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Crustal deformation and hydrothermal percolation related to the India‐Asia collision have caused extensive remagnetization of the Tethyan Himalaya Terrane (THT). The present work identified three phases of regional remagnetization during 62.3–50.0 Ma for the east‐central THT. Consequently, a model of three‐stage India‐Asia collision was proposed. The east‐central THT first collided with the southward migrated southern margin of the Lhasa Terrane (LT) at 5.4 ± 0.9°N during 62.3–60.9 Ma. Subsequently, the THT continuously moved northward and pushed the southern margin of the LT back to its original position prior to the initiation of fore‐arc and back‐arc extension on both sides of the Gangdese magmatic arc. Since the final suturing of the THT with Asia at ∼10°N during 59.8–58.0 Ma, the east‐central THT remained stationary until India collided with it at 10.9 ± 5.1°N at ∼50.0 Ma. Plain Language Summary: The collision of India and Asia caused intense tectonic deformation and hydrothermal alteration throughout the Tethyan Himalaya Terrane (THT), which resulted in the large‐scale remagnetization in the THT. The regional remagnetization of the THT can be used to constrain the India‐Asia collision process, on the premise that the time of remagnetization can be determined. Based on this assumption, we measured two representative Paleocene remagnetized components from Early Jurassic limestones in the Gyangze Basin in the east‐central THT. These remagnetized components, combined with non‐remagnetized components and remagnetization events recorded in the adjacent areas, suggest that the east‐central THT experienced three phases of regional remagnetization during 62.3–50.0 Ma. The first and second phases of remagnetization in the north‐central part of the east‐central THT occurred at the paleolatitude of 5.4 ± 0.9°N at 62.3–60.9 Ma and 10.3 ± 1.0°N–9.5 ± 1.1°N at 59.8–58.0 Ma, respectively. The third phase of remagnetization occurred in the southern part of the east‐central THT, at the paleolatitude of 10.9 ± 5.1°N at ∼50.0 Ma. Consequently, a model of three‐stage India‐Asia collision and southward spreading tectonic deformation of the THT was proposed based on these successive remagnetizations. Key Points: The east‐central Tethyan Himalaya Terrane (THT) experienced three phases of large‐scale remagnetization during 62.3–50.0 MaThe collision of THT with the Lhasa Terrane commenced at 62.3–60.9 Ma and finished at 59.8–58.5 MaIndia finally collided with the THT at the paleolatitude of ∼10.9 ± 5.1°N at ∼50.0 Ma [ABSTRACT FROM AUTHOR]

Microfaunal analysis of Cretaceous and Paleogene sediments collected over several decades from Western Siberia has illuminated the diversity, stratigraphic distribution, and paleoecological signifi cance of the rare low trochospiral benthic foraminiferal genus Bagginoides. Here, we describe two new species B. yamalensis n.sp. and B. tibeisalinella n.sp [ABSTRACT FROM AUTHOR]

Menispermaceae are an angiosperm family of mostly climbing plants distributed throughout tropical regions. The fruits in this family have a strong sclerified endocarp and can be used for reliable species‐level identifications, even in a fossilized form. New Paleocene‐age menispermaceous endocarps have recently been discovered in South America and Asia, while in Europe, they are mostly found in the Eocene. This paper focuses on the study of fossil endocarps belonging to Menispermaceae, found in the Petit‐Pâtis locality, which is one of the few Paleocene localities in Europe. The locality is dated to the late Paleocene (57–56 Ma) and the fossils represent flora and fauna in the Paris Basin before the onset of the Paleocene–Eocene Thermal Maximum. Lignitized fossils were photographed and their morphology was analyzed using computed tomography, scanning electron microscope, and geometric morphometric analysis. The analysis of 36 fossils belonging to Menispermaceae revealed that three genera (Stephania, Palaeosinomenium, Tinospora) were present in the late Paleocene of Paris Basin. Differences in internal morphology and a more inflated endocarp base led to a description of a new fossil species—Palaeosinomenium oisensis sp. nov. The occurrence of these genera in the Paleocene of France is consistent with the suggestion that megathermal flora was present in the Paris Basin before the Paleocene–Eocene Thermal Maximum. These three genera are currently the oldest occurrences in Europe, highlighting the connection between paleofloras of the Americas, Europe, and Asia in the Paleocene. [ABSTRACT FROM AUTHOR]

The post-collisional evolution of the Tibetan lithosphere is of paramount significance to our understanding of collisional orogeny. It is generally postulated that the Lhasa lithospheric mantle was horizontally shortened and thickened coherently with the overlying crust to form a physical barrier, preventing Indian subduction beneath Tibet until the thickened mantle root was foundered during the Miocene. This study first identifies post-collisional oceanic-island basalt (OIB)-type magmatism in the Lhasa Block (LB), as attested by zircon U-Pb age (ca. 58 Ma) and geochemistry--positive Nb-Ta anomalies, high La/Yb, and depleted bulkrock Sr-Nd and zircon Hf isotopes, of diabase in the northern (inboard relative to Indus Suture) part of this block. Coupled with extensive early Paleogene arc-type magmatism in the southern-central LB and thermodynamic modeling, we suggest that these diabases were formed by partially molten upwelling asthenosphere near the base of continental crust, where much of the underlying lithospheric mantle had been removed due to Neo-Tethyan slab rollback and lithospheric delamination. Compared to OIB-type magmatism worldwide, the diabases investigated here were emplaced peculiarly in a region where the continental crust was under horizontal compression and shortening by coeval thrusting. Our study thus implies a decoupled deformation between the crust and mantle of the LB during the early Indian-Asian collision. [ABSTRACT FROM AUTHOR]

We describe new macro- and microfossils recovered on a field expedition to the North Quarry of Poponguine, a locality in western Senegal that spans the Late Cretaceous through the Early Paleogene, albeit with a likely unconformity at the base of the Danian. Newly discovered macrovertebrates from the Maastrichtian Cap de Naze Formation include pycnodonts, dyrosaurids, and chelonians, the latter two the oldest and first Cretaceous representatives of these clades from Senegal. Screenwashing of this deposit revealed that the matrix also consists of abundant microscopic biological clasts comprising osteichthyan vertebrae, spines, and cranial fragments, shark dermal denticles, invertebrates, and numerous ovoid coprolites among other fragments. These microfossils document a much less conspicuous portion of the paleobiodiversity. We describe the stratigraphy of this new locality and incorporate it into a new correlated section that ties together three other Late Cretaceous–Early Paleogene fossiliferous localities of the Senegalese-Mauritanian Basin. The presence of fossiliferous Danian rocks, which are notably rare in West Africa, is elucidated in our section, which synthesizes prior geological and paleontological work around Poponguine and in the nearby coastal region of Ndayane. Primarily dated using the biostratigraphy of ostracods and foraminiferans, the correlated rock units include the Paki and the Cap de Naze Formations of the Late Cretaceous Diass Group and the Ndayane and the Poponguine Formations of the Paleocene Cap-Vert Group, deposits that capture ancient near shore marine environments. The localities described, although separated by only a few kilometers, exhibit dramatic differences in thickness across faulted blocks of the Diass Horst. Dyrosaurids, which are common fossils in nearshore marine outcrops of West Africa, have been hypothesized to be of African origin and to have dispersed to South America in the Late Cretaceous. This dyrosaurid specimen, although fragmentary, documents a western extreme in the geographic range of African dyrosaurids where the clade would have been well situated for broader trans-Atlantic dispersal. [ABSTRACT FROM AUTHOR]

The kinematics and deformation pattern along the Altyn Tagh fault (ATF), one of the largest strike‐slip faults on Earth is of great significance for understanding the growth of the Tibetan Plateau. However, the initial rupture along the ATF remains debated given the limited constraints on the depositional age of associated Cenozoic syntectonic strata. Here we investigated the syntectonic Cenozoic strata in the Xorkol Basin, associated with the strike‐slip faulting along the ATF. New uranium‐lead analyses of the carbonate deposits in the Paleogene strata yield dates of 58.9 ± 1.29 Ma, representing the initial rupture of the ATF. This first documented radioisotopic age coincides with the ca. 60 Ma onset timing of India‐Asia collision, highlighting its far‐field effect at the northern edge of the Tibetan Plateau. We infer that the deformation of the entire Tibetan Plateau started synchronously with the India‐Asia collision. Plain Language Summary: Carbonate U‐Pb dating techniques applied to rocks associated with the Altyn Tagh fault, a major fault in North Tibet, reveal that the fault started slipping about 58.9 million years ago, coinciding with the time when India collided with Asia. This finding provides new constraints on when and where this fault formed and suggests that the northern Tibetan Plateau started deformation earlier than previously thought. This result emphasizes that the entire Tibetan Plateau deformed simultaneously in the early Cenozoic. Key Points: Calcite U‐Pb dating yields ca. 59 Ma age for carbonate strata in the East Xorkol BasinXorkol Basin was a pull‐apart basin during the Paleocene due to the left‐lateral strike‐slip faulting along the Altyn Tagh faultWidespread Paleocene‐Eocene tectonism in Northern Tibet highlights the far‐field effect of the India‐Asia collision [ABSTRACT FROM AUTHOR]

The Arabia – Eurasia collision zone in the central part of the Alpine – Himalayan orogenic system has had a complex deformation history since the Palaeozoic. In Iran, the collision zone consists of the Alborz‐Talesh, Kopeh Dagh and Zagros foldbelts and the intervening Central Iran area. In this review paper, we summarize the structural architecture and tectonostratigraphic characteristics of these domains and attempt to correlate regional deformation events between them. The results show that six regional‐scale deformation phases can be recognized and correlated in Iran over a time interval extending from the Late Palaeozoic to the Late Cenozoic. Late Palaeozoic rifting in northern Gondwana and subsequent oceanic spreading resulted in the separation of the Central and North Iran blocks from the Arabian Platform. These blocks later converged and collided with the southern margin of Eurasia due to the subduction of the intervening PalaeoTethys lithosphere ("Cimmerian orogeny": Late Triassic). The convergent setting resulted in the initial development of the Alborz‐Talesh foldbelt in present‐day northern Iran, while extensional basins developed in the forebulge area in Central Iran. Continuing northward subduction of NeoTethyan oceanic lithosphere at the southern Eurasia margin produced Early Cretaceous back‐arc extension and associated volcanism in Central Iran and the Alborz‐Talesh area to the north. A phase of compressional deformation in the Late Cretaceous was related to the collision of a series of microcontinents derived from Northern Gondwana, including the Ercinjan and Bitlis massifs, with the Central Iran block, and is recorded in the Alborz‐Talesh foldbelt and in Central Iran. Further back‐arc extension in the late Paleocene – Eocene was accompanied by pervasive volcanism and volcaniclastic sedimentation throughout northern and Central Iran. The final closure of NeoTethys and convergence between the Arabian and Eurasian Plates evolved through phases of early Oligocene "soft" collision and middle Miocene "hard" collision. This was accompanied by thrusting in the internal parts of the Zagros foldbelt and by folding and subordinate thrusting in the more external parts, with related development of the flexural Mesopotamian Basin in the foreland to the SW. [ABSTRACT FROM AUTHOR]

Premise of research. Macginitiea is an iconic and easily recognizable genus well known from the Paleogene of western North America and Kamchatka. Here we recognize new species and occurrences of Macginitiea from multiple sites in North America and discuss their implications for biogeography and evolution. Methodology. We studied leaves and inflorescences from multiple sites across North America. We used microcomputed tomography scans to illustrate the floret anatomy of Macginicarpa from Texas. Cuticles were examined using epifluorescence microscopy and maceration in Jeffrey's solution. Pivotal results. The distribution of Macginitiea suggests transberingial migration, similar to that in other platanoids like Platimeliphyllum. Macginitiea lived as far south as central Texas; few fossil platanoids are known from southern latitudes, even though most living Platanus species occur below 30°N. We document the cuticle anatomy of M. whitneyi , revealing a heavily striated cuticle with circular hair bases. Newly recognized species include those with a striking proclivity for basilaminar lobe development, including M. rannii sp. nov. from the Eocene of Wyoming, M. basilobata comb. nov. from the Paleocene of Montana, and M. rileyi comb. nov. from the Eocene of Texas and New Mexico. We also revise the taxonomy of Aralia wardiana (formerly Aralia digitata) and Platanus browni and show their affinities to Macginitiea. Using a broad concept of the gene regulatory networks governing lobe and stipule development, we hypothesize how different Macginitiea leaves could have developed their basilaminar lobes. Conclusions. We find that Macginitiea had a greater taxonomic diversity and a wider geographic range than was previously known. Among this diversity are several species with substantial basilaminar lobe development. Basilaminar lobes in modern and fossil platanoids tend to develop only when a leaf produces at or near the maximum number of lobes for that taxon. Influence of the same gene regulatory network may explain the similar appearance of stipules and basilaminar extensions in Platanaceae. [ABSTRACT FROM AUTHOR]

IODP Site U1509 (Expedition 371), New Caledonia Trough, provides a rare latest Cretaceous–Paleocene record from offshore northern Zealandia. We present new palynomorph and benthic foraminiferal assemblage data that show a transition from a latest Cretaceous vegetated sediment source region to a fully oceanic environment in the Paleocene. Latest Cretaceous (c. 68–66 Ma) non-calcareous claystone was deposited in upper bathyal paleodepths, with abundant plant microfossils that were likely transported in a northwest direction through the Aotea Basin region. A 2–3 Myr unconformity is identified at the Cretaceous/Paleogene boundary. Middle early–late Paleocene (c. 63.5–56 Ma) calcareous claystone shows evidence of deepening, deposited in middle bathyal or deeper paleodepths, and terrestrial input is minor. This latest Cretaceous to Paleocene deepening trend is consistent with inferred evolution of the Aotea and Northland basins further to the east, and other sparse northern Zealandia records, which show a common pattern of post-rift transgression consistent with long-term subsidence. Site U1509 allows for biogeographic extension and modification of the New Zealand Paleocene dinoflagellate zonation, description of a new dinoflagellate and pollen species, better inference of regional paleogeography, and may provide insights into the onset of western Pacific subduction initiation. [ABSTRACT FROM AUTHOR]

There are few studies focused on spore and pollen clumps in paleopalynological samples, and these are only reports from the Northern Hemisphere. These aggregates may be of animal or floral origin. The goal of this contribution is to provide the first detailed study of spore and pollen clumps from the Southern Hemisphere, and to discuss their possible origin, botanical affinities, and pollination modes, based on their morphological characteristics, preservation and comparison with putative living representatives. Three fern spore clumps and 18 angiosperm pollen clumps were recognized in Maastrichtian-Danian La Colonia Formation sediments that outcrop at Chubut Province, Patagonia, Argentina. Most clumps are monospecific and composed of undamaged elements but some of them have two types of spore/pollen with corroded and/or fragmented exines. These findings represent the most diverse and abundant record of fern and angiosperm clumps from the Late Cretaceous and early Paleocene so far known from South America and the Southern Hemisphere. These results are indicative of the need for comprehensive large-scale studies on pollination of modern taxa and careful processing of palynological samples to lessen the already large bias in paleopalynological interpretations. The paucity of information on clumps in the fossil record has impaired our comprehension of dispersion/pollination in deep time. [ABSTRACT FROM AUTHOR]

Summary: The storied structure in wood anatomy is considered derived and highly specialized, and is present in some related families. The first records of this character are from the Cretaceous, but it is scarce in the Cretaceous and Paleocene, and apparently absent until the Eocene in the temperate Southern Hemisphere. Using standard methodology, we describe the anatomy of a fossil wood from the early Paleocene of Patagonia (Salamanca Formation). The fossil wood, assigned to a new genus and species Elizabethiaxylon patagonicum related to the Malvaceae, is characterized by its diffuse-porous wood, solitary vessels, simple perforation plates, apotracheal banded axial parenchyma, and mostly biseriate storied rays. The storied structure in this wood is one of the oldest records from Gondwana. [ABSTRACT FROM AUTHOR]

We studied at least part of Kuhnʼs original material of lizards from the Paleocene (~ MP 5) of the Walbeck locality in Germany. The collection was considered to be lost but is consistently discussed in the literature due to its importance. We restudied the type material of aff. Parasauromalus paleocenicus and aff. Glyptosaurus walbeckensis described by Kuhn in 1940. The former was originally allocated to Iguania, the latter to Anguimorpha, though later on these identifications were questioned by several authors. We show such a classification of both cannot be upheld. P. paleocenicus resembles the morphology of lacertids showing their presence in Europe already around MP 5. We consider the name P. paleocenicus as a nomen dubium. The material of aff. G. walbeckensis was later suggested to belong to Lacertidae and also considered as a potential amphisbaenian. Although it differs from modern amphisbaenians, it shares features with one supposed polyodontobaenid – Camptognathosaurus parisiensis. The Walbeck form is identical to this species. Since the Walbeck taxon was described in 1940, the principle of priority makes Camptognathosaurus parisiensis a junior synonym of the species erected by Kuhn. We propose a new combined name for this form, Camptognathosaurus walbeckensis comb. nov. The specimen figured by Kuhn is currently lost, thus we designate a neotype from Walbeck. However, this taxon differs significantly from Polyodontobaena and new data doubt the attribution of Camptognathosaurus to Amphisbaenia. This taxon is tentatively assigned here to Lacertidae, as further confirmed by phylogenetic analyses. Material of Scincoidea is also described. [ABSTRACT FROM AUTHOR]

This paper discusses the current state of knowledge of Paleogene stratigraphy of Kaliningrad Oblast, the age of local stratigraphic units and the duration of possible gaps in deposition and proposes trends for future regional Paleogene stratigraphy. The Paleocene Chistoozerskaya and Lubava formations in total probably correspond to the Danian–Selandian on the basis of foraminiferal zonation, but their relationship is not entirely clear. The Zaostrovskaya Formation probably corresponds to the upper Thanetian, but its fossil record is very poor. The Sambian Formation corresponds to the Ypresian according to foraminifera, but its precise stratigraphical interval remains insufficiently substantiated. The Alka Formation is presumably late Lutetian–early Bartonian. The age of the Prussian Formation is emended based on dinocysts as early–late Priabonian, while the Palvé Formation is now dated as the latest Priabonian. The Kurshskaya Formation corresponds to the uppermost Eocene–Oligocene–?lower Miocene. The problem of the recognition and duration of hiatuses between formations remains relevant. [ABSTRACT FROM AUTHOR]

Premise of research. Burseraceae is represented by a large number of fossils from the Cenozoic sedimentary succession and has a disjunct distribution today in the southern part of North America, Africa, Europe, and Asia. However, the family has a poor fossil record in deep time, making hypotheses concerning its origin and dispersal difficult to evaluate. Here, we report the occurrence of reproductive organs (fruit and flower) attributable to modern Burseraceae from the probable latest Maastrichtian (Late Cretaceous) to earliest Danian (early Paleocene) sediments of the Mandla Lobe of the Deccan Intertrappean beds of Madhya Pradesh, Central India. Methodology. X-ray micro–computed tomography was used to observe three-dimensional structures. The reconstruction software takes the 2D projection images acquired by the X-ray detector and generates a 3D image represented by gray values distributed in a volumetric space. The present fruit and flower fossils are identified through morphological comparison with another fossil and extant Burseraceae. Pivotal results. The fossil pyrene corresponds in morphology with that of Burseraceae (particularly some genera of Protieae and Bursereae), while the fossil flower, with the pentamerous organization of sepals and petals and ovary and 10 stamens arranged in two whorls around a prominent intrastaminal nectary disk, is particularly similar to Protium. The fossil fruit is here described as a new species of the fossil genus Bursericarpum Reid et Chandler as Bursericarpum indicum Kumar, Manchester et Khan sp. nov. and the fossil flower as Debursera indica Kumar, Manchester, Judd et Khan sp. nov. Conclusions. The present discoveries represent the oldest reliable fossil records of Burseraceae supporting its Gondwanan origin. These fossils also provide important information for tracing possible migration pathways of Burseraceae from India to Europe after the docking of the Indian subcontinent with Eurasia during the early Eocene (55 ± 10 Ma), supporting an "out-of-India" dispersal hypothesis. Based on earlier and present records we suggest that the family might have reached Europe from India and then colonized North America (during the Eocene) in areas where it is now regionally extinct. [ABSTRACT FROM AUTHOR]

We here report on the first lizard fossils from the Deccan intertrappean strata (latest Cretaceous/Palaeocene) exposed at Kesavi, District Dhar in the Malwa Plateau of lower Narmada Valley, central India. The material is only fragmentary, but tentatively three tooth morphotypes of non-acrodontan lizards can be identified. Besides these, two oblong osteoderms, resembling paramacellodid osteoderms, are described as Squamata indet. The 4th isolated tooth is questionably referred to Squamata. Although the intertrappean deposits of the Deccan volcanic province have been explored for over three decades, lizards are scarce and many aspects remain unclear. However, the tentative absence of agamids in Kesavi and other localities yielding pre-Eocene deposits (e.g., Naskal and Kisalpuri) appears to be interesting, because a high diversity of agamids has been reported from early Eocene localities of India. There is, in contrast, a total absence of non-acrodontan lizards. The contrast between pre-Eocene and Eocene localities seems to be puzzling in the context of India's supposed physical isolation from Asia during this time. Only future researches can shed light on that. Although the material described here brings only limited new insight, it supports that non-acrodontan lizards were present in India during the latest Cretaceous/earliest Palaeocene. [ABSTRACT FROM AUTHOR]

A new concept of the formation of glauconite from the Paleocene deposits of the sedimentary cover of the southwestern part of the Near-Volga monocline is presented. It was formed as a result of halmyrolysis of ash material in lake conditions. Glauconitization is confined to the Paleogene thermal maximum and the periphery of active volcanism along the entire mobile belt. In this connection, it is widely distributed in the Paleogene deposits. Structural and crystal-chemical features of glauconite, stages of crystallization of its aggregates, paragenetic associations with smectite, kaolinite, and zeolite, as well as the presence of ore minerals indicate its authigenic origin. The authigenic origin is confined to the formations of temporary, episodically submerging shallow marine basins. An applied scheme of discrimination of glauconites by their crystallochemical features has been developed. The discrimination diagram can be used to determine the specific conditions of the sedimentation basin, which are expressed in the progradation cycles of glauconite accumulation. The cycles were controlled by induced depressions within the Russian plate. [ABSTRACT FROM AUTHOR]

The Dinaric–Hellenic mountain belt bends where two fault systems transect the orogen: (1) the dextral Shkoder-Peja Transfer Zone (SPTZ), active sometime between the Late Cretaceous and middle Eocene; (2) the Shkoder-Peja Normal Fault (SPNF), which accommodated NW–SE directed orogen-parallel extension. The SPTZ dextrally offsets the Dinaric–Hellenic nappes by ~ 75 km, a displacement attributed to reactivation of an Early Mesozoic rift transfer zone in the Adriatic margin during Paleogene subduction of the Pindos Ocean. This subduction involved an initial counter-clockwise rotation of the Hellenides with respect to the Dinarides around a pole at the NW end of the Budva–Krasta–Cukali–Pindos Basin. The SPNF overprints the SPTZ and is a composite structure comprising five fault segments: four of them (Cukali–Tropoja, Decani, Rožaje, Istog) were active under ductile-to-brittle conditions. They downthrow the West Vardar Ophiolite in the hanging wall. The Cukali–Tropoja and Decani segments exhume domes with anchizonal-to-greenschist-facies metamorphism in their footwalls. These structures formed during a first-phase of extension and clockwise rotation, whose Paleocene age is constrained by cross-cutting relationships. A second extensional phase was accommodated mainly by the fifth (Dukagjini) segment of the SPNF, a subsurface normal fault bordering syn-rift, mid-late Miocene clastic and lacustrine sediments in the Dukagjini Basin (DB) that are sealed by Plio-Pleistocene strata. This later phase involved subsidence of Neogene basins at the Dinaric–Hellenic junction coupled with accelerated clockwise oroclinal bending. The driving force for clockwise rotation is thought to be bending and rollback of the untorn part of the Adriatic slab beneath the Hellenides. [ABSTRACT FROM AUTHOR]

Materials on the Paleogene biostratigraphy of the Lomonosov Ridge in the Arctic Ocean are presented, including new data on foraminifera from the M0004A borehole section (interval 390–391 m) in its polar part. The studied benthic foraminifers are united into the complex with Reticulophragmium coksuvorovae of the upper part of the Selandian Stage to the lower part of the Thanetian Stage, which includes some characteristic species of the North Atlantic and West Siberian basins. Also, information about the connection of these Arctic seas in the Paleocene time is presented. [ABSTRACT FROM AUTHOR]

On the Fueguian Atlantic coast and in the southern Andean fold and thrust belt, the Paleogene is partially represented by numerous turbiditic deposits. Between Río Bueno and Cabo Leticia in Península Mitre, the Paleocene-Eocene is represented by the Río Claro Group (Cabo Leticia, La Barca, Punta Noguera, and Cerro Ruperto formations) and the Río Bueno Formation; which together constitute a ~1050-m-thick succession. Calcareous nannofossil assemblages recovered from 95 samples were analyzed to determine their relative ages. The Cabo Leticia Formation was barren in nannofossils. The upper LB2 Member of La Barca Formation contained Rhomboaster cuspis and Fasciculithus richardii, frequent in the Paleocene/Eocene boundary (subzones NP9a/NP9b). Nannofossils recovered from Punta Noguera Formation indicate an early Eocene age (biozones NP9b-NP10), with Fasciculithus tympaniformis, Rhomboaster cuspis, and abundant Toweius spp. The Cerro Ruperto Formation provided one productive sample, with specimens of Reticulofenestra spp. and Toweius spp., which indicates an early Eocene age for the formation (biozones NP12-NP13). The Río Bueno Formation yields calcareous nannofossil assemblages characterized by Chiasmolithus eograndis, Chiasmolithus expansus, Toweius spp., and Reticulofenestra spp., which indicate a late early Eocene to middle Eocene age (biozones NP14-NP15). Although the calcareous nannofossil record is discontinuous due to preservational biases in the Río Claro Group, our data allow a better age constraint for the investigated formations and to correlate these units with other surface and subsurface units in the Austral Basin. [ABSTRACT FROM AUTHOR]

Paleocene–Eocene phosphorite beds from three major outcrops of the Maknassy-Mezzouna basin (Central-Tunisia) were geochemically investigated to improve local stratigraphy, determine their ages, and attempt to constrain ocean circulation. 87Sr/86Sr ratios of phosphate materials from the Chouabine Formation display a large variation even within the same deposit (87Sr/86Sr between 0.707782 and 0.707830). Based on the Sr isotope data obtained in teeth enamels, further supported by the results in coprolites, Sr isotope ages of 57 ± 1.1 Ma were attributed to Layers I and II, and 62 ± 1.4 Ma to Layer III. These results assign a Paleocene Epoch for the principal phosphate series. The 87Sr/86Sr value of teeth enamel sample from the upper phosphate bed suggests an age of 46 ± 1 Ma for this sequence. Furthermore, the Nd isotopic compositions of these samples were used as a tracer for seawater circulation. The 143Nd/144Nd values of the analyzed samples vary between 0.512174 and 0.512202, corresponding to εNdCHUR between – 8.1 and – 8.4. These values are similar to those reported from Gafsa-Metlaoui basin, southern Tunisia and other North African phosphorite deposits. These data are reminiscent of those of the Atlantic Ocean, which is new argument of the overturning of Tethys Circumglobal Current during the Paleogene period. The obtained results suggest water exchange between Tethys and Atlantic Ocean during this period. [ABSTRACT FROM AUTHOR]

Recognition of the existence of an Andeantype continental margin in southern Tibet prior to its collision with India has provided crucial constraints on the formation of the Tibetan Plateau and South Asian climate evolution. Here, we focused on well-dated Late Cretaceous successions in the Linzhou Basin and determined the elevation changes from sea level to high mountains in the Gangdese magmatic arc in southern Tibet. Our results show that the Linzhou Basin was still submerged in the Tethyan Sea at around 92 Ma when it accumulated Cenomanian (ca. 105-92 Ma) shallow-marine orbitolinid-bearing limestones (Takena Formation); these limestones are unconformably overlain by Campanian (83-78 Ma) fluvial-lacustrine deposits (Shexing Formation) after an ~9 m.y. depositional hiatus. A prominent unconformity between the tightly folded Shexing Formation and gently tilted overlying Paleocene--Eocene Linzizong successions represents the formation of the regional Lhasaplano ca. 70 Ma, which may have been linked to the rise of an Andean-type Gangdese mountain range along the southern margin of the Lhasa terrane. Using carbonate oxygen isotope and clumped isotope thermometry data from the Shexing Formation paleosols, we quantitatively documented the rise of the Andeantype Gangdese Mountains with a peneplain surface at an elevation of 2.7 + 0.5/-0.9 km above sea level prior to onset of the India-Asia collision ca. 65-63 Ma. This scenario of surface gain may have been an isostatic compensation response to the crust thickening to over 50 km during the Late Cretaceous. Subsequently, the surface isostatically rose to its near-present elevation of ~4.6 km due to the removal of overthickened mantle lithosphere and revival of intense Gangdese magmatism by 56 Ma. [ABSTRACT FROM AUTHOR]

After about a century, the skull of the holotype and sole specimen of Arambourgia gaudryi from the Eocene of Quercy (France) is here redescribed. Its interest resides in its extreme morphology, such as its very small skull and altirostral condition, that raises questions about its maturity, taxonomic validity, and paleoecology. In order to reveal previously hidden anatomical details, we used computed microtomography (µCT-scan) data to digitally extract the bones of the cranium and mandible, as well as the endocranial structures. Here, we update the osteological description of A. gaudryi, outlining new characters and re-evaluating anatomical details. Comparing it with ontogenetic series of extant crocodylians, we confirm that the specimen is a mature individual of a dwarf species with pedomorphic features. Our phylogenetic analyses indicate that A. gaudryi is an alligatorine. Comparisons with other small alligatoroid taxa show that it is indeed a valid species. The depositional environment of the finding site suggests a continental context with a karstic network. Taking this into account, together with its peculiar anatomy, we propose that A. gaudryi was a semi-terrestrial animal occupying an ecological niche similar to that of extant juvenile or dwarf crocodylians. Finally, we discuss the current knowledge of the paleobiogeography of Alligatoroidea in Europe during the Paleogene. Arambourgia gaudryi was likely part of a group of small early globidontans that arrived in the Old World, probably from North America, around the end of the Paleocene and that dispersed to at least four localities of the ancient European archipelago. [ABSTRACT FROM AUTHOR]

The Paleocene deposits from the Cerrejón Formation (Colombia, South America) have provided abundant snake remains. All this material has been assigned to the giant snake Titanoboa cerrejonensis. Here, we describe six vertebrae from among the referred materials that do not correspond to Titanoboa. All six vertebrae are morphologically identical, of a similar size, and display a distinct morphology that differs from Titanoboa. This new and unnamed taxon is a large snake estimated to be around eight meters in total length. Anatomical comparisons show that these vertebrae possess certain features typical of the Palaeophiidae, a group of extinct snakes of uncertain relationships known almost exclusively from vertebrae. Among the Palaeophiidae, the new taxon resembles the more generalized forms assigned to the genus Palaeophis, but lacks the extreme aquatic adaptations that define forms assigned to the genus Pterosphenus. We regard the new taxon as an undetermined palaeophiine but do not assign it to Palaeophis as a number of features differ from that described genus. This new record expands the known diversity of aquatic snakes from the Paleogene of South America and provides a substantial new record for the paleogeographic distribution of the Palaeophiidae. The oldest records are from the Cretaceous of Africa, while Paleocene records are poorly known and restricted to Africa, North America, and Europe. The majority of vertebral forms assigned to Palaeophiidae are from the Eocene, principally from the Tethys region, with only one previous South American record coming from the Eocene of Ecuador. [ABSTRACT FROM AUTHOR]

Resolving the Mesozoic and Cenozoic palaeogeography and geomorphologic development of outcropping Variscan basement is a pre-condition for the understanding of central European geodynamics. For our study, we have applied apatite fission-track (AFT) and apatite (U–Th)/He (AHe) thermochronology to surface rocks of the southeastern Bohemian Massif. 46 samples were examined by the AFT method. Additional AHe dating was applied to 8 of them. The AFT ages range from 251 ± 46 to 60.2 ± 4.8 Ma. AHe ages range from 25 to 525 Ma with rather high intra-sample scatter. On a regional scale, the AFT ages generally decrease from mainly late Variscan in the NE to Late Cretaceous and Paleocene in the SW. This regional age asymmetry relative to the NW–SE trending watershed of the Weinsberg Forest is neither compatible with regional uplift of a single block nor with large-scale lithospheric updoming. The lack of age breaks along late Variscan faults demonstrates that strong vertical offset cannot have occurred in Cretaceous and Cenozoic times. Inverse modeling of thermochronological data indicates regional Early Cretaceous cooling and subsequent reheating during the Late Cretaceous. Rocks of the present-day surface were heated up to a temperature of ca. 80 °C without full reset of the AFT system. This thermal history is compatible with the existence of a large mainland in Early Cretaceous times and a subsequent sedimentary reburial until the Campanian on the order of up to 1 km overburden. Parts of the exhumed weathering basal relief to the N and NE of the Weinsberg Forest are inherited as 'sealed relief' from Middle Cretaceous time. The observed regional asymmetry of AFT data is best explained by the development of a continental escarpment adjacent to the North Penninic Ocean in latest Cretaceous to Paleogene times. A final episode of accelerated cooling after ca. 20 Ma, as indicated by thermochronological modeling, is tentatively ascribed to either collisional coupling of the Alpine-Carpathian nappe pile with its northern foreland or to East-Alpine slab detachment. [ABSTRACT FROM AUTHOR]

Bauhinia s.s. is a large genus in the family Fabaceae, but its evolutionary and biogeographical history is still unclear due to the scarcity of fossil records compared to the highly diverse modern species in pantropic regions. Here, we report the earliest fossil record of Bauhinia s.s., namely Bauhinia tibetensis Y. Gao et T. Su sp. nov., based on leaves from the latest Paleocene of the southern Tibetan region. Combined with palaeoecological niche simulations and ancestral state reconstruction, the new fossils suggested a Paleocene origin of Bauhinia s.s. in the Afrotropical realm that subsequently dispersed to the Neotropical and Indomalayan realms. Bauhinia tibetensis belongs to the Asian clade of Bauhinia s.s. that reached the southern Tibetan region from the Afrotropical realm via the Kohistan-Ladakh Island Arc in the early Paleocene. This clade spread to south-eastern China during the Oligocene and entered northern India during the Neogene or earlier. The discovery of the oldest Bauhinia s.s. from what is now the southern Tibetan Plateau updates our understanding of the biogeographical history of this genus and demonstrates that the Kohistan-Ladakh Island Arc is an ancient corridor for floristic interchange between Africa and India. [ABSTRACT FROM AUTHOR]

The record of articulated marine fish fossils during the latest Cretaceous and earliest Cenozoic is sparse. The oldest-known definitive squirrelfishes and soldierfishes, like the first examples of many extant reef-dwelling clades, are known from early Eocene deposits of Europe. Here, we describe a new genus and species of holocentroid (Teleostei: Beryciformes: Holocentroidea) based on material from three individuals from early Paleocene (Danian) deposits of New Jersey, USA using micro-computed tomography. The specimens comprise a three-dimensionally preserved skull and partial postcranium, plus two isolated neurocrania. The new taxon, †Iridopristis parrisi, possesses a unique combination of characters, including a heterosulcoid otolith morphology and an edentulous premaxillary tooth-gap, while lacking a newly proposed character for the remainder of Cenozoic holocentroids: a lamina on the lateral surface of the anguloarticular, anterior to the jaw joint. Bayesian phylogenetic analysis of morphological, stratigraphical and molecular data under the fossilized birth-death process finds that the new taxon branches from the holocentrid stem, where it is joined by two of the three squirrelfish genera from the early Eocene (Ypresian) of Bolca, Italy. We estimate a Danian divergence between Myripristinae and Holocentrinae, the two reciprocally monophyletic subfamilies of Holocentridae. Our analysis suggests that several holocentroid lineages crossed the Cretaceous–Palaeogene boundary. [ABSTRACT FROM AUTHOR]

Premise of research. Fossil infructescences of the late Paleocene Cyclocarya brownii (Juglandaceae) and winged fruits with newly recognized anatomical structure are described from Almont and Beicegel Creek, North Dakota. These fossils demonstrate that C. brownii has many similarities to extant Cyclocarya paliurus but differs in several morphological and anatomical features. Comparisons of the fossil record of Cyclocarya document the transition of characters in the evolution of the oldest extant genus of Juglandaceae. Methodology. Fossils were digitally imaged with reflected light microscopy (LM). Some specimens were embedded in Ward's Bio-Plastic synthetic resin, sectioned into wafers, mounted on microscope slides, and imaged with reflected LM. Permineralized specimens from the Beicegel Creek site were prepared with the cellulose acetate peel technique, mounted on microscope slides, and imaged with transmitted LM. Extant fruits of C. paliurus were photographed from freshly collected material. Pivotal results. Infructescences of C. brownii differ from those of modern C. paliurus in several ways. The fossils are racemes with crowded fruits borne on elongate pedicels. In contrast, extant C. paliurus has spikes bearing fewer fruits per infructescence (10 vs. 23). Fruit wall anatomy is similar to that of extant and other known fossil species of Cyclocarya but is more complex and has a distinctive idioblast layer not present in extant Cyclocarya. Fossil pollen is triporate in contrast to the tetraporate pollen in extant Cyclocarya. As in extant Cyclocarya , pollen is borne both in staminate catkins and occasionally in stamens attached to fruits. We document a fossil fruit with stamens bearing pollen. This feature, sporadic in extant Cyclocarya , was also present in the Paleogene. Conclusions. Late Paleocene C. brownii differs in comparison with extant C. paliurus and related fossil forms in infructescence architecture, fruit size and symmetry, fruit wall anatomy, and pollen aperture number. Trends in character evolution from the Paleogene to the present day include (1) changes from helically arranged, densely distributed pedicellate fruits to sessile fruits borne singly along a slender axis, (2) transition from pyramidal to round nutlets with (3) equatorial versus basal wing attachment, (4) a change in pollen aperture number from three to four, and (5) simplification of the fruit wall in extant Cyclocarya. [ABSTRACT FROM AUTHOR]

New decapod crustaceans from ?Selandian to Thanetian strata of the Kambühel Formation, Austria, include the paguroid Squamipelta insecta new genus and species, and two new galatheoid species, Protomunida kambuehelensis and Annieporcellana paleocenica. The range of Annieporcellana is extended across the Cretaceous-Paleogene boundary. Selandian and Thanetian decapod occurrences are rare; thus, this report adds to the known decapod diversity in the recovery after the end-Cretaceous events. [ABSTRACT FROM AUTHOR]

The Paleocene lavas from Dianzhong Formation (E1d) in Linzhou basin of southern Lhasa terrane are a key target for paleomagnetic investigations into the timing and paleolatitude of the initial India‐Asia collision. Controversy exists, however, on whether these rocks preserve a primary remanent magnetization. Here we reanalyze previously published thermal demagnetization data and report detailed rock magnetic results and petrographic observations of these rocks. We find that the original magnetic carrier, a magmatic multidomain Ti‐poor titanomagnetite, underwent significant grain size reduction and was variably reacted to single‐domain maghemite and nano‐hematite. Such strong alteration may have resulted from successive hydrothermal events: a first event related to the ∼52 Ma dike intrusions into the E1d that accompanied a massive ignimbrite eruption deposited above the E1d producing heating up to 300°C; and a secondary event related to the 42–27 Ma southward overthrusting of the basin, heating the E1d up to 130–145°C. Unblocking/inversion temperature spectra of the authigenic maghemite and nano‐hematite overlap with those of the titanomagnetite, implying that the primary remanence of the E1d lavas has been contaminated or replaced by thermoviscous and chemical remanent magnetizations. Thus the isolated characteristic remanent magnetization from these rocks, whether slightly or completely altered, cannot be considered primary and should not be used for paleolatitudinal determination. Our study confirms that hydrothermal alteration can seriously jeopardize the remanence carried by titanomagnetite and thus should be tested for paleomagnetic investigations of rock units from tectonically active areas. Plain Language Summary: The study of past magnetic directions preserved in rocks (paleomagnetism) enables to constrain the paleolatitudes of the southern Asian margin where the rocks formed and is thus key to understand the timing and latitude of the initial India‐Asia collision. However, paleomagnetic studies of Paleogene rocks of southern Tibet have returned a surprisingly wide spread of paleolatitude data. One possible explanation is unrecognized remagnetization, a process that may relate to subsequent hydrothermal alteration. Such hydrothermal alteration has recently been documented for Paleocene lavas in the Linzhou basin, paleomagnetic signals from which have previously been interpreted either as primary or secondary (re)magnetizations. Here we analyze previously published and newly acquired paleomagnetic, rock magnetic, and petrographic data of these rocks to identify whether hydrothermal alteration caused remagnetization, and how such events may be fingerprinted. We find that the original magnetic carrier is Ti‐poor titanomagnetite. Hydrothermal alteration, however, led to significant changes in these titanomagnetite crystals and mineral transformations to maghemite and hematite, associated with acquisition of secondary magnetizations with the magnetic behaviors similar to the primary magnetization. This can be misleading for the interpretation of the magnetic directions obtained from these rocks and imply that they are not suitable for paleolatitudinal determination. Key Points: Hydrothermal alteration intensively destroyed the original multi‐domain titanomagnetite and recorded magnetization in the Dianzhong lavasTitanomagnetite experienced serious grain size reduction and has variably converted to authigenic single‐domain maghemite and nano‐hematitePrimary remanent magnetization has been replaced or contaminated, these lavas should not be used to determine Lhasa terrane paleolatitude [ABSTRACT FROM AUTHOR]

Premise of research. The Anacardiaceae family is distributed throughout the vegetated continents. The fossil record indicates extensive diversification of the family during the Paleogene and, in particular, during the Eocene. Despite the abundant fossil record of this period, there are only a few reliable anacardiaceous fossils in the Paris Basin. Here, we aim to document newly recognized fossils of Anacardiaceae from the Paris Basin, understand their paleoecology, and discuss their biogeographic history. Methodology. Thirty-three lignite fruits were examined from two sites, one before and one after the Paleocene-Eocene boundary (i.e., Petit Pâtis [Rivecourt] and Le Quesnoy [Houdancourt]). The specimens were photographed, and anatomy was studied using computed tomography and histological sections. Comparative analyses were undertaken using available descriptions of fossil and modern fruits of Anacardiaceae. Pivotal results. A new species, Cyrtocarpa biapertura sp. nov., is described on the basis of a unilocular fruit with two prominent apertures present on the ventral side of the endocarp, protruding into two lacunae surrounding the locule. Taphonomic analysis indicates that this plant grew close to riverbanks. Furthermore, a new record of " Lannea " europaea (Reid and Chandler) Chandler is reported for the Eocene site. Conclusions. The occurrence of Cyrtocarpa in both the Paleocene and the Eocene floras in the Paris Basin suggests similar vegetation during both time intervals. It is likely that both floras grew under similar subtropical climates. Moreover, it appears that the early Eocene shows an enrichment of the paleodiversity of Anacardiaceae and other plant families in the Paris Basin. The presence of Cyrtocarpa documents a rarely reported disjunction between the Paleogene of Europe and the recent tropical flora of South America. [ABSTRACT FROM AUTHOR]

Palaeogene sediments of the Hampshire Basin were a sensitive recorder of fluctuations in climate and eustatic sea level as Earth's climate transitioned from the global early Eocene 'hothouse' to the early Oligocene 'coolhouse', accompanied by the first permanent continent‐scale glaciation of Antarctica at the Eocene/Oligocene boundary. A study of the Palaeogene sediments of the Hampshire Basin is not only interesting from a palaeoclimate perspective, but the marine middle Eocene formations are renowned for containing some of the most abundant and diverse Palaeogene fossil assemblages in the world. In this article, I take you on a tour of the geological evolution of the Hampshire Basin from the end of the Paleocene through to the Pleistocene. I highlight some of the best exposures where sediments can be studied at outcrop and representative fossil assemblages can be collected, along with the economic and archaeological significance of these Palaeogene sediments. [ABSTRACT FROM AUTHOR]

Cimolodontan multituberculates are common in the Late Cretaceous and Paleocene of central Asia; they are rarely known from regions south of the Mongolian plateau. Here, we report a new genus and species of multituberculate, Erythrobaatar ganensis gen. et sp. nov., from the Late Cretaceous of Ganzhou, Jiangxi Province, central China, representing the southernmost record of Cretaceous multituberculates in Eurasia. The new species is based on two well-preserved specimens that include cranial and postcranial materials. With a cranium and dentary length of 83 and 66 mm, respectively; it is one of the largest known Cretaceous multituberculates from Eurasia. The present work focuses on the description of the craniodental morphology of the new species, in comparison with that of other Late Cretaceous and Paleogene cimolodontans in order to establish the new taxon. Erythrobaatar ganensis is most closely related to Yubaatar zhongyuanensis and Yubaatar qianzhouensis in the shape and size of the skull and teeth. Phylogenetic analyses place the new species within Taeniolabidoidea, which consists mostly of Late Cretaceous and Paleogene cimolodontans from Asia and North America. The new material also sheds light on tooth replacement, reduction, homologies, and occlusion of multituberculates with a focus on cimolodontans. [ABSTRACT FROM AUTHOR]

Pollen and spores were recovered from the Paleocene Fort Union Formation and Paleocene–Eocene Willwood Formation of the Bighorn Basin (BHB), northwestern Wyoming, USA. In many local stratigraphic sections in the BHB, the base of the Eocene has been identified by the characteristic negative carbon isotope excursion (CIE) that marks the beginning of the Paleocene–Eocene Thermal Maximum (PETM). The palynotaxa from outcrop samples were examined using light microscopy (LM) and scanning electron microscopy (SEM). Seven new species are formally described (Tricolpites vegrandis, Rousea spatium, Striatricolporites astutus, Striatopollis calidarius, Friedrichipollis geminus, Retistephanocolporites modicrassus and Retistephanocolporites pergrandis). The temporal and geographic distributions of many of these palynotaxa suggest that hotter and more seasonally dry climates facilitated their northward range shifts during the PETM from the tropics or subtropics of the USA. For the temperate palynotaxa, the hotter and seasonally dry conditions resulted in local extirpation. A re-evaluation of the palynostratigraphic schemes established for the Paleocene–Eocene boundary confirms that the first appearance of Platycarya platycaryoides denotes the Paleocene–Eocene boundary in the Rocky Mountains region. A new Striatopollis calidarius Subzone, associated with early Wasatchian (Wa) Wa-0 and Wa-R faunas, is also recognized for CIE body localities in the BHB. [ABSTRACT FROM AUTHOR]

We describe nearly complete skeletons of basal Anseriformes from the Latest Paleocene to the early Eocene of North America and Europe. Collectively, these birds appear to be representative of anseriforms near the divergence of Anhimae and Anseres, but their exact positions relative to these clades remains uncertain. A new family, Anachronornithidae nov. fam., is erected on the basis of one of these, Anachronornis anhimops nov. gen., nov. gen. et sp., to which the others cannot be confidently assigned. The new fossils augment a growing collection of early Pan-Anseriformes, which in their diversity do not paint an unambiguous picture of phylogeny or character state evolution on the path to or within crown-Anseriformes. Anachronornis nov. gen. is similar in some aspects of both cranial and postcranial anatomy to other well-represented early Paleogene Anseriformes and members of Anseres, such as Presbyornis Wetmore, 1926. However, it exhibits a more landfowl-like bill, like that of Anhimae and unlike the spatulate bill of Anseres. Additional specimens of similar basal Anseriformes of uncertain affinities from the early Eocene of North America and Europe further complicate interpretation of character state polarity due to the mosaicism of primitive and derived characters they exhibit. [ABSTRACT FROM AUTHOR]

A new fossil occurrence of freshwater sponges (Porifera: Demospongiae) is reported from the Deccan volcanic associated Naskal intertrappean locality, deposited in an interval of <100 kyr across the Cretaceous/Paleogene (K/Pg) boundary. This is the oldest record of siliceous fossil birotule spicules (gemmuloscleres) belonging to asexual resting stages typical of the order Spongillida. The analysis supports the ascription of these fossils to the family Palaeospongillidae. The diagnosis and description of Longibirotula Pronzato and Manconi gen. nov. and its type species Longibirotula antiqua Manconi and Samant sp. nov. from the Naskal intertrappean is based on skeletal and gemmular spicular morphotraits. The findings have provided evidence of the presence of diversified groups of freshwater sponges during the Late Cretaceous on the Indian subcontinent and Gondwanaland. From the biogeographic context, the findings track the evolutionary trends of the oldest continental sponges in the Asian and Australasian/Insular Pacific regions. [ABSTRACT FROM AUTHOR]

Sea turtles (Testudines: Chelonioidea) represent a monophyletic group with a global distribution in the fossil record, although most of our knowledge of their diversity and morphology comes from the Atlantic Ocean and Tethyan Sea (Evers and Benson 2019; Gentry et al. 2019). By contrast, the record of their evolution in the vast Pacific Basin prior to the Neogene is paltry. Late Cretaceous chelonioids are known from northern California (Parham et al. 2003), British Columbia (Nicholls 1992; Nicholls and Meckert 2002), Japan (Hirayama and Chitoku 1996), Australia (Kear et al. 2003; Kear 2006; Kear and Lee 2006), and New Zealand (Wiffen 1981), indicating they occupied at least portions of both the North and South Pacific Oceans at this time. Pacific Paleocene specimens are entirely unknown, and to date the Eocene record has been limited to chelonioids and dermochelyids (Köhler 1995a, b; Grant-Mackie et al. 2011) from New Zealand, and an informal report of a sea turtle from Oregon (Orr and Orr 2009). Here, we describe a new sea turtle fossil from the middle Eocene of California which demonstrates that members of this clade were indeed present in the North Pacific at this time. [ABSTRACT FROM AUTHOR]

In contrast to the rich collections of articulated spiny-rayed fishes from early Late Cretaceous and Eocene Lagerstätten, similar skeletal remains are sparse in Maastrichtian-Paleocene strata. Here we coin this poorly understood span "Patterson's Gap" and review known articulated skeletons from it, summarizing available information on their phylogenetic affinities, age, and environmental context. Roughly fifty percent of taxa in both the Maastrichtian and Paleocene come from Europe and North America, with percomorphs representing around 60% of the skeletal acanthomorph taxa in each interval. This is higher than the only pre-Maastrichtian assemblage with a reasonable sample of percomorphs, but lower than most Eocene and younger sites. Fossils from Patterson's Gap show a steady accumulation of the principal lineages of spiny-rayed fishes. Material from Paleocene or older strata provides evidence for most of the roughly 20 major acanthomorph divisions recovered by molecular studies. Many fossils from Patterson's Gap remain undescribed and unnamed, and almost none have been included within formal phylogenetic analyses. Revision of existing material, combined with additional fieldwork, should be a priority for future efforts seeking to clarify this murky but significant interval in the evolutionary history of a major vertebrate radiation. [ABSTRACT FROM AUTHOR]

The Maastrichtian-Thanetian Ncham Member of the Aurès Basin in northeastern Algeria contains variably lenticular microfossils, which are defined as Laffitteina genera. It is a large benthic foraminifera, studied from three stratigraphic sections. The systematic study of Laffitteina led identification of nine species, four of which, Laffitteina marsicana, L. aff. mengaudi, L. monodi and L. erki were recorded for the first time in the study area. Various species of Laffitteina were used as index fossils to recognize the K/Pg transition in the Aurès Basin. The last occurrence of L. oztuerki marks the end of the Maastrichtian, which was found in association with Omphalocyclus macroporus (Lamarck) in the upper Maastrichtian. The first occurrence of L. erki marks the Thanetian age, while the Danian and Selandian deposits are missing in the field after a general emersion in the early Paleocene, despite the absence of a visible non unconformity. It could therefore be proposed that the K/Pg transition may occurs in the last 10 m of the Ncham Member. Concerning the paleoenvironment, Laffitteina is considered as a resistant foraminifera and thrives in meso-to eutrophic environments (lagoon). Microfacies data from the Upper Cretaceous-Lower Paleogene limestones of the Aurès Basin suggests a carbonate platform with shallowing-up sequential organization, mainly controlled by fluctuations in sea-level. [ABSTRACT FROM AUTHOR]

A new fossil occurrence of freshwater sponges (Porifera: Demospongiae) is reported from the Deccan volcanic associated Naskal intertrappean locality, deposited in an interval of <100 kyr across the Cretaceous/Paleogene (K/Pg) boundary. This is the oldest record of siliceous fossil birotule spicules (gemmuloscleres) belonging to asexual resting stages typical of the order Spongillida. The analysis supports the ascription of these fossils to the family Palaeospongillidae. The diagnosis and description of Longibirotula Pronzato and Manconi gen. nov. and its type species Longibirotula antiqua Manconi and Samant sp. nov. from the Naskal intertrappean is based on skeletal and gemmular spicular morphotraits. The findings have provided evidence of the presence of diversified groups of freshwater sponges during the Late Cretaceous on the Indian subcontinent and Gondwanaland. From the biogeographic context, the findings track the evolutionary trends of the oldest continental sponges in the Asian and Australasian/Insular Pacific regions. [ABSTRACT FROM AUTHOR]

Sea turtles (Testudines: Chelonioidea) represent a monophyletic group with a global distribution in the fossil record, although most of our knowledge of their diversity and morphology comes from the Atlantic Ocean and Tethyan Sea (Evers and Benson 2019; Gentry et al. 2019). By contrast, the record of their evolution in the vast Pacific Basin prior to the Neogene is paltry. Late Cretaceous chelonioids are known from northern California (Parham et al. 2003), British Columbia (Nicholls 1992; Nicholls and Meckert 2002), Japan (Hirayama and Chitoku 1996), Australia (Kear et al. 2003; Kear 2006; Kear and Lee 2006), and New Zealand (Wiffen 1981), indicating they occupied at least portions of both the North and South Pacific Oceans at this time. Pacific Paleocene specimens are entirely unknown, and to date the Eocene record has been limited to chelonioids and dermochelyids (Köhler 1995a, b; Grant-Mackie et al. 2011) from New Zealand, and an informal report of a sea turtle from Oregon (Orr and Orr 2009). Here, we describe a new sea turtle fossil from the middle Eocene of California which demonstrates that members of this clade were indeed present in the North Pacific at this time. [ABSTRACT FROM AUTHOR]

The Cretaceous-Paleogene (K-Pg) contact interval is constrained by vertebrate fossil sites at seven sites in the Tornillo Group and lies within an 80–100-m stratigraphic section between the top of the Javelina Formation and the base of the “log jam sandstone” marker bed in the Black Peaks Formation. In western exposures of this interval, the highest occurrence of in situ dinosaur specimens and the lowest occurrence of Paleocene mammal specimens are separated by an unusual conglomerate bed. This thin conglomerate bed coincides with the contact between Cretaceous and Paleogene strata and contains reworked Cretaceous fossils. It is superficially similar to conglomerate beds elsewhere attributed to the effects of tsunamis generated by the Chicxulub impact; however, the maximum depositional age of ca. 63 Ma based on detrital zircons indicates that the conglomerate was deposited about three million years after the K-Pg boundary event. Paleocene mammalian fossils from immediately above the conglomerate bed represent a fauna that can be no older than the middle Torrejonian (To2 interval zone). The contact between Cretaceous and Paleocene strata is therefore disconformal and represents a hiatus of at least three million years. A condensed section occurs at the westernmost exposure of the K-Pg contact, where at least 80 m of strata are absent below the conglomerate bed; these strata are present in exposures farther east. This condensed section likely records an erosional event resulting from uplift and deformation of the nearby Terlingua monocline. Although the 80 m of strata below the conglomerate bed are poorly fossiliferous, several clearly in situ dinosaur specimens indicate that this entire interval is Late Cretaceous in age. There is no compelling evidence for preservation of the K-Pg boundary event horizon at any of the seven sites in the Tornillo Group, and so the hiatus represented at the Cretaceous/ Paleocene contact here likely also includes some part of latest Cretaceous time. Mammalian specimens from sites in the “log jam sandstone,” ~40 m above the middle Torrejonian sites, represent an early Tiffanian fauna (Ti1 interval zone). Latest Torrejonian (To3) sites have not been recognized, and therefore a second disconformity likely coincides with the base of the “log jam sandstone” marker horizon in the Black Peaks Formation. [ABSTRACT FROM AUTHOR]

The absolute age of the Brusilovka paleo-caldera volcanic rocks has been determined for the first time through the LA-ICP-MS U–Pb zircon isotopic dating. The obtained results (~61–56 Ma) are consistent with the Paleocene magmatic stage of the Sikhote-Alin, and the patterns of their structural localization and spatiotemporal distribution are consistent with left-lateral displacements along the Vostochny and Pribrezhny faults. The distribution characteristics of early Paleogene igneous rocks both in the frontal and rear continental-marginal parts of the southern Sikhote-Alin can serve as a basis for paleogeodynamic reconstructions and subsequent metallogenic analysis. [ABSTRACT FROM AUTHOR]

Numerous bottom current-controlled depositional and erosional features, which together form Contourite Depositional Systems (CDS), have been recognized in deep-water settings over the past decade. Most of these systems are described based on two-dimensional (2D) seismic data, whereas only a few CDS have been characterised from high-resolution 3D data. Here we document a newly identified CDS that formed during the Paleocene within the Morondava Basin, offshore west Madagascar, through analysis of a depth-migrated 3D seismic survey, enhanced by the implementation of seismic attributes. Three seismic units (SU) mark the main evolutionary stages of the CDS: (a) the onset (SU1), (b) drift growth (SU2), and (c) burial (SU3) stages. The growth stage documents lateral upslope migration of a mounded drift and its associated moat. The increasing, long-term influence of bottom currents along the foot of the slope occurred simultaneously with plate tectonic, climatic and oceanographic changes. Evidence amassed from the CDS highly erosive bounding discontinuities, internal discontinuities, and moat architecture all indicate the intermittent behaviour of the currents over shorter time frames during its formation. Drift deposits form under the influence of weaker currents, while discontinuities appear to record the most vigorous currents, producing the large-scale morphology of the system. [ABSTRACT FROM AUTHOR]

The Cretaceous/Paleogene (K/Pg) boundary fern-spore spike concept was first introduced by R.H. Tschudy 40 years ago and established the precept that ferns are so-called 'disaster taxa' that flourish after natural disasters because of the high colonization potential of their wind-blown spores. Among the least understood topics at the crux of contemporary K/Pg boundary fern-spike studies is whether Stenochlaena-like or other stem blechnaceous ferns were among those that flourished after the K/Pg event, contributing to the Laevigatosporites-dominated phase of the fern-spore spike. For instance, dispersed Stenochlaena-like spores – e.g. Polypodiisporites usmensis (van der Hammen) Khan & Martin – first appear in the upper Eocene stratigraphic record of South America, whereas megafossils of Stenochlaena-like ferns are first known from the upper Paleocene strata of this same region. Beyond this traditional focus on taphonomic and taxonomic uncertainty regarding the identity of dispersed spore producers, however, there is further discordance between contemporary paleobotanical and molecular phylogenetic perspectives on the timing of diversification of stem lineages of blechnaceous ferns in relation to the K/Pg boundary. This investigation reconciles these two perspectives by constraining molecular clock divergence time estimates using contemporary fossil data. If this reconciliation is correct, then paleopolyploidization (whole genome duplication or WGD) associated with the origin of the genus Stenochlaena J. Sm. predated evolution of polocytic stomata, a heavily ornamented exospore, and a hemi-epiphytic to epiphytic habit observed in the crown lineage. Critically, this novel perspective elicits a merger of the K/Pg boundary fern spike and K/Pg boundary-WGD concepts, suggesting a link between polyploidy and the 'disaster taxon' concept. [ABSTRACT FROM AUTHOR]

Volkheimerites labyrinthus gen. et sp. nov., an angiosperm pollen type, is fully described and illustrated. The specimens were retrieved from lower Paleocene strata of the Salamanca Formation (San Jorge Basin) located in Chubut Province, Patagonia, Argentina. Medium to large size (25.5–55 μm) and a characteristic striate sculpture with labyrinthine pattern are distinguishing features of the grains in this new genus. Volkheimerites labyrinthus shares the presence of a coarse tectum supported by columellae with the fossil genera Dichastopollenites, Trisectoris and Periretisyncolpites, and some Winteraceae and Annonaceae species, but differs in having striate-labyrinthine as opposed to reticulate ornamentation. Our material is also distinguishable by the occurrence of solitary grains (monads) instead of the more frequent presence of tetrads in the Winteraceae family or grains separated into two hemispheres as in Dichastopollenites. The large thin area of the exine appears to be morphologically analogous to the proximal thinning of the exine in the monads of tetrads as occurs in some Annonaceae species. The detailed description and illustration of this new angiosperm pollen genus expands our understanding of Paleocene plant diversity in Patagonia. [ABSTRACT FROM AUTHOR]

The earliest phases of mammalian recovery following the Cretaceous–Paleogene (K–Pg) mass extinction are incompletely known but crucial to understanding the development of modern terrestrial ecosystems. Here we report new mammalian faunal data from three vertebrate microfossil assemblages in the Hell Creek region of northeastern Montana, the deposition of which we constrain to within the first 28–80 ka of the Paleocene using new stratigraphic observations within a high-resolution chronostratigraphic framework. We quantified the taxonomic diversity among these three assemblages and five other assemblages from both the Hell Creek region and Denver Basin, together spanning the first ca. 300 ka post-K–Pg mass extinction. Our results allowed us to sub-divide the established 'disaster' and 'recovery' phases of recovery into the following sub-phases: (i) early disaster, characterized by the presence of 'dead clades walking,' high relative abundance of bloom taxa, and the appearance of post-mass-extinction immigrants, (ii) late disaster, characterized by a reduction in the number of 'dead clades walking,' continued high relative abundance of bloom taxa, and a more diverse assemblage of immigrants, (iii) early recovery, characterized by decreased relative abundance of bloom taxa, and continued immigration, and (iv) late recovery, characterized by the onset of in situ diversification. We note important differences in the pattern and timing of mammalian faunal succession between the Hell Creek and Denver Basin, suggesting that post-K–Pg mammalian recovery was spatially heterogeneous. Our results provided a new model for post-K–Pg mammalian biotic recovery that can now be tested with other earliest Paleocene assemblages across western North America. [ABSTRACT FROM AUTHOR]

Investigating the fossil record to provide evidence about the response of shallow-water tropical calcifiers to past warming events is crucial considering that they are severely threatened by current global warming. We thus focused our attention on scleractinian reef corals (SRC) and shallow-water foraminifera (SWF), both mainly symbiont-bearing organisms, analysing and comparing their diversity patterns during the Paleocene and Eocene from datasets of the Neothetyan circum-Mediterranean region. In particular, at both genus and species level, we analysed changes in diversity, together with origination and extinction rates. Despite some biases related to biostratigraphic resolution of SRC and SWF data, our results show that they reacted differently to the major warming events of the Early Paleogene. The K/Pg mass extinction caused the disappearance of almost all SWF, whereas several corals passed this crisis and persisted up to the end of Paleocene, when the Paleocene-Eocene Thermal Maximum (PETM) caused a relatively small decrease of coral diversity together with a global collapse of coral reefs. On the other hand, a rapid radiation of nummulitids and alveolinids occurred at species level. The impact of the Early Eocene Climatic Optimum (EECO) was apparently more severe for SWF than for SRC. After the Middle Eocene Climatic Optimum (MECO), the general cooling trend led to the fading of Eocene SWF genera and species, whereas SRC began their rapid diversification from Bartonian to Priabonian and culminating in the Chattian. Our results suggest that rapid warming events favoured speciation in SWF, whereas the slow cooling trend (e.g., after the MECO) favoured diversification of SRC. Our data also underline that SWF reacted differently to warming events as compared to deep-sea smaller benthic foraminifera and that the same events in the planktonic realm are not strictly coeval with those occurring in shallow-water environments. On a wider perspective, we observe that shallow-water calcifiers demonstrate a good degree of resilience to global temperature increases, even if undergoing to more or less marked reduction of biodiversity. We provide evidences for the recovery of past ecosystems from both short and long stressors. • Paleocene-Eocene foraminifera and corals biodiversity is affected by warming events. • The response of foraminifera and corals to the same warming events is different. • Foraminiferal diversity is more susceptible to rapid warming events. • Coral diversity appears less influenced by warming events. • Generally, both groups exhibit quite high resilience to warming events. [ABSTRACT FROM AUTHOR]

The advent of the Paleocene-Eocene Thermal Maximum (PETM), a ∼ 200 kyr period of global warming ca. 56 Ma, caused sea-levels to rise, transgressing near-coastal environments in southeastern (SE) Australia over >55,000 km2. During the PETM, warming tropical climates may have extended south to ≥60°S paleolatitude. The PETM in SE Australia is corroborated primarily by stable carbon isotope chemostratigraphy and detailed palynology records in four geological basins. Previous work showed that, in addition to the globally recognised carbon isotope excursion, the PETM interval in coastal SE Australia can be identified using the dual occurrence of the tropical mangrove Nypa palm pollen (Spinizonocolpites prominatus) accompanied by thermophilic marine dinoflagellate cysts (mainly Apectodinium hyperacanthum). We here document a total of twenty-six Gippsland Basin wells that record this Nypa - A.hyperacanthum association in the earliest Eocene Kingfish Formation (Lower Malvacipollis diversus Zone). In the Bass Basin, eight wells record Nypa-A.hyperacanthum association within the Eastern View Group basal Koorkah Formation, or lower part of the Lower M. diversus Zone (earliest Eocene). In the Bass Basin a further thirteen wells with Nypa occurrences near the top of the Cormorant Formation are found, which might be associated with the longer-term warmth of the Early Eocene Climatic Optimum (EECO, ∼53–49 Ma). Government bores and petroleum wells across the Otway Basin record the Nypa-A. hyperacanthum PETM association within the Pember Mudstone Lower M. diversus Zone in twenty-one bores. Nine horizons with Nypa occurrences occur within the Burrungule Member (EECO) at the top of the Dilwyn Formation. In western Tasmania, Nypa occurs in the Sorell Basin and Macquarie Harbour area within the Lower M. diversus Zone. Together, these observations show the remarkable extent of the mangrove-coasts that were established across the mid-high paleolatitudes in SE Australia during the warmest intervals of the Cenozoic, the PETM and EECO. • In 56 Ma SE Australia, 5 m sealevels flooded >55,000km2 of near-coastal environments. • PETM megathermal biota are found at localities down to ∼65°S paleolatitude. • The PETM is identified in the Gippsland, Bass, Otway and Sorell basins. • The PETM contains mangrove Nypa palm pollen (Spinizonocolpites prominatus). • Thermophylic marine dinoflagellates include Apectodinium hyperacanthum. [ABSTRACT FROM AUTHOR]

It has long been debated whether the India-Eurasia collision was a single-stage event that began 60-55 million years ago, or whether it was a two-stage process that involved a collision between India and the Trans-Tethyan Arc before the early Paleocene, and the collision of India with Eurasia during the middle Eocene. Here, we report a late Paleogene angular unconformity (ca. 40-28 Ma) in western Myanmar. This angular unconformity developed around the same time as the Assam unconformity (NE India) but is younger than those found in northern Myanmar. Development of these unconformities indicates that an oblique convergence margin in western Myanmar formed before the middle Eocene, with a major dextral strike-slip fault (proto-Sagaing/Shan Scarp Fault) in the backarc. We interpret this oblique convergence margin to be partial continental collision between the West Myanmar Terrane (WMT) and NE India. In backarc regions, syn-rift successions of the Shwebo sub-basin have formed as a consequence of transtensional tectonics along the proto-Sagaing/Shan Scarp Fault since at least the late Paleocene. The syn-rift successions consist of Asian-derived materials that were not identified in the forearc because of the Wuntho-Popa Arc served as a geographical barrier. The presence of the unconformities and tectonic configuration of the Myanmar backarc sub-basins are inconsistent with the scenario inferred from paleomagnetic data, in which the WMT was part of an intra-oceanic arc at near-equatorial latitudes before the late Oligocene. Instead, we propose that the WMT has been part of continental SE Asia since at least the Paleocene (ca. 60-58 Ma). We reconsider the paleomagnetic data and suggest that the Mawgyi Arc, rather than the WMT, is the oceanic fragment that rifted from the northern Gondwana margin during the Late Jurassic. The Mawgyi Arc collided with continental SE Asia (WMT) during the Late Cretaceous, and then with India during the early Eocene (ca. 51-49 Ma). Our results support the collision between India and Eurasia is a multistage event. • Our findings support a two-stage India-Asia collision scenario. • The unconformities propagated from north to south over western Myanmar during the Paleogene. • Syn-Rift sequences of the Shwebo basin were produced by transtensional faulting. • The West Myanmar Terrane was anchored on SE Asia before the late Paleocene (60-58 Ma). [ABSTRACT FROM AUTHOR]