L.) Merr. [Fabaceae]), one nonnative plant (Lespedeza bicolor Turcz. [Fabaceae]), and 7 native species had the 10 highest values of crude fat. Glycine max, along with 2 nonnative Lespedeza species and 7 native species had the greatest values of crude protein. For gross energy, Helianthus annuus and Glycine max, along with 8 native species, ranked highest. When comparing the combination of gross energy, crude fat, and crude protein for all species, 4 nonnative and 6 native species ranked highest. This study, the first to test these species with identical procedures, provides comparative evidence that supports using native plants, rather than agricultural and other nonnative species, to supply wildlife winter food. [ABSTRACT FROM AUTHOR]

The lack of adequate documentation of wildlife translocations, particularly details regarding the source stock used, have potentially serious implications for wildlife management. Poor documentation of translocations may lead to unintentional mixing of distinct types, potentially causing problems for future management, design of harvest programs, and evolutionary stability. In Kansas, we employed molecular tools and assignment methods to uncover the cryptic distribution of wild turkey subspecies resulting from decades of poorly documented translocations. Pure forms of Eastern (Meleagris gallopavo silvestris) and Rio Grande turkeys (Meleagris gallopavo intermedia) remain in many portions of the state, and future translocation programs now have the option to keep these distinct by prohibiting translocations between regions containing different subspecies. In addition, we documented 3 zones of hybridization: 1) at the interface between Eastern and Rio Grande subspecies, 2) in southwest Kansas where immigrant Merriam's turkeys (Meleagris gallopavo merriami) are mixing with translocated Rio Grande turkeys, and 3) surrounding an undocumented translocation of Eastern turkeys within a region characterized primarily by Rio Grande turkeys. The DNA-based techniques employed in this study were extremely informative tools for characterizing the distribution of wild turkeys in Kansas and suggest that such tools could be applied in a multitude of similar situations in other wildlife species. [ABSTRACT FROM AUTHOR]

We discuss the necessity of a paradigm shift among managers toward dealing with the recovery and management of prairie grouse (Tympanuchus spp). To assure the future of these species, we will need to test dogmatic assumptions about grouse and their management and challenge the demagogues who insist on perpetuating untested "principles." Tolerance for descriptive and qualitative studies is needed. Additionally, managers will need to remove themselves from the box and embrace landowners, theoretical biologists, economists, human-dimensions researchers, marketing and advertising specialists, and many other professionals outside the normal sphere of wildlife management. There is crucial need for a willingness to devise, test, and apply innovative ideas that are not normally considered in the management of grouse species, especially applying management to large areas within ecosystems. A comprehensive plan is needed to guide rangewide prairie grouse management. Planned management systems are needed to provide operational guidance in implementing species plans. [ABSTRACT FROM AUTHOR]

Deals with a study which compared the attitudes of members of the National Wild Turkey Federation (NWTF) from other turkey hunters in Kansas. Differences between NWTF members and other wild turkey hunters; Attitude toward hunting-related issues; Influence of the mission of NWTF on the attitudes of its members.

A male Odocoileus virginianus (White-tailed Deer) had a 1.42-m segment of large/ small intestine entrapped in its scrotum. This report is the second documented instance of this anomaly, which is common in inbred lines of domestic animals and man. [ABSTRACT FROM AUTHOR]

The article discusses the late flowering of the Tennessee purple coneflower or Echinacea pallida. The critically endangered and federally listed coneflower occurs in three countries in the Nashville Basin, middle Tennessee and its populations can be found in cedar glades, barrens and closely associated prairie. On November 9, 2007, newly emerged flowers on the plant at a cedar glade in Wilson County were found. It points out the insufficiency of heat radiation from exposed rock and gravel to protect plants from frost on November 15-16.

Deals with the lek surveys involving the use and misuse of prairie chickens and other species. Information on density of animal population; Views on the problems regarding the use of lek surveys; How lek surveys are useful in detecting population trends or presence-absence in local areas; Discussion on how it can be valuable in tracking population distribution.

Presents information on a study done on Greater Prairie-chickens. Reference to the observation of 4 Greater Prairie-chicken males; Indication of where the study took place; Indepth look at the study; Findings of the study.

Presents a letter to the editor about the use of volunteer and intern positions by natural resources agencies.

Rural mail carrier surveys (RMCS) have been used to track populations of game animals in the United States for many years, but few states employ RMCS data to track populations of wild turkeys (Meleagris gallopavo). Fourteen years (1987-9000) of RMCS data and hunter harvest data were examined for trends in wild turkey population dynamics in Kansas. Exponential expansion was seen in five of the six regions of the state, the only exception being the southwest, where the population dynamics did not differ significantly from a zero-growth model. Although no population estimates exist for comparison with the relative indices generated from the RMCS data, the residuals from respective exponential models, which represent stochastic fluctuations in turkey abundance, were correlated among adjacent regions of the RMCS, suggesting that the RMCS is internally consistent. Comparisons of RMCS indices with hunter harvest survey data are suggestive of a Type-I functional response curve in the fall hunting season and a Type-II functional response curve in the spring hunting season. Changes in current management strategies are proposed to increase harvest in response to the observed growth of wild turkey populations in Kansas. [ABSTRACT FROM AUTHOR]

Cyclic dynamics of bird and mammal populations are commonly reported in northern latitudes throughout the world, and recent European observations on rodents and grouse suggest that cycle periods decline towards southern latitudes. To investigate latitudinal patterns of cyclic dynamics in North America, we assembled 27 long-term data sets collected between 1939 and 2001 for three grouse species. By fitting the data with autoregressive models to measure direct and delayed density dependence, we show that, in contrast to European studies, North American grouse exhibit period increases from north to south, with cycles collapsing via period lengthening. This occurs because delayed density dependence decreases in southern latitudes, whereas direct density dependence increases. These results show that cyclic dynamics can dissipate by period lengthening as well as the period shortening postulated for European grouse and rodents. [ABSTRACT FROM AUTHOR]

Northern bobwhites (Colinus virginianus) are one of the most broadly researched and intensively managed species in North America. However, we argue that a disadvantage of this status is that traditional management principles currently are incompatible with the spatial scale necessary to address the nationwide decline in bobwhite abundance. We maintain that halting or reversing this decline will entail 2 principal changes in the scale of management. Primarily we suggest that habitat oversight must switch from historical fine-scale management (promotion of edge habitat, weedy fencelines, disked strips, living hedges, and food plots) to regional management of usable space. Secondly, within these regional management areas, we should apply harvest management that employs risk-sensitive strategies that conservatively avoid undermining the primary goal. This entails narrowing the scale of harvest management from statewide to regional levels. If these ideological changes cannot be made and historical policies remain in force, we risk failing to stabilize, let alone increase, bobwhite populations. [ABSTRACT FROM AUTHOR]

We examined how harvest affected natural mortality rates and how the dynamics of northern bobwhite (Colinus virginianus, hereafter bobwhite) covey membership might affect the additive nature of harvest. We conducted repeated searches for coveys and radiomarked all coveys found on 12 259-ha study areas in east-central Kansas, USA. We simulated a harvest in 6 randomly selected areas each year during November-January, 1997-2000, by trapping and removing 60% of bobwhites in each covey. We used flush counts and radiotelemetry to measure harvest effects on natural mortality rates, overall winter survival, covey size, and densities. Observed natural mortality rates of radiomarked individuals were similar between harvested (50.6 ± 4.3%) and unharvested areas (52.1 ± 4.7%). Estimated winter survival was 47.9% on the unharvested areas and 20.9% on the harvested areas. Harvest did not affect the number of coveys, average covey size, or density of bobwhites, indicating that coveys coalesced through local movement. Our results indicate that harvest is additive to natural mortality and suggest that local movement to maintain optimal group sizes can mask the true effect of harvest on observed densities in small areas. [ABSTRACT FROM AUTHOR]

Little is known about how density-independent and density-dependent processes affecting population dynamics vary geographically across species' ranges. To address this problem for three vertebrate species (Northern Bobwhite [Colinus virginianus], Ring-necked Pheasants (Phasianus coichicus], and eastern cottontails [Sylvilagus floridanus]), we examined spatially subdivided, long-term (1966-2001), seasonal (January, April, July, and October), time-series data from the Kansas Rural Mail Carrier Survey, USA. All three species have range boundaries in Kansas, with population abundances declining toward the periphery of their ranges. We quantified the strengths of density-dependent and density-independent processes affecting the dynamics of 10 populations of each species ranging from low (peripheral) to high (central) mean abundance using first-order autoregressive models that incorporate measurement error. For all three species, peripheral populations with lower mean abundance tended to have greater population variability. This pattern could potentially be explained by peripheral regions experiencing either weaker density dependence or greater environmentally driven density-independent fluctuations in per capita population growth rates. In general, density dependence did not vary among geographic regions, although there was a trend for smaller, peripheral populations to exhibit stronger density dependence. Density-independent variability in per capita population growth rates was higher in peripheral populations. Furthermore, density-independent fluctuations in per capita population growth rates were weakly correlated with temperature and precipitation and were highest for the period October through January, identifying fall as the period of greatest environmentally driven variability in population dynamics. Per capita population growth rates fluctuated in moderate synchrony among regions, especially for more abundant, nonperipheral populations in close geographical proximity. The strong density-dependent and stronger density-independent processes in smaller, peripheral populations suggest that the greater variability in peripheral populations' densities is caused by greater population sensitivity to environmental fluctuations. This may make peripheral populations more likely to go extinct and leads to the prediction that, if these species decline to the point of becoming endangered, this decline will be accompanied by a contraction in their geographical ranges. [ABSTRACT FROM AUTHOR]

Northern bobwhite, Colinus virginianus, form social units, called coveys, during the nonbreeding season (approximately September–April). Because the evolutionary advantage of this behaviour is generally unknown, we used controlled group size manipulations within an aviary to investigate whether group size influences (1) the time that the covey spends feeding, (2) the percentage of the covey that is vigilant, (3) the overall vigilance of the group and (4) the time to predator detection. We found that increasing group size increased the time that coveys spent in an exposed feeding area, reduced individual vigilance, improved group vigilance and decreased the time to detection of a potential predator. Additionally, we used experimental reductions of wild northern bobwhite coveys to test whether groups size influences (1) individual and covey survival, (2) daily movement in maintaining covey size and (3) mass change. We conducted field research on 12 independent 259-ha study areas (6 control plots and 6 treatments, where 60% of the population was removed) in east-central Kansas, U.S.A. between 9 November and 31 January, 1997–2000. We radio-marked 386 radiocollared individuals that comprised 137 groups on the study areas. Covey size did not differ between or within years or treatments (X±SE: 10.98±0.22 individuals). Our results indicate that a stable group size existed between 1 and 22 individuals, with 11 being an optimal group size. Small coveys (1–7 individuals) had lower group persistence and individual survival, and used increased movement to create or join larger groups where survival was higher. Large groups (15–22) had lower individual survival, increased group movement and individual mass loss. Density-dependent feedbacks (e.g. lower survival and increased competition) may have lowered larger coveys to a stable size. Our results suggest the regulation of an optimal covey size of 11 was promoted by high group persistence, low group movement, improved feeding efficiency, improved individual predator detection and improved individual survival. Copyright 2003 Published by Elsevier Ltd on behalf of The Association for the Study of Animal Behaviour. [Copyright &y& Elsevier]

The article examines the host range and geographic distribution of Ophidiomyces ophiodiicola, the fungal pathogen responsible for snake fungal disease (SFD), in Tennessee, U.S. It highlights the prevalence of the pathogen among various snake species across different counties and identifies new host species affected by SFD, underscoring the need for comprehensive sampling and standardized detection methods to better understand the pathogen's impact.

ABSTRACT Oak ( Quercus spp.) woodlands and savannas throughout the Mid-South, USA have become closed-canopy forests through succession in the absence of fire, contributing to declines in disturbance-dependent avian species. Restoration could ameliorate these trends, but effects of such management practices on avian communities of the Mid-South are poorly understood. We implemented an experiment on the Cumberland Plateau, Tennessee with treatments involving fall (Oct) or spring (Mar) fire with woodland (14 m2/ha) or savanna (7 m2/ha) residual basal area and unmanaged control stands (20 ha each). We conducted fixed-radius point counts 3 times annually (2010-2012) and analyzed trends in naïve occupancy for 41 species. For 20 species with adequate data (10 early-, 10 late-successional), we performed multiple-season, robust-design occupancy modeling and accounted for detection bias. We then used mixed-model polynomial regression to define relationships between occupancy and measures of live and dead basal area, midstory density, and herbaceous groundcover. Restoration substantially altered forest structural characteristics but did not affect the occupancy of most late-successional species. In contrast, the presence of early-successional species increased as live basal area decreased and herbaceous groundcover increased. Only 3 of 41 species (hooded warbler [ Setophaga citrina], ovenbird [ Seiurus aurocapilla], and worm-eating warbler [ Helmitheros vermivorum]) exhibited reduced occupancy as woodland and savanna conditions were approached. The presence of all other species, including 13 regionally and 2 continentally declining, remained constant or increased as restoration progressed. Woodland conditions of 10 m2/ha live basal area and 20% herbaceous groundcover maximized occupancy of early-successional species with little consequences for late-successional species. Proceeding further toward savanna conditions negatively affected species associated with lower strata of mature-forests but further increased the presence of some early-successional species. Midstory density and dead basal area were poor predictors of occupancy and thus may be necessary but not sufficient in promoting early-successional species. Our results suggest implementing oak woodland and savanna management in the Mid-South could benefit disturbance-dependent birds of high conservation priority with minimal negative impacts on the presence of late-successional species. © 2016 The Wildlife Society. [ABSTRACT FROM AUTHOR]

ABSTRACT Grassland birds have declined more than any other guild in the United States because of loss and degradation of native grasslands. The United States Department of Agriculture Farm Bill programs have restored some native warm-season grasses (NWSG), but populations of many grassland bird species continue to decline. Market-based NWSG uses focused on hay, pasture, and biofuel feedstock may be more appealing to landowners, and may still provide grassland bird habitat on the landscape. We examined breeding grassland bird occupancy of 102 NWSG production fields including 5 treatments (control [ n = 37], grazing [ n = 7], hay [ n = 22], seed [ n = 21], and biofuel [ n = 15] production) in Kentucky and Tennessee during 2009-2010 breeding seasons. We used a robust design model in Program MARK to determine occupancy and detection rates for grassland birds. We modeled occupancy differences among treatments and sites (i.e., KY, TN), and included covariates (i.e., field-level vegetation metrics and landscape composition at 3 scales [250 m, 500 m, and 1 km]) for eastern meadowlark ( Sturnella magna), field sparrow ( Spizella pusilla), grasshopper sparrow ( Ammodramus savannarum), northern bobwhite ( Colinus virginianus), and red-winged blackbird ( Agelaius phoeniceus). Covariates that influenced occupancy included treatment (field sparrow, 2009 only), site (eastern meadowlark, grasshopper sparrow, and northern bobwhite), percent NWSG cover (positive, red-winged blackbird), and percent forest cover within 250 m (negative for eastern meadowlark, grasshopper sparrow, northern bobwhite, and red-winged blackbird) or within 1 km (negative, field sparrow). Our results suggest that forest cover in the surrounding landscape negatively influences species occupancy, and species occupancy generally did not differ among NWSG production treatments. These treatments could be an alternative means to provide grassland bird habitat within an agriculture production landscape. © 2016 The Wildlife Society. [ABSTRACT FROM AUTHOR]

The article presents a study which examines the distribution of mitochondrial DNa (mtDNA) variation of the lesser prairie-chicken in southern Great Plains. The declining population of the species in the area calls for a better understanding of its genetic variation to derive efficient conservation measures. The study found out that there is no connection between haplotype phylogency and geography, however, suggests to evaluate conservation actions to increase genetically viable population.

The lesser prairie-chicken (Tympanuchus pallidicinctus) is currently considered a candidate for protection under the Endangered Species Act. To identify potential limiting factors for lesser prairie-chicken populations, we developed an age-based matrix model of lesser prairie-chicken population dynamics to compare the relative importance of components of reproduction and survival, and determine if various management alternatives stabilize or increase rates of population change. We based our analyses on an intensive 6-year population study from which demographic rates were estimated for each age class in Kansas. We used deterministic models and elasticity values to identify parameters predicted to have the greatest effect on the rate of population change (l) at 2 study sites. Last, we used life-stage simulation analysis to simulate various management alternatives. Lambda was ,1 for both populations (site 1: l 5 0.54, site 2: l 5 0.74). However, we found differences in sensitivity to nest success and chick survival between populations. The results of the simulated management scenarios complemented the lower-level elasticity analysis and indicated the relative importance of female survival during the breeding season compared with winter. If management practices are only capable of targeting a single demographic rate, changes to either nest success or chick survival had the greatest impact on l at site 1 and 2, respectively. Management that simultaneously manipulated both nest success and chick survival was predicted to have a greater effect on l than changes in survival of adult females. In practice, our demographic analyses indicate that effective management should be based on habitat conservation measures to increase components of fecundity. [ABSTRACT FROM AUTHOR]

Declines in native warm season grasslands have been linked to grassland bird population decline due to habitat loss including conversion to non-native grasses. Rotational grazing (ROT) and patch-burn grazing (PBG) are two possible tools to restore native warm-season grasses (NWSG) on working-lands in the Mid-South USA and thus aid in the recovery of grassland bird populations. This project compares ROT, PBG, and before treatment implementation to assess their effects on grassland-associated bird species. At three research sites between KY and TN, 14–10 ha NWSG pastures were established and randomly assigned 7 pastures each to ROT and PBG treatment and monitored avian relative abundance during the breeding season from 2014 to 2017. Avian call count data and vegetation characteristics were collected in 2014 and treated as a before treatment year. Following 2014, ROT and PBG treatments were implemented across each respective research site. We used the open N-mixture model framework to estimate avian relative abundance related to year, treatments, research site, and landscape and within-field variables. Avian species richness and diversity were calculated for each treatment, research site, and year. Landscape variables, within-field variables, and research sites exerted more influence on relative abundance than ROT or PBG. Grassland-associated bird species relative abundance and species richness/diversity were affected by habitat disturbances (both ROT and PBG) but varied by species and site. Field sparrows [ Spizella pusilla ] had the highest increase in relative abundance (9.68 ± 1.24 birds/point count location or 1.77 ha) while northern cardinals [ Cardinalis cardinalis ] exhibited a significant decrease in relative abundance (3.44 ± 1.54 birds/point count location) following treatment implementation on specific research sites. Species diversity and richness did not differ between ROT and PBG treatments. However, a site and year difference were observed for both estimates. Using ROT and/or PBG to create habitat disturbances can alter within-field variables (i.e., vegetation height) which, taken into context with landscape variables, could impact grassland bird populations and diversity depending on grassland bird species habitat requirements. Our research provided the baseline information for ROT and PBG impacts on grassland birds in the east/southeastern USA. However, we believe future research should focus on breeding and annual fecundity to better understand how populations will change over time and how working lands conservation might aid this conservation effort without a reduction in livestock productivity. • Grassland bird populations decline has been linked to habitat loss. • Utilizing private working-lands management could increase potential avian habitat. • Rotational grazing and patch-burn grazing create vegetation structural diversity. • Increased vegetation structural diversity can benefit avian relative abundance. • Habitat disturbance/landscape can impact grassland bird richness and diversity. [ABSTRACT FROM AUTHOR]

The northern bobwhite (Colinus virginianus) is an economically important gamebird that is currently undergoing widespread population declines. Despite considerable research on the population ecology of bobwhites, there have been few attempts to model population dynamics of bobwhites to determine the contributions of different demographic parameters to variance of the finite rate of population change (λ). We conducted a literature review and compiled 405 estimates of 9 demographic parameters from 49 field studies of bobwhites. To identify demographic parameters that might be important for management, we used life-stage simulation analyses (LSA) to examine sensitivity of λ to simulated variation in 9 demographic parameters for female bobwhites. In a baseline LSA based on uniform distributions bounded by the range of estimates for each demographic parameter, bobwhite populations were predicted to decline (λ = 0.56) and winter survival of adults made the greatest contribution to variance of λ (r² = 0.453), followed by summer survival of adults (r² = 0.163), and survival of chicks (r² = 0.120). Population change was not sensitive to total clutch laid, nest survival, egg hatchability, or 3 parameters associated with the number of nesting attempts (r² <0.06). Our conclusions were robust to alternative simulation scenarios, and parameter rankings changed only if we adjusted the lower bounds of winter survival upwards. Bobwhite populations were not viable with survival rates reported from most field studies. Survival rates may be depressed below sustainable levels by environmental conditions or possibly by impacts of capture and telemetry methods. Overall, our simulation results indicate that management practices that improve seasonal survival rates will have the greatest potential benefit for recovery of declining populations of bobwhites. [ABSTRACT FROM AUTHOR]

Long-term population declines and habitat reductions have increased concern over the status of the lesser prairie-chicken (Tympanuchus pallidicinctus). Robust estimates of demographic parameters are essential for identifying population declines and planning effective management. We evaluated the effects of age and season on the survival of female lesser prairie-chickens at 2 sites in southwestern Kansas, USA. Using telemetry data from a 7-year field study (from 1997 to 2003), we estimated seasonal (Apr-Sep) and annual (Apr-Mar) survival. We also examined daily survival rates of females attending nests during the 26-day incubation period and young during the 14-day early brood-rearing period. We evaluated the probable mortality causes of radiomarked birds by examining evidence at recovery sites. We captured 227 female lesser prairie-chickens (87 yearlings, 117 ad, and 23 age undetermined) and fitted them with radiotransmitters. Estimates of 12-month survival were lower among yearlings (Ŝ12 = 0.429, SE = 0.117) and adults at site I (Ŝ12 0.302, SE = 0.080) than among yearlings (Ŝ12 = 0.588, SE = 0.100) and adults at site II (Ŝ12 = 0.438, SE = 0.083). The patterns in timing of mortality and age-specific 6-month survival were consistent with those of 12-month estimates at site I from 1998 to 2002, with a peak in mortality during May and June. Females tending to nests or to prefledged chicks had lower daily survival (DŜRtend = 0.993, SE = 0.001) than females not involved in these activities (DŜRfailed-breeder = 0.997, SE = 0.002). We recorded 92 mortalities from April 1997 to March 2003, and 59% and 11% were attributed to predation by mammals and raptors, respectively. Our research suggests that predation during the nesting season can have a major impact on lesser prairie-chicken demography, and conservation efforts should focus on enhancing female survival during the nesting and brood-rearing seasons. [ABSTRACT FROM AUTHOR]

Radiotelemetry has provided wildlife biologists with a tool to estimate survival where fates of individuals likely are known. Analyses of known-fate data can yield accurate survival estimates if 5 assumptions are met. Two of these assumptions are rarely tested: that transmitters have no effect on survival of study animals and that right-censoring (i.e., any animal not located is as likely to be alive as dead) is random with respect to the survival of study animals. Using joint-models originally developed for live-encounter and dead-recovery data, we examined the potential for bias in survival estimates of radiomarked male lesser prairie-chickens (Tympanuchus pallidicinctus) in a 3-year study in southwestern Kansas, USA. Additionally, we examined the potential bias of right-censoring by comparing the return rates of known-fate and right-censored individuals. We captured 216 male lesser prairie-chickens and marked them with a combination of leg bands and a radio (n = 72) or leg bands only (n = 144). We applied joint-models to capture histories based on live-capture and telemetry data. The model best supported by the data indicated that 6-month survival was constant (Ŝc 0.679, SE 0.050) across radiomarked and banded birds. Eight of 16 (50%, SE = 12.5%) right-censored birds not detected because of radio failure were subsequently recaptured, which was not different from the return rates for known-fate birds (23 of 59; 39%, SE = 6.3%). Survival estimates of male lesser prairie-chickens in this study were not measurably biased by radiomarking, as their survival was greater than or equal to those of banded birds, and right-censored birds had similar return rates to those of known-fate individuals. Our results are encouraging because they indicate that 2 critical assumptions underlying analyses of known-fate data can be met with radiotransmitters and attachment techniques currently used in field studies of wild populations of lesser prairie-chickens. [ABSTRACT FROM AUTHOR]

Juvenile survival has been identified as the most critical demographic parameter influencing grouse populations. Little information currently exists on survival of juvenile lesser prairie-chickens (Tyrnpanuchus pallidicinctus). We regularly flushed 51 individually identifiable lesser prairie-chicken broods over a 6-year period to estimate survival from hatch to 14 days post-hatch (early period) and from 15 to 60 days post-hatch (late period). Estimates of overall daily survival rates were 0.949 (95% CI = 0.932-0.966) for the early period and 0.978 (95% CI = 0.908-0.989) for the late period. Overall survival from hatch to 60 days posthatch was 0.177 (95% CI = 0.028-0.376). We used encounter histories of 31 transmitter-equipped juveniles to estimate survival from 1 August to 31 March (overwinter) using known-fate models. Juvenile overwinter survival was 0.70 (95% CI = 0,47-0.86), and chicks heavier than average for their age at 50-60 days posthatch were more likely to survive the 8-month overwinter period. Survival of juveniles from hatch to 31 March of the following year was 0.12 (95% CI = 0.01-0.32). We compared overwinter survival of juveniles and 93 transmitter-equipped full-grown lesser prairie-chickens using a second set of models. Overwinter survival rates for juveniles (0.64) and full-grown (0.63) birds were similar, but the timing of mortality events differed between age-classes. We recommend that managers in Kansas, USA, focus on improving early survival of juveniles by providing additional food resources to chicks. This can be accomplished by manipulating vegetation to increase forb cover, which will result in increased invertebrate biomass. [ABSTRACT FROM AUTHOR]

Wildlife managers depend on accurate information regarding wild turkey (Meleagris gallopavo) survival patterns to properly manage turkey populations. Survival patterns of male Rio Grande wild turkeys (M. g. intermedia) have not been studied intensively. Wildlife managers in the Texas Panhandle, USA, and southwestern Kansas, USA, suspected that turkey populations were declining. From January 2000 through August 2002, we studied survival and movement of 107 juvenile male and 115 adult male rediomarked Rio Grande wild turkeys on 4 study sites in the Texas Panhandle and southwestern Kansas. We predicted that males would experience lowest survival during spring and that there would be no difference in survival between age classes. We also predicted that greater male movement rates would lead to lower survival rates. Juvenile males had a higher annual survival rate (0.597 [95% Cl hereafter: 0.478-0. 716]) than adults (0.364 [0.257-0.472]). Juvenile male survival did not differ among seasons, with survival rates of 0.813 (0. 736-0.891), 0.904 (0.837-0.972), and 0.917 (0.838-0.996) for spring, summer, and autumn, respectively. Adult male turkey survival was higher during summer (0.915 [0.859-0.972]) than during spring (0. 725 [0.651-0. 799]), autumn (0.671 [0.536-0.807]), and winter (0. 792 [0.732-0.851]). Males had lower survival rates during seasons when long-distance movements were common. The annual survival rate for turkeys that moved to new core-use areas (0.383 [0.282-0.484]) was lower than that for turkeys that did not (0.535 [0.460-0.609]). Also, survival rates increased with time since relocation of core-use areas. Hunting accounted for 18.5% of all mortalities. However, most (80.7%) mortality was attributed to natural causes, mostly mammalian predation. We suspected most predation was the result of coyotes (Canis latrans) and bobcats (Lynx rufus). Managers in the northern portion of the natural range of Rio Grande wild turkeys should be aware of the presence of natural mortality factors that are evident in lightly hunted populations. Managers interested in increasing the survival of male Rio Grande wild turkeys should concentrate on efforts that will provide needed resources in close proximity to roosts. [ABSTRACT FROM AUTHOR]

Despite the fact that the Lesser Prairie-Chicken (Tympanuchus pallidicinctus) is a species of conservation concern, little is known about its nesting ecology, particularly in sand sagebrush (Artemisia filifolia) habitats. To find and monitor nests, we captured and equipped 227 female Lesser Prairie-Chickens with transmitters (87 yearlings, 117 adults, and 23 of unknown age) from 1997 to 2002 in southwestern Kansas. Apparent nest success was similar for yearlings (31%, n = 74) and adults (27%, n = 97) but differed marginally (P = 0.090) between first nests (29%) and renests (14%). An estimated 31% of females that were unsuccessful in their first nesting attempt initiated a second nest. The probability that a female would initiate a second nest after failure of the initial attempt was negatively influenced by the day of incubation on which the initial attempt failed. Over 95% of all nests were initiated and completed between 5 May and 2 July. The primary cause of nest failure was predation by cayotes (Canis latrans) and gopher snakes (Pituophis melanoleucus). Mean clutch size, egg fertility, hatching success, nesting and renesting frequency, and incidence of interspecific parasitism were all similar across years and between yearlings and adults. Distances between nest sites were used as an index to nest-site fidelity between first nests and renests and for across-year nesting attempts. Mean distances between first nests and renests were similar for yearlings (1,071 m) and adults (1,182 m). Mean distance between nests constructed by the same female in subsequent years (918 m) did not differ between age classes or success of the first year's nest. Most females (80%) nested closer to a lek other than the lek where they were captured. [ABSTRACT FROM AUTHOR]

invertebrates are an important food source for grouse chicks, especially within the first 2 weeks of life, invertebrate abundance is highly patchy and dependent upon herbaceous cover and vegetation structure. We examined the relationship between invertebrate bio- mass (from sweepnet samples) and habitat structure at lesser prairie-chicken (Tympanuchus pallidicinctus) brood-use and non-use areas during 2001 and 2002 in a sand sagebrush (Artemisia filifolia) prairie vegetation community of southwestern Kansas. We delineated use and non-use areas from paired sampling points within and outside 95% utilization distributions of radiomarked brood females, respectively, during the first 60 days post-hatch. We measured vegetation cover and invertebrate biomass (Acrididae and "other" invertebrates) at 71 paired points on 2 study sites (Site 1=4 broods, Site 11= 12 broods). Both Acrididae and other invertebrate biomasses were greater at brood areas than non-use areas on both study sites, suggesting this food source likely had a greater influence on brood habitat use than vegetation type. Vegetation structure described brood-use areas better than vegetation type because brood-use areas had greater visual obstruction readings (VORs) than non-use areas regardless of dominant cover type. We also examined the predictive relationship between vegetation type and invertebrate biomass. Sand sagebrush density was the best linear predictor of Acrididae biomass, with lower densities having the greatest Acrididae biomass. We propose experiments to deter- mine best management practices that produce abundant invertebrate biomasses for lesser prairie-chicken brood habitat, using our study as a baseline. [ABSTRACT FROM AUTHOR]

Lesser prairie-chicken (Tympanuchus pallidicinctus) populations have declined rangewide, and one of the principal causes is thought to be low nest success. Little is known about the relationship of vegetation structure and human intrusion to lesser prairie-chicken nest location and success. We conducted our study from 1997 to 2002 in southwestern Kansas, USA, on 2 sand-sagebrush (Artemisia filifolia) prairie areas managed for livestock production. We determined apparent nest success (26%) for 200 of 209 lesser prairie-chicken nests located. Nest sites had taller grass, greater sand-sagebrush density, and higher visual obstruction than random locations in the surrounding prairie. We recorded the distances from nests to 6 anthropogenic features (wellheads, buildings, improved roads, unimproved roads, transmission lines, center-pivot irrigation fields) to determine whether the features were related to nest location and success. Sand-sagebrush habitat around S of 6 features (all except unimproved roads) was avoided for 80 m (wellheads) to >1,000 m (buildings) by nesting lesser prairie-chickens, but distances to the features were not substantial predictors of apparent nest success. Grass height, sagebrush plant density, and sagebrush height were the most important vegetation characteristics influencing nest success. [ABSTRACT FROM AUTHOR]

We used mark-recapture methods to estimate age-specific apparent survival rates for male Lesser Prairie-Chickens (Tympanuchus pallidicinctus), a gamebird of conservation concern. A total of 311 male prairie-chickens (135 yearlings, 176 adults) were captured and banded during a 5-year study in southwest Kansas. Time-since-marking models were used to estimate apparent survival after first capture (φ¹), apparent survival among returning birds (φ2+), and probability of capture (p) for yearling and adult prairie-chickens. Apparent survival is the product of true survival and site fidelity, and our model-averaged estimates of this parameter were ranked: yearlings after first capture (φ¹yr = 0.60 ± 0.12) > adults after first capture (φ¹ad = 0.44 ± 0.10) > returning birds (φ2+ = 0.36 ± 0.10). In contrast, movement data showed that site fidelity to communal display sites (or leks) increased with male age; yearlings returned to leks at lower rates (80%, n = 60) than adults (92%, n = 65). Thus, true survival rates of male Lesser Prairie-Chickens likely decline with increasing age, an unusual pattern found in few species of birds. We hypothesized that declines in survival as males' age may be a feature of promiscuous mating systems where competition for mating opportunities are intense. A review of annual survival rates for holarctic grouse did not support this idea; age-specific declines in male survival were not restricted to lek-mating species, and appear to be a general feature of most grouse populations. [ABSTRACT FROM AUTHOR]

We derived consequences (realizations of hunter efficiency, relative harvest rates) of fixed, liberal quail (northern bobwhite [Colinus virginianus], Gambel's quail [Callipepla gambelii], and scaled quail [C. squamata]) harvest regulations applied at large scales from time series on quail abundance, total harvest, and hunter participation. Data came from Kansas (1966-2001), Missouri (1983-2001), Oklahoma (1990-2001), north and south Texas (1986-2001), and Arizona (1982-1999), USA, where harvest regulations were liberal (season length 2.5-4 months, daily bag limit 8-15 birds) during the periods of record. For all study regions, hunter-days were expressible as a linear function of quail abundance, and total harvest was expressible as a linear function of hunter-days. These results implied that hunter efficiency (harvest/hunter-day/index bird) declined monotonically and curvilinearly as quail populations increased. Likewise, relative harvest rate declined monotonically and curvilinearly as abundance increased, which implied that harvest was not self-limiting; however, the rate of decline generally was low because harvest rate was the product of an increasing (hunter-days) and a decreasing function (hunter efficiency) of quail abundance. Under fixed, liberal regulations, variations in quail abundance seem to govern harvest rates at the state or regional level; the regulations per se probably are biologically inconsequential. [ABSTRACT FROM AUTHOR]