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In this paper, a novel multihop virtual multiple-input-multiple-output (MIMO) communication protocol is proposed by the cross-layer design to jointly improve the energy efficiency, reliability, and end-to-end (ETE) QoS provisioning in wireless sensor network (WSN). In the protocol, the traditional low-energy adaptive clustering hierarchy protocol is extended by incorporating the cooperative MIMO communication, multihop routing, and hop-by-hop recovery schemes. Based on the protocol, the overall energy consumption per packet transmission is modeled and the optimal set of transmission parameters is found. Then, the issues of ETE QoS provisioning of the protocol are considered. The ETE latency and throughput of the protocol are modeled in terms of the bit-error-rate (BER) performance of each link. Then, a nonlinear constrained programming model is developed to find the optimal BER performance of each link to meet the ETE QoS requirements with a minimum energy consumption. The particle swarm optimization (PSO) algorithm is employed to solve the problem. Simulation results show the effectiveness of the proposed protocol in energy saving and QoS provisioning. [ABSTRACT FROM AUTHOR]